间伐和去除凋落物对马尾松林细根生产力和土壤有机碳含量的影响

Soils play a critical role in the global carbon cycle, and can be major source or sink of CO₂ depending upon land use, vegetation type and soil management practices. Fine roots are important component of a forest ecosystem in terms of water and nutrient uptake. In this study the effects of thinning and litter fall removal on fine root production and soil organic carbon content were examined in 20-year-old Masson pine (Pinus resinosa) plantations in Huitong, Hunan Province of China in the growing seasons of 2004 and 2005. The results showed that fine root production was significantly lower in the thinning plots than in the control plots, with a decrease of 58% and 14% in 2004 and 2005 growing seasons, respectively. Litter fall removal significantly increased fine root production by 14% in 2004. Soil temperature (T_soil) and soil moisture (M_soil) were higher in the thinning plots than those in the controls. Litter fall removal had significant effiects on T_soil and M_soil. Soil organic carbon content was higher in the thinning plots but was lower in the plots with litter fall removal compared with that in the controls. Our results also indicated that annual production of fine roots resulted in small carbon accumulation in the upper layers of the soil, and removal of tree by thinning resulted in a significant increase of carbon storage in Masson pine plantations.     

Response of soil respiration to precipitation during the dry season in two typical forest stands in the forest-grassland transition zone of the Loess Plateau

Forest ecosystems on the Loess Plateau are receiving increasing attention for their special importance in carbon fixation and conservation of soil and water in the region. Soil respiration was investigated in two typical forest stands of the forest-grassland transition zone in the region, an exotic black locust (Robinia pseudoacacia) plantation and an indigenous oak (Quercus liaotungensis) forest, in response to rain events (27.7 mm in May 2009 and 19 mm in May 2010) during the early summer dry season. In both ecosystems, precipitation significantly increased soil moisture, decreased soil temperature, and accelerated soil respiration. The peak values of soil respiration were 4.8 and 4.4 μmol CO₂ m⁻² s⁻¹ in the oak plot and the black locust plot, respectively. In the dry period after rainfall, the soil moisture and respiration rate gradually decreased and the soil temperature increased. Soil respiration rate in black locust stand was consistently less than that in oak stand, being consistent with the differences in C, N contents and fine root mass on the forest floor and in soil between the two stands. However, root respiration (R_r) per unit fine root mass and microbial respiration (R_m) per unit the amount of soil organic matter were higher in black locust stand than in oak stand. Respiration by root rhizosphere in black locust stand was the dominant component resulting in total respiration changes, whereas respiration by roots and soil microbes contributed equally in oak stand. Soil respiration in the black locust plantation showed higher sensitivity to precipitation than that in the oak forest.     

生态抚育对华北落叶松幼龄林枯落物和土壤水文效应的影响

生态抚育是培育森林的基础和关键性技术环节,也是森林生态系统经营的重要组成部分。该文比较了生态抚育措施对华北落叶松(Larix principis-rupprechtii Mayr)幼龄林森林枯落物和土壤水文效应的影响。研究结果表明:生态抚育后改变了林地光照、温度等环境条件,有利于枯落物的分解,抚育后华北落叶松幼龄林林地枯落物总储量减少41%,枯落物自然持水量、最大持水量、有效持水量和实际拦蓄量也明显减小。抚育改变了表土层的结构,从而降低了土壤的容重,提高了林地表层的毛管孔隙度和总孔隙度,改善了土壤的通气性能,提高了土壤的持水量,使抚育后的林地土壤稳渗速率增加(0.33mm/min),林地的土壤水分渗透能力增强,有利于减小或遏制水土流失。     

Short-term effects of thinning on soil respiration in a pine (Pinus tabulaeformis) plantation

Respiration was measured at daytime during the growing seasons (May–October) of 2011 and 2012 in a young Pinus tabulaeformis plantation with heavy, medium and light intensity thinning and unthinned control plots in Shanxi province in northern China. Soil temperature, moisture, fine root biomass, amounts of soil organic C and litterfall biomass were also measured. We found that immediately following thinning treatments, soil respiration increased by 8 %–21 % compared with the unthinned control plots during both growing seasons. Thinning significantly affected soil respiration and soil temperature with different thinning intensities, while there were no significant differences in soil moisture among the various treatments. During the growing seasons, the soil respiration rates were positively correlated with the soil moisture: the 19.4 %–54.0 % variation in soil respiration rates in the four thinning regimes are explained by the changes in soil moisture. Meanwhile, a positive correlation was found between soil temperature and soil respiration rates at all sites. The best fitting model with temperature and moisture explained 44.3 % of the variation in soil respiration in the high thinning treatment, 27.6 % in the light thinning treatment, 18.6 % in medium thinning and in the control sites during the measuring periods. Overall, soil respiration is better predicted by soil moisture, soil organic C, live fine root biomass and soil temperature when data are pooled for all thinning treatments over the two growing seasons. The best regression model explained 74.7 % of the total variation in soil respiration over the different thinning intensities for the two sampling periods.     

大兴安岭林区落叶松林地不同发育阶段土壤肥力演变与评价

 以大兴安岭林区不同发育阶段兴安落叶松人工林和天然次生林为研究对象,采用野外调查和定量分析相结合的方法,对林地土壤有机质、土壤腐殖质组成、土壤酸度指标及其主要养分含量的变化进行测定和分析。利用主成分分析的方法,以各主成分特征贡献率为权重,加权计算各林地土壤肥力综合指标值。结果表明:随着林龄的增长,不同发育阶段落叶松人工林土壤酸度总体呈现升高的趋势,土壤有酸化的迹象;全磷和速效钾呈现降低的趋势;有机质、水解氮、胡敏酸和胡敏素含量在近熟林略有升高,但总体呈现降低的趋势。土壤肥力评价结果为:天然林>幼龄林>近熟林>中龄林>成熟林,即随着林龄的增长,土壤肥力迅速下降,其主要原因是林地凋落物分解缓慢。因此,建议通过适度提高林地抚育强度,调整群落结构,增加林地光照,诱导天然阔叶树种进入,形成针阔混交林的调控措施,调节落叶松人工纯林凋落物分解与积累的矛盾,以维持地力平衡和提高林地生产力。     

内蒙古自治区多伦县不同林地枯落物持水性能研究

[目的]探讨多伦县不同林分类型的枯落物持水性能差异,为该区森林土壤持水性能提供基础数据理论。[方法]以内蒙古自治区锡林浩特市多伦县为研究区,选取代表性的10块林地,收集林地地表枯落物,测定其现存量、持水性能及拦蓄能力等,旨在分析不同林分类型下枯落物持水性能的差异及其对表层土壤(0—20cm)含水率的影响。[结果](1)乔木林的枯落物现存量、持水能力和拦蓄能力均高于灌木林。(2)天然林的枯落物现存量、持水能力和拦蓄能力均高于人工林。(3)各林地枯落物厚度显著影响表层土壤的含水率,即枯落层越厚,表层土壤含水率越高。[结论]由于树种组成、年龄、林分密度及立地条件的影响,不同林分类型的持水能力差异较大,但变化规律为基本一致,同时,林地对土壤水分的影响高于草地。     

改变碳输入对南亚热带海岸沙地典型天然次生林土壤呼吸的影响

土壤呼吸是陆地生态系统碳循环的一个重要过程,开展环境因子和改变碳输入对土壤呼吸影响的研究具有重要意义.2015年3月-2016年2月,在南亚热带海岸沙地典型天然次生林中设置去除根系、去除凋落物、加倍凋落物和对照4种处理,采用LI-8100连续观测改变碳输入对土壤呼吸的影响.结果表明:改变碳输入没有显著影响l0cm土壤温度和湿度(P＞0.05);不同处理土壤呼吸速率存在明显的季节变化,表现为夏高冬低,最大值出现在5月或者6月,最小值出现在11月或12月;土壤呼吸速率的年均值为加倍凋落物＞对照＞去除根系＞去除凋落物,不同改变碳输入方式均降低了土壤呼吸的Q10值;矿质土壤呼吸、凋落物呼吸和根系呼吸对土壤总呼吸的贡献分别为41.24％、43.29％和15.45％;不同处理土壤呼吸速率分别与土壤温度和湿度呈显著的指数和线性正相关(P＜0.05),双因子模型能解释土壤呼吸变异的45％ ～ 69％;改变碳输入影响土壤可溶性有机碳和微生物生物量碳,不同处理土壤呼吸速率与可溶性有机碳和微生物生物量碳呈正相关.因此,改变碳输入引起土壤易变碳的变化进而影响土壤呼吸.     

The effects of mixed broad-leaved trees on the collembolan community in larch plantations of central Japan

The effects of broad-leaved trees on the collembolan community in larch plantations were investigated at the foot of Mt. Yatsugatake (1200–1400 m a.s.l.) in Japan. The study sites comprised five pure larch plantation plots (larch dominated more than 95% of the area at breast height) and five mixed forest plots (larch dominated between 50% and 80% of the area at breast height). We compared the collembolan community structures between stand types and related them to the plant community composition and soil properties at each plot. Density and species richness of Collembola were not significantly different between pure larch and mixed plots. Using partial redundancy analysis (pRDA), the variance of collembolan species data in the litter layer was explained by the biomass of grass on the forest floor, and the variance in the soil layer was explained by the biomass of total forest floor plants. These results suggest that the biomass or the composition of forest floor plants influence the collembolan community more than the crown trees in this area.     

冀北山地天然次生林与人工林地上植被层的水文调控功能

[目的]比较冀北山地天然次生杨桦林(13,18和28a)和人工落叶松林(9,13,15和30a)地上植被层(林冠层、凋落物)的水文调控功能,为评估两种森林类型的转化对该地区水文过程的影响提供科学依据。[方法]采用标准样地和测定分析法对林分的生物量、凋落物及其持水量的进行测算。[结果](1)不同林龄的人工落叶松林地上植被层的降水截留量均大于天然次生杨桦林,且差异显著。(2)两种林分类型不同地上植被层的截留量均表现为:凋落物林冠层灌木层。(3)幼、中龄人工落叶松林的林冠截流量明显大于同龄的天然次生杨桦林,而近熟林,由抚育间伐导致的林分密度下降使得人工落叶松林低于天然次生杨桦林。(4)在各个林龄上,人工落叶松林的凋落物持水量均显著大于天然次生杨桦林,其中13a落叶松为35.36±6.50t/hm2,13a杨桦林为15.79±4.85t/hm2。[结论]冀北山地大面积的人工落叶松林地上植被层具有良好的水文调控功能,其水文调控功能不低于甚至高于当地的天然次生杨桦林,对人工林的抚育间伐在一定程度上会使地上植被层的水文调控功能出现一定程度的下降。     

The microbial communities of aspen and spruce forest floors are resistant to changes in litter inputs and microclimate

Previous studies have shown that forest floors from stands dominated by trembling aspen (ASPEN; Populus tremuloides Michx.) tend to support a greater microbial biomass with a different microbial community structure than forest floors from stands dominated by white spruce (SPRUCE; Picea glauca (Moench) Voss). A reciprocal transfer experiment, in concert with coarse and fine mesh bags that allowed or excluded fine root in-growth, was used to examine how the composition of these forest floor microbial communities respond to changes in belowground inputs from fine roots, aboveground inputs (e.g. from litter and through-fall) and soil microclimatic conditions over 1 year. Neither the microbial biomass nor the microbial community structure (assessed using phospholipid fatty acid analyses and substrate-induced respiration techniques) of forest floors of ASPEN or SPRUCE origin were altered by reciprocal transfer to SPRUCE or ASPEN stands, with or without fine root inputs. Despite the lack of changes in microbial community structure, the stand type during incubation had a strong effect on forest floor moisture content and concentrations of nitrate, while mesh size had a significant effect on forest floor pH and the abundance of mesofauna. Thus, changes in microbial community structure did not co-occur with changes in other characteristics of these forest floors. The resistance of the forest floor microbial communities to change may be a function of the high C contents of these soils. Further treatment effects may have been detected if the study had been extended beyond 1 year. Reciprocal transfer studies using coarse and fine mesh bags allow transferred soils to respond to fluctuations in microclimate, organic inputs and soil biota and, therefore, hold considerable promise for studies examining the influence of disturbances on soil properties.     

华北落叶松人工林短周期间伐与林分生产力的关系

为了研究短期间伐对华北落叶松人工林林分生长和生产力的影响,设置不同保留密度(450、600、750、900株/hm2)的间伐样地以及相应的对照样地,对间伐前后林分的树高、胸径、单株材积、蓄积量、生物量和生产力进行测量与计算。结果表明:不同保留密度下,各间伐样地胸径、树高和单株材积的生长量均大于对照样地;由于保留密度的减小,各间伐样地蓄积量和生物量小于对照样地,各间伐样地生产力均小于对照样地(7 969 kg/(hm2.a));但是其生长率大于对照样地。抚育间伐改变了林分的空间结构和生长环境,明显促进了林木的生长,增加了林地的经济效益。     

Fine Root Dynamics in Thinned and Limed Pitch Pine and Japanese Larch Plantations

ABSTRACT

To investigate fine root dynamics after thinning (50% of standing tree) and liming calcium magnesium carbonate[CaMg(CO₃)₂] 2 Mg ha^{? 1}, a 2-year study was performed in 40-year-old pitch pine (Pinus rigida Mill.) and 44-year-old Japanese larch (Larix leptolepis Gord.) plantations in central Korea. Mean total fine root mass (kg ha^{? 1}± SE) in the control, thinned, and limed plots were 1234 ± 32, 1346 ± 67, and 1134 ± 40 for the pitch pine plantation and 1655 ± 48, 1953 ± 58, and 1868 ± 70 for the Japanese larch plantation, respectively. Live fine root mass of pitch pine at 0-10 cm soil depth decreased after thinning and liming. In addition, liming significantly increased dead fine root mass of Japanese larch. Fine root production (kg ha^{? 1} yr^{? 1}± SE) in the control, thinned and limed plots was 1108 ± 148, 2077 ± 262, and 1686 ± 103 for the pitch pine plantation and 1762 ± 103, 1886 ± 277, and 2176 ± 271 for the Japanese larch plantation, respectively. Fine root turnover rates increased after liming for both plantations. Fine root nitrogen (N) and phosphorus (P) concentrations of Japanese larch (1.012% of N and 0.073% of P) were higher than those of pitch pine (0.809% of N and 0.046% of P) in the control. Also N and P inputs into soil through fine root turnover increased after treatments. Results indicated that comparing fine root dynamics among forest types and after forest management practices might influence differences in soil fertility and underground nutrient cycling.     

Contribution of plant root respiration to the CO2 emission from soil

The respiration activity of roots was studied in field experiments on gray forest and soddy-podzolic soils and under cropland and natural vegetation. It was shown that the contribution of roots to the CO₂ emission from the soil surface depends significantly on the method of determination. The contributions of fine and coarse roots to the total root respiration were approximately similar in forest ecosystems. The use of the method of substrate-induced respiration made it possible to obtain the best estimates of the contributions of root respiration and respiration of microorganisms. The application of glucose in the form of a dry mixture with sand or talc instead of in the water-soluble form appeared to be the optimal procedure for determining the root respiration under field moisture conditions.     

Tree species effects on microbial respiration from decomposing leaf and fine root litter

Tree species have an impact on decomposition processes of woody litter, but the effects of different tree species on microbial heterotrophic respiration derived from decomposing litter are still unclear. Here we used leaf and fine root litter of six tree species differing in chemical and morphological traits in a temperate forest and elucidated the effects of tree species on the relationships between litter-derived microbial respiration rates and decomposition rates and morphological traits, including specific leaf area (cm² g⁻¹) and specific root length (m g⁻¹) of litter at the same site. Litterbags set in forest soil were sequentially collected five times over the course of 18 months. During litter decomposition, microbial respiration from leaf and fine root litter differed among the six tree species. Temporal changes in the remaining mass and morphology (specific leaf area and specific root length) were observed, and the magnitude of these changes differed among species. Positive correlations were observed between respiration and mass loss or morphology across species. These results revealed that litter mass loss and morphological dynamics during decomposition jointly enhanced microbial respiration, and these carbon-based litter traits explained species differences in decomposition of leaves and fine roots. In conclusion, tree species influenced the magnitude and direction of microbial respiration during leaf/fine root litter decomposition. Tree species also affected the relationship between microbial respiration and litter decomposition through direct effects of litter traits and indirect effects mediated by regulation of heterotroph requirements.     

间伐对杉木林土壤CO2通量的影响

Forest management is expected to influence soil CO₂ efflux (FCO₂) as a result of changes in microenvironmental conditions, soil microclimate, and root dynamics. Soil FCO₂ rate was measured during the growing season of 2006 in both thinning and non-thinning locations within stands ranging from 0 to 8 years after the most recent thinning in Chinese fir (Cunninghamia lanceolata (Lamb.) Hook) plantations in Huitong Ecosystem Research Station, Hunan, China. Soil temperature and moisture were also measured to examine relationships between FCO₂ and soil properties. Forest thinning resulted in huge changes in FCO₂ that varied with time since cutting. Immediately following harvest (year 0) FCO₂ in thinning area increased by about 30%, declined to 20%-27% below pre-cutting levels during years 4-6, and recovered to pre-cutting levels at 8 years post-cutting. A similar temporal pattern, but with smaller changes, was found in non-thinning locations. The initial increase in FCO₂ could be attributed to a combination of root decay, soil disturbance, and increased soil temperature in gaps, while the subsequent decrease and recovery to the death and gradual regrowth of active roots. Strong effects of soil temperature and soil water content on FCO₂ were found. Forest thinning mainly influenced FCO₂ through changes in tree root respiration, and the net result was a decrease in integrated FCO₂ flux through the entire felling cycle.     

晋西黄土区不同地类土壤抗冲性分析

为了评价植被对土壤抗冲性的影响,以晋西黄土区蔡家川流域刺槐林、油松林、次生林、退耕荒草地、农地、土质路面为研究对象,采用野外实地放水冲刷法,以侵蚀量为指标,研究不同地类土壤的抗冲性,并分析地被物、根系与土壤抗冲性的相关性.结果表明:不同地类的土壤抗冲性为次生林＞刺槐林＞油松林＞退耕荒草地＞土质路面＞农地;单位面积上枯落物量及其厚度对土壤抗冲性的影响最大;土壤表层根系对土壤抗冲性也具有较大影响,能够有效提高土壤抗冲性的根系结构是0～30 cm土层中的毛根,其中,影响最大的是表层0～10 cm的根系,单位体积土壤内的根系越多,土壤抗冲性越强.     

The effect of boreal forest composition on soil respiration is mediated through variations in soil temperature and C quality

Getting a better understanding of CO₂ efflux from forest soils is critical for increasing our comprehension of the global C cycle. We examined the influence of two common boreal tree species, either in pure stands (BS = black spruce; TA = trembling aspen) or in mixtures (MW = BS + TA mixedwood), on total (R_S), heterotrophic (R_H) and autotrophic soil respiration (R_A) and their relationship with soil temperature and moisture, distance to the nearest tree, labile and total soil organic C (SOC), and root content. Stand-specific soil respiration–temperature models were developed to estimate annual soil CO₂ efflux. Soil temperature was the main factor explaining R_S and its components, followed by labile and total SOC. These three variables were significantly affected by forest composition, while no difference in soil moisture, distance to the nearest tree and root content was observed between stand types. A reciprocal forest floor transplant experiment showed that the influence of stand types on mineral soil temperature was due to a difference in light penetration rather than forest floor characteristics. Annual R_S and R_H were significantly greater in MW and TA than in BS, whereas annual R_A was greater in BS and MW than in TA. Temperature sensitivity (Q₁₀) of both R_S and R_H was significantly higher in BS than in MW and TA, suggesting that CO₂ efflux from BS soils could be increased more under climate warming than that from the other stand types. Our results show evidence that boreal forest composition affects soil CO₂ efflux and that litter quality is not the only factor explaining the differences between stand types. The influence of forest composition on soil CO₂ efflux would be mediated through effects on soil temperature as well as on factors affecting the accumulation and the quality of SOC.     

The bacterial community inhabiting temperate deciduous forests is vertically stratified and undergoes seasonal dynamics

Bacterial communities living in forest soils contribute to the decomposition of organic matter and the recycling of nutrients in these ecosystems and form one of the most diverse habitats on Earth. Unfortunately, due to difficulty in culturing soil bacteria, the understanding of their ecology is still limited. In the case of temperate deciduous forests, soil microbial communities face large seasonal variations in environmental conditions, such as temperature or moisture. Moreover, the supply of nutrients also differs due to seasonal processes, such as the allocation of photosynthates into soil by the roots of primary producers or the seasonal input of fresh litter. The aim of this study was to reveal how the bacterial community responds to these seasonal processes in the litter and soil of a Quercus petraea forest. Bacterial communities from litter and from the organic and mineral horizons of soil were analyzed during the four seasons of the year by 16S rRNA gene pyrosequencing. The results revealed that the composition of the bacterial community is horizon specific. The litter horizon had a higher relative abundance of Proteobacteria and Bacteroidetes than soil, while the organic and mineral horizons had a higher abundance of Acidobacteria, Firmicutes and Actinobacteria than litter. Moreover, the bacterial community was significantly affected by seasonality in all horizons. Bacterial communities in the litter showed significant differences between the vegetation season (May and July) and the autumn and winter seasons (October, February). In mineral soil, bacterial community composition was specific in the summer, when it was significantly different from all other seasons, with a larger number of taxa described as rhizosphere and mycorrhizosphere inhabitants. The results indicate that litter decomposition is the main driver of bacterial community composition in litter horizon. In contrast to reports on fungal communities, bacterial community composition in mineral soil responds to the seasonal peaks of rhizodeposition in the summer.     

Variation of soil respiration at three spatial scales: Components within measurements, intra-site variation and patterns on the landscape

Soil respiration is an important component of terrestrial carbon cycling and can be influenced by many factors that vary spatially. This research aims to determine the extent and causes of spatial variation of soil respiration, and to quantify the importance of scale on measuring and modeling soil respiration within and among common forests of Northern Wisconsin. The potential sources of variation were examined at three scales: [1] variation among the litter, root, and bulk soil respiration components within individual 0.1 m measurement collars, [2] variation between individual soil respiration measurements within a site (<1 m to 10 m), and [3] variation on the landscape caused by topographic influence (100 m to 1000 m). Soil respiration was measured over a two-year period at 12 plots that included four forest types. Root exclusion collars were installed at a subset of the sites, and periodic removal of the litter layer allowed litter and bulk soil contributions to be estimated by subtraction. Soil respiration was also measured at fixed locations in six northern hardwood sites and two aspen sites to examine the stability of variation between individual measurements. These study sites were added to an existing data set where soil respiration was measured in a random, rotating, systematic clustering which allowed the examination of spatial variability from scales of <1 m to 100+ m. The combined data set for this area was also used to examine the influence of topography on soil respiration at scales of over 1000 m by using a temperature and moisture driven soil respiration model and a 4 km² digital elevation model (DEM) to model soil moisture. Results indicate that, although variation of soil respiration and soil moisture is greatest at scales of 100 m or more, variation from locations 1 m or less can be large (standard deviation during summer period of 1.58 and 1.28 μmol CO₂ m⁻² s⁻¹, respectively). At the smallest of scales, the individual contributions of the bulk soil, the roots, and the litter mat changed greatly throughout the season and between forest types, although the data were highly variable within any given site. For scales of 1-10 m, variation between individual measurements could be explained by positive relationships between forest floor mass, root mass, carbon and nitrogen pools, or root nitrogen concentration. Lastly, topography strongly influenced soil moisture and soil properties, and created spatial patterns of soil respiration which changed greatly during a drought event. Integrating soil fluxes over a 4 km² region using an elevation dependent soil respiration model resulted in a drought induced reduction of peak summer flux rates by 37.5%, versus a 31.3% when only plot level data was used. The trends at these important scales may help explain some inter-annual and spatial variability of the net ecosystem exchange of carbon.     

晋西黄土高原水土保持林适宜密度研究

黄土高原区水土流失严重,由于降水、温度等自然气候的限制因素,关于适宜的林分密度的研究至关重要.从林分生长状况,林下枯落物蓄水能力,林下草本植物多样性和林地土壤物理特性等方面,对山西省吉县蔡家川流域不同密度的人工林进行了研究.结果表明,从林分生长状况,恢复地表草本植物多样性,改良土壤物理性质的角度出发,该地区营造刺槐林时密度以1 325株/hm~2为宜,油松林的密度以1733株/hm~2为宜,侧柏刺槐混交林的密度以2 089株/hm~2为宜.从增加林地枯落物的蓄水能力考虑,该地区营造刺槐林时密度以2 133株/hm~2为宜,油松林的密度以2222株/hm~2为宜,侧柏刺槐混交林的密度以2 356株/hm~2为宜.刺槐、油松、侧柏刺槐混交林中,侧柏刺槐混交林地草本植物多样性指数和均匀度指数最高,刺槐林地土壤的非毛管孔隙度最大,土壤涵养水源能力最强.