Influence of environmental productivity levels and yield stability on selection strategies in soybean

Summary Yield data were collected for soybean (Glycine max (L.) Merr.) lines in maturity groups III and IV in 14 environments from 1985 to 1989. The lines in each maturity group were subdivided into three different groups based on the mean yield over all environments, and the genotype × environment interactions were studied for each group. Yield stability of the lines was determined. Effectiveness of selection based on different types of environments was examined.Productivity level of the environment did not influence the relative ranking of the lines. Significant rank correlations occurred between mean yields in most environments and the overall mean yield, but few similarities occurred in the line rankings among individual environments. High-yielding lines contributed a significantly smaller proportion to the genotype × environment interaction than medium- and low-yielding lines.A small proportion of the lines were below or above average stability. Significant correlations occurred among stability, overall mean yield, and mean yield in high- and low-yielding environments. However, few significant correlations occurred between individual environment yields and stability in high- or low-yielding environments. Maturity groups differed in yield and stability relationships. Stability in high- and low-yielding environments did not adequately predict stability for each other.Contribution No. 93-445-J from the Kansas Agricultural Experient Station     

我国水稻种植户风险偏好：理论模型与定量测算

为深入了解我国不同地区水稻种植户风险偏好类型和风险偏好程度,推动粮食生产稳定发展,基于预期效用函数构建理论模型,采用面板数据模型进行回归估计,利用回归结果测算2008~2015年我国23个省市以及分东部、中部、西南、东北四个区域的水稻种植户风险偏好指数。结果表明：我国不同地区水稻种植户都是风险规避者,并且水稻产出越大的地区,农户风险规避程度也越高。因此,对于风险规避的水稻种植户而言,降低产量风险是实现稳粮增收的有效管理手段。由此从加强产量保险推广、增加政府公共品供给、规范农资市场等方面提出了相关建议。     

Genotype × environment interaction for yield and reaction to leaf spot infections in groundnut in semiarid West Africa

Capitalizing on the yield potential in available groundnut germplasm, and high stability of kernel yield are important requirements for groundnut producers in semiarid environments. Forty-seven groundnut genotypes were evaluated from 2003 to 2005 at 4 locations representative of the Guinea and Sudan savanna ecologies in Ghana. The objectives were to assess genotypic differences in reaction to early and late leaf spot infections under natural field conditions, assess the extent of genotype × environment (G × E) interaction for kernel yield, and determine the relationship between yield potential and yield stability. Genotypes differed significantly in their reaction to leaf spot infections indicated by the area under disease progress curve (AUDPC). Genotypic AUDPC was negatively correlated with maturity period (P < 0.01), with kernel yield (P < 0.05) at each of the 3 locations in the Guinea savanna ecology but not in the Sudan savanna ecology and with each of four stability parameters (P < 0.05). High or low yielding genotypes were grouped based on Dunnett’s test at P < 0.10. High yielding groups had significantly low AUDPC, high biomass, high partitioning of dry matter for kernel growth, and were later in maturity compared to low yielding genotypes. Significant G × E interaction effect for kernel yield was dominated mainly by the lack of correlation among environments variance (76–78%) relative to the heterogeneity of genotypic variance component (22–24%). Stability of yield assessed through the among-environment variance, Wricke’s ecovalence, and Finlay-Wilkinson regression coefficient revealed that genotypes in the higher yielding group were relatively unstable compared to the low yielding group. Indicated by the Kataoka’s index of yield reliability, however, relatively unstable genotypes in the high yielding group are expected to be more productive even under assumptions of high risk aversion (P = 0.75–0.95) compared to the more stable, low yielding genotypes. The findings indicate that deploying these recently developed germplasm in semiarid regions in West Africa provides a better match to farmers’ risk-averse strategies compared with the use of existing earlier maturing cultivars.     

Ranking parents by best linear unbiased prediction (BLUP) breeding values in oil palm

Summary Breeding values obtained from best linear unbiased prediction (BLUP) were used to rank nine pisifera (P) male parents belonging to two different but related genetic groups, utilising highly unbalanced data from three D (Dura) × P progeny-test trials of oil palm. The traits studied were fresh fruit bunch yield (FFB), total number of bunches (BNo), average bunch weight (BW), oil to bunch percentage (OB) and average annual height increment (HINC). Four P's from the AVROS genetic group had higher breeding values for FFB, BNo and OB while five P's from the Dumpy-AVROS group were superior in breeding values for BW and HINC. Selection of parents based on the average of the rankings in breeding value for BNo, OB and HINC for each parent tended to favour AVROS P's. However, selection of parents based on the aggregate genetic worth function involving relative economic values, tended to favour Dumpy-AVROS P's. The latter method is preferred to the average ranking method because of its emphasis on relative economic values and the selection index approach.It is suggested that the BLUP technique may prove to be a very useful tool in oil palm breeding.     

Non-parametric measures of phenotypic stability in chickpea genotypes (<Emphasis Type="Italic">Cicer arietinum</Emphasis> L.)

Multi-environment trials (MET) play an important role in selecting the best cultivars and/or agronomic practices to be used in future years at different locations by assessing a cultivar's stability across environments before its commercial release. Objective of this study is to identify chickpea (Cicer arietinum L.) genotypes that have high yield and stable performance across different locations. The genotypes were developed by various breeders at different research institutes/stations of Iran and the International Center for Agricultural Research in Dray Areas (ICARDA), Syria. Several statistical methods were used to evaluate phenotypic stability of these test chickpea genotypes. The 17 genotypes were tested at six different research stations for two years in Iran. Three non-parametric statistical test of significance for genotype × environment (GE) interaction and ten nonparametric measures of stability analyses were used to identify stable genotypes across the 16 environments. The non-parametric measures (Kubinger, Hildebrand and Kroon/Van der) for G × E interaction were highly significant (P < 0.01), suggesting differential responses of the genotypes to the test environments. Based on high values of nonparametric superiority measure (Fox et al. 1990) and low values of Kang's (1988) rank-sum stability parameters, Flip 94-123C was identified as the most stable genotype. These non parametric parameters were observed to be associated with high mean yield. However, the other nonparametric methods were not positively correlated with mean yield and were therefore not used in classifying the genotypes. Simple correlation coefficients using Spearman’s rank correlation, calculated using ranks was used to measure the relationship between the stability parameters. To understand the nature of relationships among the nonparametric methods, a hierarchical cluster analysis based on non weighted values of genotypes, was performed. The 10 stability parameters fell into three groups.     

Investigation of yield and persistence of white clover using cluster analyses

Summary A visual growth score method was used to monitor the growth performance of 24 populations of white clover (Trifolium repens) over a three year period. The mean growth scores for each year were used as population x year data for cluster analyses. Cluster analysis methods based on a regression model where genotypes were grouped for similarity of both intercepts and slopes (Method 1) or slopes only (Method 2), were able to differentiate the persistent populations from those that showed a yield decline over time. The deviations from regression mean square was used as an indication of stability. Method 1 was able to cluster the high yielding persistent populations into a separate cluster, thus facilitating the selection of desirable populations. Deviations from regression identified the stable populations from the cluster. In combination with the simple growth score system, this analytical method is a powerful tool for evaluating a large number of breeding families of perennial pasture clovers and grasses for yield and persistence.     

Genotype × environment interactions for tea yields

Several methods were used to evaluate phenotypic stability in 20 tea (Camellia sinensis) genotypes, many of which are cultivated widely in East Africa. The genotypes were evaluated for annual yields at two sites over a six year period. Data obtained were used to compare methods of analysis of G × E interactions and yield stability in tea. A standard multi-factor analysis of variance test revealed that all first order interactions (genotypes × sites; genotypes × years; sites × years) as well as second order interactions (sites × genotype × years) were significant. Regression analysis was used to assess genotype response to environments. Regression coefficients (b_i) obtained ranged from 0.78 to 1.25. Deviations from regression (S²d) were significant (p < 0.05) from 0.0 for all the test genotypes. Analysis for sensitivity to environment change (SE² _i) revealed that the test genotypes differed in their level of sensitivity. The hierarchical cluster analysis method was used to assemble the test genotypes into groups with similar regression coefficients (b_i) and mean yield, which proved useful for the identification of high yielding genotypes for breeding purposes as well as for commercial exploitation. Rank correlation between yield and some stability parameters were significant. Mean yield was significantly correlated to b_i (r = 0.80^***) and SE² _i(0.74^***) which is an indication that selection for increased yield in tea would change yield stability by increasing b_i and SE² _i leading to development of genotypes that are specifically adapted to environments with optimal growing conditions. Genotypes differed in response to years and sites. As stand age increased, genotype yields generally increased though annual yield fluctuations were more pronounced in some genotypes than others. This response was not consistent across the sites for all genotypes indicating the need to test clones at multiple sites over longer periods of time. This revised version was published online in August 2006 with corrections to the Cover Date.     

Yield and adaptation of hexaploid spring triticale

Summary Gains in yield and stability were analysed using grain yield data from the 8th to the 14th International Triticale Yield Nursery (ITYN) and differences in adaptation between complete and substituted triticales (X Triticosecale Wittmack) were examined. A trend of improving yield and adaptation was observed. As highly significant genotype × site interaction each year was not adequately described using regression analysis, yield stability was examined by ranking genotypes within sites and summarizing scores for individual genotypes across sites. Cluster analysis was used to study the relationships among sites and among genotypes. There was little evidence of sites grouping on a geographical basis. Grouping of genotypes showed a tendency for complete and substituted triticales to perform differently, suggesting that diversity for adaptation is maximized by utilizing both types in breeding.Progress for grain yield is compared with more spectacular gains in test weight and reasons for the different rates of improvement are postulated.Abbreviations CIMMYT International Maize & Wheat Improvement Centre - ITYN International Triticale Yield Nursery contribution from CIMMYT/Agriculture Canada (No. R-002)     

土地整治重大项目社会稳定风险评估初探

构建土地整治重大项目社会稳定风险评估机制,对规范土地整治重大项目建设和土地整治相关管理具有重要的现实意义。本文立足重大项目的全生命周期管理要求,综合分析了土地整治重大项目社会稳定风险的内涵,提出了相应的风险评估过程,并采用PSR模型建立了风险评估体系。研究认为：（1）重大项目的社会稳定风险在项目的立项论证、建设实施和管护运营等阶段均能以不同形式存在,风险评估应根据项目各管理阶段的重点任务及特征,系统分析土地整治过程中可能存在的各类风险因素;（2）风险评估过程应包含风险调查、风险识别、风险评价及风险化解四个程序;（3）基于PSR模型的风险评估体系可反映出土地整治社会稳定风险的产生、传导、作用和控制过程,可为重大项目风险评估指标的选取提供逻辑框架。     

Adaptation to day length and yield stability of families from 4x×2x crosses in potato

Summary The objectives of this research were to evaluate the response to day length and the yield stability of environments of 4x families from 4x×2x crosses in potato. Twenty families from crosses between 4x female parents with different genetic backgrounds and 2x parents, which formed 2n pollen by first division restitution and were selected under short days, were evaluated in four locations in Peru (short days) and during two years at Rhinelander, USA (long days). Results from the USA indicate that the 4x progeny from 4x×2x crosses were very late maturing and poor yielding.Further, vine maturity, tuber size and tuber number are highly correlated with the percentage of Tuberosum germplasm in the parentage. Results from Peru indicate that most 4x×2x families had high tuber yield, stability for yield and high dry matter content. The lack of correlation between rank order of the parental value, based on progeny testing, of the male and female parents among short day locations and between short and long day environments indicates that selection of parents must be done in each location in which the final product, either a family or a clone, will be used.International Potato Center (CIP)     

Associations between nitrogen harvest index and yield characteristics of oats

Summary Fifty lines of oats (Avena Sativa L.) with a broad range of nitrogen harvest indexes (NHI) and nine check lines were evaluated in 15 environments to study the association between NHI and adaptability of oat lines to soils with different productivity levels due primarily to different amounts of N. Three yield characteristics (i.e., mean across environments, regression response to improving environments, and stability of response) were used to measure adaptability. The lines were significantly variable for means of grain and straw yield, for responsiveness to improving environments, and for stability of yield. Among the yield characteristics, only the mean of grain yield was significantly correlated with NHI.Journal Paper No. J-13336 of the Iowa Agric. and Home Econ. Exp. Stn., Ames, Iowa, 50011. Project 2447.     

内蒙古河套灌区主要作物气象产量的风险分析

为弄清作物产量面临的风险水平,通过分析内蒙古河套灌区1953—2011年间12个旗县（市、区）的粮食、小麦、玉米和向日葵的产量资料,利用5年直线滑动平均模型分离实际产量,计算了气象产量的变异系数、年平均减产率、风险概率和综合风险指数等指标,对该区域作物产量灾害风险进行研究。结果表明：河套灌区粮食单产的4种风险指数总体小于小麦、玉米和向日葵的,以向日葵的最大,小麦和玉米的风险指数介于粮食和向日葵的风险指数;地区间以准格尔旗的几种单产的风险要高于其他地区;4种风险指数的最小值,在不同作物上没有明显区域特点。从减产风险概率看,河套灌区作物增产的地区多于减产地区的概率略高一些。     

Stability and genetic improvement of maize yield in Argentina

Summary Yield stability and the genetic improvement of maize (Zea mays L.) grain yield in the humid pampas of Argentina were evaluated.Stability parameters were computed for 15 varieties grown in 35 environments. To analyze genetic improvement of grain yield, data were obtained from trials grown in four locations and twelve years. Two locations represented the typical maize area, while the remaining two were considered marginal. Simple correlation and linear regression coefficients were computed to study the relationships between yield, stability parameters, number of days to 50% tassel emergence, and year of release. Genetic improvement of grain yield was analyzed from linear regression of the average yield of the three highest-yielding varieties (as percentage of the average yield of five common checks) on year of trial.Yield differed significantly among varieties. Significant variety x environment (linear) interaction was also detected. Significant linear relationships were found between regression coefficient for yield (stability parameter 1) and days to tassel emergence, stability parameter 1 and yield, year of release and days to tassel emergence, and year of release and stability parameter 1. Thus, newer maize varieties tended to flower later and had greater responsiveness to favorable environments than did older varieties. Yields have increased for both the typical and marginal areas, with average yearly increases of 114 and 182 kg/ha, respectively.Breeding programs with nurseries located in the typical maize area have raised yield potential in both areas studied. However, if present trends continue, future releases could prove inadequate in low-yielding or short season environments.     

Assessment of drought resistance in 20 accessions of Triticum aestivum and related species I. Total dry matter and grain yield stability

Summary Interspecific hybridisation has been suggested to improve the resistance of crops to biotic and abiotic stresses. To identify sources of resistance to drought which could be used in a breeding programme, an assessment of a large number of wheat species was carried out. The evaluation was based on the estimation of a drought susceptibility index and stability in total dry matter and grain yields of various species grown across a range of soil moisture conditions. On the basis of stability in total dry matter yield, T. aestivum was relatively more drought resistant. However, on the basis of grain yield stability, T. sphaerococcum, T. vavilovii and T. aestivum cv. C 306 exhibited better yield stability and drought resistance than the other species tested. It is suggested that the above species could be utilized as parental material in a hybridisation programme aimed at improving the drought resistance of wheat.     

Selection environment and environmental sensitivity in barley

Summary Groups of 10 barley genotypes were selected for high grain yield under either high yielding (two groups) or low yielding conditions (two groups). The genotypes had a similar average grain yield across a wide range of yielding conditions, but differed in their linear response over environments (environmental sensitivity). The genotypes selected for high grain yield under low yielding conditions were less sensitive to changing environments than genotypes selected for high grain yield under high yielding conditions. The higher stability of genotypes selected under low yielding conditions was shown by both the linear regression analysis and the comparison of coefficients of variation. The use of a safety-first index showed that the probability of a crop failure of genotypes selected for high grain yield under high yielding conditions was between 1.8 and 2.7 times higher than for genotypes selected for high grain yield under low yielding conditions. The development of new cultivars for areas where a large proportion of the crop is grown by subsistence farmers should therefore be based on selection under low yielding conditions.     

The potential for early generation selection to identify potato clones with resistance to Verticillium wilt

Verticillium wilt (VW) of potato, caused primarily by the fungus Verticillium dahliae, results in yield loss and is therefore an important soil-borne disease. Resistance to VW exists in potato germplasm and is used by breeders during cultivar development. Breeders could make more rapid progress toward the development of VW resistant clones if they had an effective early generation selection strategy. The purpose of this study was to determine whether selection for VW resistance could be carried out in the first tuber generation on single hills. One hundred and fifty-two clones from 19 families were planted as single hills on a V. dahliae-infested field. Each plant was scored for vine maturity, VW symptom expression, yield, stem colonization (colony forming units (cfu), in dried basal stem segments) and incidence (percent infected stems). In the second clonal generation, which consisted of replicated four-hill plots, stem colonization scores and incidence values were used to identify clones which were more resistant than a moderately resistant cultivar and others which were more susceptible than a susceptible cultivar. The efficiency and reliability of the single-hill selection strategy, based on symptoms and yield, was then determined by comparison to the four-hill results. We determined that the best single-hill selection strategy was negative selection (discard clones with the lowest performance) with low stringency, based on yield.     

中国马铃薯生产风险区划实证研究

为探寻中国马铃薯生产风险的分布特征及适宜的种植区域,以1992—2013年中国19个马铃薯主产省的数据,综合考虑马铃薯生产过程中自然、市场和技术等风险,以减产率、变异系数、减产率小于10%和30%的发生概率以及种植规模指数等9个变量构建风险区划的指标体系,利用非参数核密度法计算风险发生概率,基于聚类法进行风险区划的实证。结果表明：中国马铃薯种植区被分为5个风险区,中国北部地区马铃薯生产风险较高,中西部地区的生产风险较低,南部地区的生产风险中等。究其原因,是由于马铃薯品种的混种、南北地区气温、降水迥异导致。研究结果为未来中国马铃薯生产布局的调整和防灾减灾措施的制定提供决策依据。     

Simple approach to the analysis of GxE interactions in a multilocational spaced plant trial with timothy

Summary Sixty genotypes of timothy, comprising 12 genotypes originating from each of five locations, were grown at the same locations in northern Scandinavia and Iceland, as a part of a joint breeding project in timothy. The trials were observed for two years and various characters were evaluated. Large genotype x environment interactions occurred for most characters and the response of individual genotypes. irrespective of origin contributed most to the variation. A superiority measure (P.), based on the genotype's response and the maximum response, averaged over all locations, was used to rank the genotypes with respect to yield and other morphological attributes. Based on this ranking it is possible to choose genotypes which are high yielding and show good stability over the area.     

Evaluation of Brassica genotypes for resistance to shatter. I. Development of a laboratory test

Summary A new test procedure for measuring shatter-resistance of siliquae of Brassica spp. is described. A siliqua with its proximal end clamped is loaded as a cantilever in a testing machine. The maximum bending moment, the corresponding stiffness and the energy to cause the siliqua to rupture were taken as measures of the resistance of the siliqua to shatter. Studies showed that ranking of accessions based on either bending moment or energy closely approximated the rankings based on observations of field shatter. A reasonably high significant negative correlation was found between percentage of shattered siliquae and both bending moment and energy (r=–0.595 and –0.531 respecitvely, P=0.001). The above results indicated that bending moment and energy could be used as criteria in evaluating rapeseed genotypes for shatter-resistance in breeding programmes.     

Wheat grain yield and stability assessed through regional trials in the Eastern Gangetic Plains of South Asia

Improving the level and stability of grain yield is the primary objective of wheat breeding programs in the Eastern Gangetic Plains (EGP) of South Asia. A regional wheat trial, the Eastern Gangetic Plains Yield Trial (EGPYT), was initiated by CIMMYT in collaboration with national wheat research programs in Bangladesh, Nepal, and India in 1999–2000 to identify wheat genotypes with high and stable grain yield, disease resistance, and superior agronomic traits for the EGP region. A set of 21 wheat experimental genotypes selected from a regional wheat screening nursery in South Asia, three improved widely grown cultivars (Kanchan, PBW343 and Bhrikuti), and one long-term cultivar (Sonalika) were tested at 9–11 sites in six wheat growing seasons (2000–2005) in the EGP. The 21 experimental genotypes were different in each year, whereas the four check cultivars were common. In each year, one or more of the experimental genotypes showed high and stable grain yield and acceptable maturity, plant height, and disease resistance compared to the check cultivars. Three improved cultivars have already been commercially released in the region through EGPYT and many germplasm lines have been used in the breeding programs as parents. Identification of wheat genotypes with high-grain yield in individual sites and high and stable yield across the EGP region underlines their value for regional wheat breeding programs attempting to improve grain yield and agronomic performance.