Trees,grasses, and weeds: species choices in farmed-developed contour hedgerows

Innovations intended to conserve soils and improve soil nutrient cycling have not been widely adopted because of technical problems and lack of fit with farmers' cricumstances. Research in Claveria, the Philippines, facilitated and monitored farmers' adaptation of contour hedgerows to fit their particular needs. Farmers tested different establishment methods and many hedgerow species. They initially planted a combination ofGliricidia sepium (a legume tree) andPennisetum purpureum. (grass). Later adoptors chose fodder grasses (especiallySetaria spp.) or naturally occuring vegetation in their hedgerows-either solely or in combination with other species, including weeds such asRottboellia cochinchinensis identified as crop problems. Upland rice and maize farmers who adopted contour hedgerows from 1987 through 1991 were interviewed in 1992. Although hedgerow-crop competition, grazing by neighbors' cattle, and added labor were problems, farmers viewed hedgerows as a way to reduce soil erosion and provide fodder. Farmers planting mulberry were disappointed after a silkworm project folded. Farmers now face the problem of soil nutrient depletion, leading to fallowing of fields with hedgerows and shifting to other parcels. A farmer decision tree model of the minimum necessary criteria for sustainable adoption of contour hedgerows is hypothesized.     

Nitrogen fixation capacity in the component species of contour hedgerows: how important?

The choice of an appropriate hedgerow species is one of the most critical decisions in exploiting the value of a contour hedgerow system. The implications of hedgerow species with nitrogen (N)-fixation capacity on hedgerow-crop competition and crop productivity have been widely debated. We examined the agronomic significance of N-fixation by comparing the performance of species representing three classes of hedgerow vegetation: A nitrogen-fixing tree legumeGliricidia sepium), a non-nitrogen fixing tree (Senna spectabilis syn.Cassia spectabilis), and a forage grass (Pennisetum purpureum). The 4-year study investigated the hedgerow biomass and nutrient yields, and their relative effects on the performance of two annual crops commonly grown in alley farming systems, with emphasis on hedgerow-crop interference. The work was done on an Ultic Haplorthox (pH 4.8, organic C 1.9%, total N 0.18%).Senna produced 46% more pruning biomass on an annual basis than didGliricidia; N supplied to the alley crops was similar toGliricidia in the first year of observation, but 20–30% higher in the succeeding years. Upland rice and maize grain yields and total dry matter were unaffected by tree species, but the nitrogen-fixing tree exerted less competitive effects on the annual crops growing in adjacent rows. Grass hedgerows reduced maize yields 86% by the second year, indicating an unsustainable drawdown of nutrients and water. We conclude that hedgerow systems composed of a nitrogen-fixing tree did not exert significant advantages compared to a non-fixing tree species, and that factors other than N-fixation were more important determinants for the choice of hedgerow species.     

Productivity of alley farming with leucaena (Leucaena leucocephala Lam. de Wit) and Napier grass (Pennisetum purpureum K. Schum) in coastal lowland Kenya

Inadequate supply of fodder is a serious constraint to the potentially-promising small-holder-dairy production system in coastal Kenya. Alley farming could be an approach to addressing this problem. A study of forage production based on Napier grass and leucaena in an alley cropping system was conducted on an infertile sandy soil in lowland coastal Kenya. The effects of leucaena hedgerows,Clitoria ternatea (L.) intercropping, addition of slurry (110 t ha^–1 yr^–1) and two harvesting managements (severe and lenient) on the yield of Napier grass fodder, were assessed. The study was initiated in 1989 and three years results are reported.Napier grass fodder yields did not decline in the presence of leucaena heggerows in most harvests. Generally, the Napier rows adjacent to the hedgerows gave greater yields than in either the centre or sole Napier rows. Interplanting clitoria between Napier rows had a beneficial effect on Napier yields in the later harvests. Compared with sole Napier, the hedgerow treatment receiving slurry increased yield ha^–1 of Napier by 50%, of total forage by 80%, and of nitrogen (measured over two harvests) by 200%.During the first two years of cropping most treatments did not affect the OM and nitrogen content of the soil but by the end of the third cropping year, slurry application increased soil OM by 21% and slurry and clitoria treatments each increased soil nitrogen by 33%.The study demonstrated the increases in the supply and quality of forages for dairy cattle that can be achieved by planting complementary forages, legume and grass species in an intercropping system like alley farming and the benefits to soil composition by the return of slurry.     

Impact of contour hedgerows on upland maize yields in the Philippines

An agricultural production function is used to examine the yield impacts of contour hedgerows constructed using double rows of the N₂-fixing species Desmodium rensonii and Flemengia macrophylla. The analysis uses data from a sample of upland Philippine maize farms and controls for input use and plot characteristics. Results support a hypothesis that the long-run impact of hedgerows on maize yields is positive. However, results suggest that in the short run, hedgerows not only reduce area available for cultivation on a plot, but also reduce the performance of corn in the remaining alleys. For newly established hedgerows an increase in hedgerow area is associated with a reduction in maize yield. The time required for hedgerows to compensate for the area they occupy is found to be approximately eight years. This revised version was published online in August 2006 with corrections to the Cover Date.     

Agroforestry for soil and water conservation in the western Himalayan Valley Region of India 2. Crop and tree production

A ten-year-study (1983 to 1992) conducted on nine 15 × 90 m runoff plots at 4% slope compared production efficiency of Leucaena leucocephala and Eucalyptus hybrid based agroforestry as well as monocropping landuse systems in the warm, subhumid climate of the western Himalayan region of India. Treatments for the first sequence were: monocropping systems of leucaena, eucalyptus, Chrysopogon fulvus grass and maize – wheat rotation, and alley cropping systems of grass and crops at 4.5 and 10.5 m alley widths with paired contour tree rows of leucaena and eucalyptus. In the second sequence, alley width increased to 22.5 m in 1989, grass was replaced by turmeric Curcuma longa and paired contour rows of leucaena hedges were introduced in monocropping systems of grain crops and turmeric. Integration of leucaena and eucalyptus trees with crops caused severe reduction of crop yields ranging from 21 to 92% for wheat grain, 59 to 69% for maize grain, 60 to 67% for dry grass and about 50% for turmeric rhizome depending upon the age of trees and alley width. The grain yield of crops stabilized at about 50% reduction with 22.5 m alley width. Total crop biomass (grain + straw) also revealed a similar trend; however, its magnitude of reduction was less severe than for grain. Production of biomass was much lower near the tree rows than in mid alleys. Managing leucaena as contour hedgerows eliminated crop yield reduction in alleys. Performance of grass and turmeric in alleys was not found to be satisfactory. Biomass produced from trees adequately compensated the crop yield reduction. Land equivalent ratios of agroforestry landuses were comparable or even better than monocropping systems indicating suitability of these systems for the western Himalayan valley region. This revised version was published online in June 2006 with corrections to the Cover Date.     

The effects of between-row (alley widths) and within-row spacings ofGliricidia sepium on alley-cropped maize in Sierra Leone

The effects of spacings between hedgerows (alley widths) and the spacings of trees within hedgerows ofGliricidia sepium on growth and grain yield of maize were investigated at Senehun in southern Sierra Leone. Four between-row spacings (2, 4, 6 and 8 m) were combined with three within-row spacings (0.25, 0.50 and 1.00 m) in a split block design. Maize, at densities of 20,000, 40,000 and 53,333 plants ha^–1, was established in the alleys and also as pure crops. N, P and K fertilizers were applied to all plots before pruning of the trees began. When pruning started, only the pure maize plots received fertilizer; prunings from the hedgerows were returned to the appropriate alleys in the other plots.Plots with the highest maize populations consistently gave the best yields before pruning started, but lower populations gave improved yields after pruning. Yields of maize increased with increasing alley widths before the start of pruning, after which the narrower alleys of 2 and 4 m outyielded the wider ones by almost double, probably because of the large amount of nutrients applied in prunings. Lack of light limited grain yields before the start of pruning, when there was some shading by the hedgerows. Alleys of 2–4 m wide, planted no closer than 0.50 m within rows, resulted in more than twice the yields of maize than in the 8-m alleys planted at 0.25 m within rows, once the hedgerows were well established and were being managed.     

Tree-crop interactions in hedgerow intercropping with different tree species and tea in Sri Lanka: 1. Production and resource competition

Incorporation of tree hedges along contours has been proposed as a means of reducing soil erosion and increasing soil fertility of tea (Camellia sinensis (L.) O. Kuntz) plantations on sloping terrain in high-rainfall zones of Sri Lanka. Tea yields in these hedgerow intercrops are determined by the balance between the positive (i.e., increased soil fertility) and negative (i.e., resource competition) effects of hedgerows. Therefore, the objectives of the present study were to determine: (a) the net effect of six different hedgerow species (Calliandra calothyrsus Meissner, Senna [Cassia] spectabilis (DC.) H. Irwin and Barneby, Euphatorium innulifolium (R.M. King and H. Rob.) H.B.K., Flemingia congesta Aiton F., Gliricidia sepium (Jacq.) (Kunth.) Walp. and Tithonia diversifolia (Hemsl.) A. Gray) on long-term tea yields; (b) the effect of incorporating hedgerow prunings as mulch; and (c) the relationship between hedgerow characteristics (i.e., biomass production, canopy and root growth) and tea yield variation. Tea yields, measured over one complete pruning cycle from October 1998 to September 2001, showed reductions relative to a sole tea crop under all hedgerow species except Euphatorium. The yield reductions ranged from 22 to 40%. Tea yields under Euphatorium showed increases up to 23% relative to the sole crop control. Addition of hedgerow prunings as mulch increased tea yields in all hedgerow intercrops. The yield increases ranged from 11 to 20%, with the highest being under Euphatorium. Tea yields showed a negative relationship (R²=0.38) with the pruned biomass of hedgerows. Limitation of environmental resources (e.g., water and light) and hedgerow characters which intensified resource competition (i.e., greater canopy lateral spread and height and greater root length densities, especially in the top soil layer) were responsible for observed tea yield reductions in hedgerow intercrops.     

Alley cropping Sesbania sesban (L) Merill with food crops in the highland region of Rwanda

Food production in the densely populated Rwandan highlands is impeded by soil erosion and loss in fertility. Alley cropping leguminous shrubs with food crops on contours is purported to minimize the problem and to provide wood and forage. This study reports the effect of Sesbania prunings plus moderate levels of N and P on bean (Phaseolus sp) and maize (Zea mays) yields in alley cropping. Experimental design was a randomized complete block with split-split plots. Main plots were alley width: 2, 4, 6 and 8 m. Phosphorus (P) at 0, 30 and 60 kg P₂O₅/ha occupied the subplot and nitrogen (N) at 0, 30 and 60 kg/ha were assigned at the sub-sub plot level. No P was applied to maize during the second cropping season. Crop yield in kg/ha included the land space taken by hedgerows. Bean yield in 6 m alleys (1100 kg/ha) was about twice that in 2 m alleys (500 kg/ha). Bean responded to N and P. Optimum alley width and N for bean yield were 6 m and 30 kg/ha, respectively. Cuttings from alley hedgerows provided stakes for climbing beans. Maize responded to N but not to residual P. The highest maize yield came from 8 m alleys with 40 kg/ha, but yields from 8 and 6 m alleys with the same N treatment were not significantly different. Maize plants in middle rows were significantly taller than plants in rows adjacent to hedgerows. Maize rust development showed significant alley width and row position effect. There were significantly fewer uredinia in the Sebania alleys relative to the control plots without shrub hedgerows. Rust development on maize in middle rows was significantly greater than development in border rows.     

Cost-benefit analysis of alternative forms of hedgerow intercropping in the Philippine uplands

Considerable resources have been expended promoting hedgerow intercropping with shrub legumes to farmers in the Philippine uplands. Despite the resources committed to research and extension, persistent adoption by farmers has been limited to low cost versions of the technology including natural vegetation and grass strips. In this paper, cost-benefit analysis is used to compare the economic returns from traditional open-field maize farming with returns from intercropping maize between leguminous shrub hedgerows, natural vegetation strips and grass strips. An erosion/productivity model, Soil Changes Under Agroforestry, was used to predict the effect of erosion on maize yields. Key informant surveys with experienced maize farmers were used to derive production budgets for the alternative farming methods. The economic incentives revealed by the cost-benefit analysis help to explain the adoption of maize farming methods in the Philippine uplands. Open-field farming without hedgerows has been by far the most popular method of maize production, often with two or more fields cropped in rotation. There is little persistent adoption of hedgerow intercropping with shrub legumes because sustained maize yields are not realised rapidly enough to compensate farmers for establishment and maintenance costs. Natural vegetation and grass strips are more attractive to farmers because of lower establishment costs, and provide intermediate steps to adoption. Rural finance, commodity pricing and agrarian reform policies influence the incentives for maize farmers in the Philippine uplands to adopt and maintain hedgerow intercropping. This revised version was published online in August 2006 with corrections to the Cover Date.     

Alley cropping in the moist savanna of West-Africa: II. Impact on soil productivity in a North-to-South transect in Togo

The moist savanna of West-Africa is characterized by a wide range of climates and soil types. The impact of the biophysical environment on hedgerow N uptake, wood production and maize grain yield was assessed for three years in three alley cropping trials with a selected number of hedgerow species in Glidji (Southern Togo), Amoutchou (Central Togo), and Sarakawa (Northern Togo). Senna siamea hedgerows accumulated significantly more N in the first pruning in Glidji (129−138 kg N ha−1) and Sarakawa (102−185 kg N ha−1) than in Amoutchou (17–26 kg N ha−1). This difference in N uptake was attributed to the infertile subsoil in Amoutchou, which was sandy up to 1 m and had a shallow groundwater-table. The amount of N accumulated in the Gliricidia sepium biomass varied between 38 kg N ha−1 in Glidji and 142 kg N ha−1 in Amoutchou. Averaged over all species and sites, 9 to 29% and 9 to 39% of the annual N accumulation in the hedgerow biomass is incorporated in the second, respectively third pruning. The Gliricidia trees produced between 12 and 26 ton fresh matter ha−1 of wood and the Senna trees between 4 and 38 ton fresh matter ha−1. Maize grain yield in Glidji was not affected by treatments (3196 kg ha−1, on average). In Amoutchou, the highest grain production was observed in the Gliricidia treatment (2774 kg ha−1 vs 1007 kg ha−1 in the control), while in Sarakawa, the Gliricidia (3786 kg ha−1) and Senna (3842 kg ha−1) plots produced a greater grain yield than the control plots (2123 kg ha−1). Maize yield increase in the alley cropping systems relative to the control plots was related to the soil total N content. Top and sub-soil characteristics were shown to be an important modifier of the functioning of alley cropping systems and should be taken into account when deciding on whether to use alley cropping and when selecting the hedgerow species. This revised version was published online in June 2006 with corrections to the Cover Date. This revised version was published online in June 2006 with corrections to the Cover Date.     

Pattern of soil moisture depletion in alley cropping under semiarid conditions in Zambia

The pattern of soil moisture changes was studied during a cropping season in an alley cropping experiment of maize withLeucaena leucocephala andFlemingia macrophylla at the SADC/ICRAF Agroforestry Research Station in a semiarid region near Lusaka, Zambia (28°2956 East and 15°2132 South). Soil moisture potential was monitored at regular intervals using tensiometers installed at 15, 30 and 45 cm depths in fertilized and unfertilized alleys within the double hedgerow, and the first, second and third rows of maize in the alleys.Soil moisture moved mostly towards the top horizon during very dry conditions. Alleys that had received a combination of fertilizer and hedgerow prunings depleted more moisture than those that had only hedgerow prunings. There were no differences in moisture utilization pattern between leucaena and flemingia hedgerows. The hedgerows depleted the same amount of moisture as the maize plants. However, during dry conditions, there was a higher soil moisture content under the hedgerows than in maize rows, indicating that there was no apparent competition for moisture between the hedgerows and the maize plants.     

Potential contribution of Leucaena hedgerows intercropped with maize to the production of organic nitrogen and fuelwood in the lowland tropics

Available information is applied to formulate quantitative hypotheses on the impact of intercropping Leucaena hedgerows with maize upon the physical productivity of grain and fuelwood. Data would indicate that productivity of organic nitrogen (N) by Leucaena hedgerows cut approximately every 8 weeks at a height of 15–30 cm and planted at a distance between rows wider than 150 cm is 45 g m^?1 yr^?1. When soil-N availability is the limiting factor, utilization of Leucaena-N by the maize crop appears to be negatively related to baseline maize production. The grain: Leucaena-N ratio declines from 20:1, when maize productivity is in the order of 500 kg ha^?1, to 3:1, when the 4000 kg level is achieved. Hedgerow N productivity, N utilization by the maize crop, and proportion of land planted to maize were used to derive yield estimates per area of intercropped land under different intercropping arrangement. It seems that the impact of hedge intercropping on maize productivity, although substantial, would be limited to systems where existing production levels of maize are lower than 1500 kg ha^?1. As expected, production per hectare decreases as spacing between Leucaena hedgerows increases. For a 1000 kg ha^?1 baseline, hypothetical increments expressed as percentage of baseline production range from 112% to 28% for between-hedgerow spacings of 1.5 and 6 m respectively. Information analyzed would indicate apotential for Leucaena hedgerow intercropping to increase maize productivity. Research required to substantiate the formulated hypotheses is discussed.     

Variable responses of the depth of tree nitrogen uptake to pruning and competition

Trees in cropped fields may improve nitrogen (N) use efficiency by intercepting leached N, but crop yield will be reduced if the trees compete strongly with crops for N. Ideal trees for intercropping will take up N from deeper soil layers not accessed by the crop species. Spatiotemporal aspects of tree nitrogen capture niches were investigated within a hedgerow intercropping system by placing 15N at three depths and monitoring 15N uptake by trees pruned either 25 or 4 days before application of 15N. Trees with contrasting rooting patterns (Gliricidia sepium L. and Peltophorum dasyrrachis (Miq.) Kurz) were grown in mixed hedgerows and intercropped with maize (Zea mays L.). Neither species showed significant N uptake during the 5-14 days after pruning, even though some shoot regrowth occurred during this time. Mean topsoil (0-5 cm depth) root length density of G. sepium was 520% greater than that of P. dasyrrachis, whereas total root length (0-100 cm depth) of G. sepium was only 450% greater. On average, G. sepium recovered 15 times as much 15N as P. dasyrrachis, following application of 15N at 5 cm depth, but the two species recovered a similar amount following application at 80 cm depth, suggesting that P. dasyrrachis had better niche complementarity with shallow rooting crops. However, both species showed strong plasticity in vertical N uptake pattern in response to competition from establishing maize plants. The species differed in their response: N uptake activity by G. sepium shifted down the soil profile in response to increasing competition from a growing maize crop (uptake from 80 cm depth changed from 4 to 9% of uptake from 5 cm depth), whereas N uptake by P. dasyrrachis became relatively shallow (uptake from 80 cm depth changed from 305 to 25% of uptake from 5 cm depth). Niche avoidance and increased competitiveness within the topsoil represent alternative responses to competition. The response displayed may be related to soil fertility in the species' natural habitats.     

Soil property changes in contour hedgerow systems on sloping land in the Philippines

The impact of contour hedgerow systems on soil sustainability under acidic conditions has been widely criticized. A study was undertaken to determine the effects of management and hedgerow species on soil properties. Cassia spectabilis (a non-N-fixing tree legume), Gliricidia sepium (an N-fixing tree legume), Pennisetum purpureum (a forage grass), and Stylosanthes guyanensis (a forage legume) contour barriers were compared with an open field (non-hedgerow treatment) over 1 cowpea and 2 rice seasons. Three types of management viz.: prunings applied + N₀P₀K₀, prunings applied + N₅₀P₂₀K₂₀ and prunings removed + N₅₀P₂₀K₂₀ were used as subplot treatments. The soils were strongly acidic (pH 4.5) and classified as clay Orthoxic Palehumult. Cassia performed better than the other species in terms of pruning biomass, N and P contributions over a period of 20 months. There was a combined positive effect of pruning biomass and fertilizers on rice and cowpea yields in Pennisetum and Gliricidia systems, while a tendency towards a positive effect of pruning biomass on rice was found in the Cassia system. The pruning biomass and/or fertilizer application did not significantly influence the top soil organic C, N and available P in the hedgerow systems. Soil bulk density was significantly reduced by the application of Cassia prunings after 12 months. Organic C, N and P dynamics indicated that in situ pruning biomass was not sufficient to maintain their level in the soil. But the cassia systems with prunings applied + N₅₀P₂₀K₂₀ experienced the lowest degradation in soil organic C (2.1 t ha⁻¹) followed by the Gliricidia systems (4.1 t ha⁻¹). The overall results imply that the application of pruning and inorganic fertilizer is imperative to conserve soil resources, and non- N-fixing tree species can exert a significant advantage in biomass and thereby in soil N-recycling under acidic soil. This revised version was published online in June 2006 with corrections to the Cover Date.     

Above- and below-ground biomass dynamics in a sole cropping and an alley cropping system withGliricidia sepium in the semi-deciduous rainforest zone of West Africa

A considerable amount of data is available about above-ground biomass production and turnover in tropical agroforestry systems, but quantitative information concerning root turnover is lacking. Above- and below-ground biomass dynamics were studied during one year in an alley cropping system withGliricidia sepium and a sole cropping system, on aPlinthic Lixisol in the semi-deciduous rainforest zone of the Côte d'Ivoire. Field crops were maize and groundnut. Live root mass was higher in agroforestry than in sole cropping during most of the study period. This was partly due to increased crop and weed root development and partly to the presence of the hedgerow roots. Fine root production was higher in the alleys and lower under the hedgerows compared to the sole cropping plots. Considering the whole plot area, root production in agroforestry and sole cropping systems was approximatly similar with 1000–1100 kg ha^–1 (dry matter with 45% C) in 0–50 cm depth; about 55% of this root production occured in the top 10 cm. Potential sources of error of the calculation method are discussed on the basis of the compartment flow model. Above-ground biomass production was 11.1 Mg ha^–1 in sole cropping and 13.6 Mg ha^–1 in alley cropping, of which 4.3 Mg ha^–1 were hedgerow prunings. The input of hedgerow root biomass into the soil was limited by the low root mass ofGliricidia as compared to other tree species, and by the decrease of live root mass of hedgerows and associated perennial weeds during the cropping season, presumably as a result of frequent shoot pruning.     

The effects of alley cropping withLeucaena leucocephala and of different management practices on the productivity of maize and soil chemical properties in lowland coastal Kenya

The effects of leucaena hedgerows, mulching with leucaena foliage (0,50 and 100% of harvested foliage), cowpea intercropping and adition of dairy cattle slurry (55 t ha^–1 per maize crop) on the yield of maize from a sandy soil were assessed. The four-year results from five maize crops are reported.Except in the first year, yields of maize grain and stover were significantly reduced by 30% in the presence of leucaena hedgerows. Use of leucaena mulch eliminated this effect; application of all the harvested leucaena mulch (100%) increased the total maize grain yield of the five crops by 44% over sole maize. Hedgerow and mulching management required an additional 36 mandays labour ha^–1 which was more than compensated by the increased maize yields. Furthermore leucaena hedgerows substantially depressed the growth of weeds between cropping seasons.Intercropping with cowpea significantly depressed yields of maize grain and stover when both crops were sown together, but not in later seasons when cowpea was sown four weeks after the maize. Application of slurry increased the total yields of maize grain and stover by 35 and 37%, respectively. The grain yield of maize in leucaena hedgerow treatments fertilized with slurry did not respond to application of more than 50% of leucaena foliage, which suggested that half of the foliage could be spared for feeding to livestock. The cumulative yield of maize grain from the highest yielding organic system was 85% of the yield from the fertilizer treatment.The study, which is continuing, demonstrates that large increases in agricultural productivity are possible through the intercropping of maize with woody forage and grain legumes and the integration of dairy cattle production into the system. It thus shows the importance of exploiting crop/livestock interactions.     

Insect pests and beneficial arthropods population under different hedgerow intercropping systems in semiarid Kenya

Exploiting the principles of biological control of crop pests is an important management strategy for the resource-poor smallholder farmers of the tropics. The effects of nine hedgerow species on the abundance of major insect pests of beans and maize, and predatory/parasitic-arthropods were monitored over two cropping seasons and the intervening dry period. The tree/shrub species evaluated were Gliricidia sepium, Grevillea robusta, Senna siamea, Senna spectabilis, Flemingea congesta, Croton megalocarpus, Morus alba, Calliandra calothyrsus and Lantana camara. Arthropod abundance was monitored through counts of infested plants and using yellow pan and pitfall traps. Beanfly (Ophiomyia spp.) infestation was significantly higher in the presence of hedgerows (35%) than in their absence (25%). Hedgerows did not influence aphid (Aphis fabae) infestation of beans. In contrast, maize associated with hedgerows experienced significantly lower stalk borer (Busseola fusca and Chilo spp.) and aphid (Rhophalosiphum maidis) infestations than pure maize, the margin of difference being 13% and 11% respectively for the two pests. Ladybird beetles closely followed their prey, aphids, with significantly higher catches in sole cropped-plots than in hedgerow-plots and away from hedgerows. Activity of wasps was significantly greater close to hedgerows than away from them. Spider catches during maize season were 77% greater in the presence of hedgerows than in their absence, but catches during other seasons were similar between the two cropping systems. Differences among hedgerow species were not significant for most of the arthropods monitored, except that beanfly infestation was greater with Gliricidia hedgerows. The study indicates that the effect of hedgerows on pest infestations of crops and their role as refugia for predators cannot be generalized but depends on the specific arthropods.This revised version was published online in November 2005 with corrections to the Cover Date.     

Calliandra calothyrsus (Meissn.) in an alley cropping system with sequentially cropped maize and cowpea in southwestern Nigeria

Alley cropping is increasingly becoming accepted as an appropriate technology with the potential to provide stable and sustainable food production in the tropics. However, only a few of the potential trees/shrubs have been tested. The performance of Calliandra calothyrsus (Meissn) as an alley cropping species was evaluated on an Oxic Paleustalf. The treatments were: prunings removal, prunings application; and three N levels, 0,45, and 90 kg N ha^–1, in a factorial arrangement. The cropping sequence was maize (main season) follwed by cowpea (minor season). Nitrogen fertilizer treatments were applied to the maize crop only. Four annual prunings of Calliandra hedgerows produced a total of 6 t ha^–1 of dry matter prunings containing about 200 kg ha^–1 of N. Maize yields were increased by the application of prunings but no benefits were obtained by supplementing the prunings with inorganic N. An average maize grain yield of 3.1 t ha^–1 per year was maintained without any chemical fertilizer input. However, without any prunings, maize yields were substantially increased by the application of inorganic N. Cowpea yield did not respond to application of prunings but plants grown adjacent to the hedgerows had reduced yield probably due to shading. Our results suggest that six rows (0.57 m inter-row spacing) between Calliandra hedgerows spaced at 4 m are optimum for this cowpea variety. The performance of Calliandra was comparable to that of Leucaena which has been widely shown to be effective in alley cropping systems of the region.     

A study of the potential of hedgerow intercropping in semi-arid India using a two-way systematic design

The potential of hedgerow intecrropping with Leucaena leucocephala was explored on vertic Inceptisols over 4 years at ICRISAT Center, Patancheru, India. The study was conducted using a systematic layout involving different alley widths ranging from 1.35 to 4.95 m and with varying distances between hedge and crops. The alleys were cropped with alternate rows of sorghum and pigeonpea. Hedges composed double Leucaena hedgerows 60 cm apart were periodically harvested for fodder. Sole crops of all components and a sorghum/pigeonpea intercrop were included in all four replications of the study.Starting in the second year, Leucaena was progressively more competitive to annual crops, causing substantial yield reduction. Competition (primarily for moisture) was most severe in narrow alleys and was greatest on pigeonpea.The growth of Leucaena was not sufficient to compensate for reduced crop yields. Land equivalent ratios (LERs) calculated on the basis of grain yield of crops and Leucaena fodder yields showed that hedgerow intercropping (HI) was advantageous over sole crops only during the first two years using wide alleys, but disadvantegeous in the last two years. LERs calculated on the basis of total dry matter indicated only a small advantage for HI (13–17 percent) over sole crops in wider (>4 m) alleys. Average returns per year from HI exceeded those of the most productive annual crop system (sorghum/pigeonpea intercropping) by 8 percent in 4.05 m alleys, and by 16 percent in 4.95 m alleys. Fodder production during the dry season was 40 percent of the annual total in these alley widths. Thus hedgerow intercropping at 4–5 m alley width is not very attractive for farmers in semi-arid India, which has 600–700 mm of annual rainfall. There is a need to examine the potential of HI in wider alleys. The merits and limitations of the systematic design are discussed.     

Cutting management of alley cropped leucaena/gliricidia-Guinea grass mixtures for forage production in southwestern Nigeria

A study was conducted at Fasola (7°45 N and 3°5 E) in southwest Nigeria to determine the best tree cutting scheme for forage production and the effects of hedge configurations on tree, grass and total forage productivity of 6–8-year-old leucaena (Leucaena leucocephala Lam. de Wit) and gliricidia (Gliricidia sepium Walp.)-Guinea grass (Panicum maximum Jacq. cv. Ntchisi) mixture. After a uniform cut at the end of January 1990 (mid-dry season), the trees were cut according to the following cutting regimes: one cut after a 12-month regrowth (12M); two cuts after three- and nine-month regrowth (3–9M); two cuts every six months (6-6M); two cuts after nine- and three-month regrowth (9-3M); three cuts, two cuts every three months and the third cut after a six-month regrowth (3-3-6M), and four cuts every three months (3-3-3-3M). Grass was cut every six weeks between April and October followed by a cut in January. The hedge configuration was either one or three hedgerows of mixed stands of leucaena and gliricidia. There were twice as many trees and one-third less grass in the triple than in the single hedgerow hedge configuration.The 3-3-3-3M, 9-3M and 3-3-6M cutting regimes produced the highest total forage (tree foliage + grass) dry matter yields (DM) of 6.54, 5.80 and 5.77 t DM ha^–1 annum^–1, respectively. The magnitude of the difference between the tree forage yields of the triple and single hedgerow plots (16%) did not reflect the theoretical difference in the number of trees (33%) in the two arrangements.