Energy requirements of young female cattle. 6. Energy requirements

Results taken from 6 experiments with young female cattle comprising 477 metabolism periods served the derivation of the animals energy requirement in the development range greater than 125 kg live weight according to the factorial method. The energy requirement per kg LW0.75 and day, calculated from metabolism data, was independent of the stage of development and the intensity of feeding. It averaged 455 +/- 66 kJ metabolizable energy/kg LW0.75.d and 250 +/- 37 kJ NEFcattle/kg LW0.75.d respectively. The partial energy requirement for live weight gain, expressed in net energy fat, was equivalent to the energy content of the live weight gain. Energy retention and thus energy requirement per kg live weight gain increased with the live weight and reached a maximum of 26 MJ. Energy retention per kg live weight gain largely depended on the intensity of feeding and the stage of gravidity. Restrictive energy supply and progressing gravidity decreased energy content in the weight gain. The influences mentioned were taken into consideration on the derivation of the partial requirement for live weight gain. Equations were developed for the estimation of the energy requirement of young female cattle, which can be applied to both gravid and non-gravid cattle.     

Growth studies of bulls of black-and-white dairy cattle during low feed levels. 4. Derivation of energy requirements

On the basis of two long-term individual feeding experiments the growth-related energy requirement values for low live weight gains (500, 600 and 700 g daily cumulative live weight gain per animal derived by Pieper et al. in 1984 (1984b) were ascertained more precisely. The differences from the recommendations of the GDR feed evaluation system found earlier (Pieper et al. 1984) were confirmed. In general one can say that the differentiation of energy requirement in dependence of live weight gain with live weight being equal is greater than according to the GDR feed evaluation system (1986). Under consideration of a norm factor of 1.05 this results in the live weight range of between 50 and 500 kg and with a cumulative live weight gain of 500 g/animal and day in an energy expenditure of 4.44 kEFUcattle/kg live weight gain, of 600 g/animal and day i an energy expenditure of 4.08 kEFUcattle/kg live weight gain and of 700 g/animal and day in an energy expenditure of 3.82 kEFUcattle/ kg live weight gain.     

Joint research on the precise determination of the energy and protein requirements of fattening pigs. 2. Energy and nitrogen metabolism of fattening hybrids in the fattening range of 30 to 120 kg

In 3 experiments a total of 242 total metabolism experiments with ad libitum feeding (experiment 1), 75% (experiment 2) and 62% (experiment 3) of the energy level of the 1st experiment and approximately equal protein and amino acid doses in experiments 1-3 were carried out with 8 castrated male fattening hybrids each (large white X land race pig) X line 150) in the live weight range between 30 and 120 kg. On average, feed intake over the whole live weight range was 2.24; 1.79 and 1.50 kg/animal and day, the corresponding daily live weight gain was 729, 533 and 396 g. With regard to the digestibility of the energy and the nutrients and the metabolisable energy in % of the gross energy there was no relation to the development of the animals. The intake of metabolisable energy per kg live weight decreased with ad libitum feeding and with the advancing development at the end of the experiment in contrast to the beginning of the experiment to 53%, energy retention to 56%. The utilisation of metabolisable energy for body energy retention, taking account of a maintenance requirement of 450 kJ/kg live weight 0,62 on the average of the 3 experiments was 68.4 +/- 1.9, 70.3 +/- 2.0 and 64.3 +/- 2.6%. Energy retention in experiments 1 to 3 amounted to 8.6, 6.8 and 5.3 MJ at the beginning and to 18.1, 12.2 and 8.0 MJ per animal and day at the end of the experiment. Protein energy retention of the pigs (live weight 40 kg) was 26% of the total energy retention in experiments 1 and 2 and 49% in experiment 3. In experiments 1 and 3 protein retention decreased to 15% of the total energy retention, in experiment 2 protein retention remained constantly at 22% between 60 and 110 kg live weight and then decreased to 18%. Consequently, the N-balances were 23-16 g, 16-20 g and 16-9 g/animal and day. The chemical composition of the carcasses was strongly influenced by the level of nutrition. At the beginning of the experiment the protein content averaged between 49 and 57% and the total fat content between 31 and 38% of the dry matter. At the end of the experiment the carcasses of the animals from experiments 1-3 had crude protein contents of 28, 33 and 42% and total fat contents of 66, 61 and 50% of the dry matter.     

Energy requirement of young female cattle. 3. Reproduction and products of conception

The results with regard to reproduction obtained from a study lasting several years of the energy requirement of young female cattle reared with different intensity are described. It becomes apparent that there are relations between feeding, rearing intensity and insemination expenditure. Age and weight at the time of conception in the individual test series averaged between 290 and 630 days and 270 and 430 kg respectively. The weight of the conceptional products of 31 calvings was registered and, according to Jakobsen (1956, 1957), the energy and N content of the conceptional products was proportionally assigned to the prenatal test periods. In the last 140 days of gravidity the N balance per animal and day increases from 16 to 23 g. N retention of the cows is low in the last days of gravidity. The utilization of metabolizable energy for the development of conceptional products is calculated as a conventional comparative value provided the maintenance requirement per unit metabolic body weight of the cows and the utilization of metabolizable energy for energy retention in the cows on an average of 156 individual values of 12 +/- 8% are the same and correspond in their magnitude to analogous findings of other authors, mainly compiled from dairy cows.     

The energy requirements of Holstein-Friesian heifers during rearing and pregnancy under the tropical conditions of Cuba. 1. Determination of dry matter and energy assimilation and of the live weight development and energy expenditure

In an individual feeding experiment with 48 Holstein-Friesian heifers divided into 3 groups, the effect of various energy levels on live weight development and energy expenditure in the age bracket of 97 days (78 kg live weight) to conception (350 kg LW) and from conception to calving (500 kg LW) was investigated. Rations with differing concentrate--roughage ratios were used. The differentiation of energy intake per day between groups 1 and 2 and group 3 amounted to 7.5% in the 1st development period and to 20% between group 1 and groups 2 and 3 in the 2nd development period. There were significant differences in the live weight gain of 95 g/animal and day in the first development period (up to conception) and of 81 g/animal and day in the second period of development (from conception to calving). Under the tropical climatic conditions the energy expenditure in these studies was approximately 10% under the energy expenditure recommended by the NRC (1978). While there was a clear reciprocal relation between energy expenditure and live weight gain in the first period of development, this relation was less distinct in the second period of development.     

The rearing of female twin cattle using only feed pellets in comparison with straw pellets plus a concentrate mixture. 2. Weight gain and feed consumption as well as rumen fermentation in the 1st half of the 2d year of life

In investigations with 30 young cows (twin cattle) the applicability of straw materials as sole roughage was tested during the first half of their second year of life (183 feeding days). In the course of the experiment sole feed pellets (I) and partly pelleted rations of dried feed (straw pellets plus loose wheat and loose wheat straw resp.; II) were used. Approximately 72% of the dry matter intake was provided from straw materials in both groups and on an average of the test period. Weight gain and feed expenditure were ascertained and at the end of the test period also the parameters of rumen fermentation and of the concentration of some metabolism parameters in the blood serum. With feed supply being restrictive (2.2 kg dry matter/100 kg live weight), a high dry matter intake from straw (1.6 kg/100 kg live weight or 4.8 kg/animal and day resp.) and average weight gains of between 574 (I) and 582 g (II) were achieved. The standard expenditure of energy (kEFUcattle) and protein stated in the GDR feed evaluation system for this period of development was undercut by 27 (II) and 29 (I)% and 12 (II) and 18 (I)% resp. On average the expenditure of kEFUcattle/kg weight gain was 4.75 (I) and 4.98 (II) and that of digestible crude protein/kg weight gain 679 (I) and 733 g (II). Compared to standard values the expenditure of energy and feed expenditure from concentrates was 80%. Rumen fermentation conditions corresponded to those of a high cellulolytic activity (C2: C3 ratio of greater than 4: 1). The only significant differences between the groups at the time of sampling were to be observed in the total concentration of volatile fatty acids and in the pH-value in the rumen. The absolute (1/animal) and relative (1/kg dry matter intake) volume of rumen fluid varied between 65 and 70 and between 10.5 and 12.5 1. The ascertained concentration of selected metabolism parameters were within the physiologic standard range.     

Rearing of female cattle twins using complete feed pellets in comparison to straw pellets plus concentrate mixture. 1. Weight gain performance and feed expenditure as well as rumen fermentation conditions during the 4th-12th months of life

The investigations with young female cattle (twin cattle, n = 30) the applicability of straw as sole roughage after the end of the suckling period was tested. Completely (I) and partly (II) pelleted rations of dried feed with a straw component of 65% each in the total rations were used. Feed intake, development of body weight, feed expenditure as well as indexes of rumen fermentation and of the concentration of some metabolism parameters in the blood serum were measured. In the complete test period (between 5th and 12th months of life, 243 days) there were not any significant differences between the groups in all registrations. Average daily weight gains of 570 g were achieved with an overall high dry matter (2.6 kg/100 kg live weight) and roughage (straw) intake (1.6 kg dry matter/100 kg live weight). The standard expenditure of energy and protein given for the stage of development was differentiatedly undercut. The energy expenditure being below the standard (ca. 10%), the total energy expenditure (kEFUcattle) was by 20 (II) and 25% (I) below standard expenditure. Energy expenditure aimed at weight gain reached very satisfactory values with 3.68 (I) and 3.79 kEFUcattle/kg weight gain (II). Protein expenditure fell short of the standard for the development stage by 14 (II) and 20% (I) resp. All parameters ascertained of the rumen and intermediate metabolism were in the physiologic standard range, as is characteristic of feeding regimes with a relatively high quota of roughage.     

生长期秦川牛能量代谢规律与需要量研究

本试验旨在研究生长期秦川牛能量代谢规律与需要量。选择30头体况良好、体重[(336.33±18.28)kg]相近的生长期秦川牛公牛,随机分为5组,每组6头牛,分别饲喂按我国《肉牛饲养标准》(NY/T 815—2004)提供的预期平均日增重900 g/d所需净能的85.0%(Ⅰ组)、92.5%(Ⅱ组)、100.0%(Ⅲ组)、107.5%(Ⅳ组)、115.0%(Ⅴ组)配制的5种试验饲粮。采用饲养试验和消化代谢试验测定秦川牛生长性能及能量代谢指标,并建立消化能和代谢能需要量预测模型。预试期10 d,正试期42 d。结果表明,Ⅲ组秦川牛平均日增重为880.15 g/d,较预期的结果略低;Ⅳ组平均日增重达到最大值(1 160.10 g/d),能量利用效率最高;总能消化率、总能代谢率和消化能代谢率平均值分别为(76.44±3.23)%、(66.75±3.16)%、(87.31±0.54)%;秦川牛的消化能和代谢能需要量的回归方程分别为:DER=0.778W0.75+37.05ADG;MER=0.668W0.75+33.49ADG[DER为消化能需要量(M J/d),MER为代谢能需要量(M J/d),W0.75为单位代谢体重(kg),ADG为平均日增重(kg/d)]。综合得出,生长期秦川牛的维持消化能和代谢能需要量分别为0.778、0.668 MJ/(kg W0.75·d),每千克增重的消化能和代谢能需要量分别为37.05、33.49 M J。     

Energy and protein concentration of broiler rations in Cuba. 2. Female broilers in winter

Four energy concentrations (10.5, 11.5, 12.6 and 13.6 MJ ME) and four protein concentrations (15.0, 17.5, 20.0 and 22.5% CP) were tested with a total of 1,900 female broilers of the breed White Plymouth Rock in their first 56 days of life in 16 experiment groups. The animals were kept in cages and were directly exposed to the natural Cuban winter temperatures from their 15th day of life. As six repetitions were carried out in every test group, the results could be variance-analytically calculated. In winter the female broilers of the breed White Plymouth showed a lower intake of feed, metabolic energy and crude protein than male animals. They achieved a lower live weight, lower amount of body protein and fat and worse feed, protein and energy utilisation. Possibilities are shown to meet the crude protein and energy requirement in monophase feeding systems with 11.5 MJ ME and 20% CP or with 12.6 MJ ME and 17.5 or 20% CP.     

The effect of live weight gain and live weight loss on body composition of Merino wethers: dissected muscle, fat, and bone.

Sheep were slaughtered after live weight gain (LWG) and live weight loss (LWL) to evaluate the effect of different periods of normal and retarded growth on dissected carcass components. Thirty-five Merino wether sheep were allowed ad libitum access to the experimental diet of 80% alfalfa chaff and 20% cereal grain (17.23% CP and 18.10 MJ/kg of GE) to grow from 23.0 to 33.0 kg live weight and then fed to lose 10 kg at 133 g/d. Five animals were slaughtered at live weights of 23.0, 26.3, 29.6, and 33.0 kg during LWG and 29.6, 26.3, and 23.0 kg during LWL. Fleece-free empty BW was similar in both LWG and LWL animals at all common slaughter weights because of the lesser and greater weights of the alimentary tract contents and the fleece, respectively, in the LWL animals. Carcass weight or dissected side weight was greater in LWL animals at each common slaughter weight but significantly so (P less than .05) at the 23.0 kg live weight only. Total side muscle weight was similar in LWG and LWL animals at each common slaughter weight, but the proportion of muscle in the dissected side weight was lower (P less than .05) in LWL animals at 23.0 kg live weight. Total side fat, subcutaneous fat, intermuscular fat, and kidney and channel fat weights were all greater in LWL animals at each common slaughter weight, but significantly so (P less than .05) at the 23.0 kg live weight only. Total side bone weight was higher at both 26.3 kg (P less than .01) and 23.0 kg (P less than .05) in the LWL animals. Weight loss in young sheep mobilized more weight of muscle than fat from the carcass, whereas bone weight was constant.     

Energy and protein concentration of broiler rations in Cuba. 4. Female broilers in the summer

A total of 1,900 female broilers of the variety White Plymouth Rock was tested with 4 energy concentrations (10.5, 11.5, 12.6 and 13.6 MJ metabolizable energy/kg feed) and 4 protein concentrations (15.0, 17.5, 20.0 and 22.5% CP) in their first 56 days of life in 16 test groups. The animals were kept in cages and were directly exposed to the natural Cuban summer temperatures from their 15th day of life onwards. As every test group was run with 6 repetitions, the results could be variance analytically calculated. In a comparison of male and female broilers in summer one can state that the female animals achieved a lower intake of feed, energy and crude protein as well as a lower live weight than the male animals. In winter the female broilers consumed between 15.7 and 36.4% more feed and achieved a live weight gain between 11.1 and 43% higher than in summer. It could be shown that the maximum live weight of the female broilers could be achieved in summer if rations containing 12.6 MJ metabolizable energy and 20% CP are fed in a one-phase feeding system.     

Feed intake of young double-muscled bulls fed on grass and supplemented with sugar-beet pulp.

Daily dry matter intake in young growing double-muscled bulls, fed indoors on grass, was estimated based on forty-four intake data from 28 animals, ageing at least five months and weighing up to 400 kg live weight. Intake was measured during five consecutive days using one of eighteen cuts of grass. Fresh meadow grass (mainly Lolium perenne) was fed ad libitum and two kg dried sugar-beet pulp was offered per animal and per day. Animal live weight averaged 278 +/- 82 kg and mean total daily dry matter intake amounted to 5.05 +/- 1.59 kg or 73.6 +/- 13.7 g per kg metabolic weight, while pulp dry matter intake amounted to 1.49 +/- 0.50 kg per day. Regression analysis showed that animal as well as feed characteristics could explain up to approximately 90% of the variation in daily dry matter intake. The supplementation resulted in an extra daily dry matter intake of 0.68 g per g pulp dry matter. Intake of double-muscled animals was considerably lower than previously reported for non-double-muscled cattle. An extra supplementation of young grazing double-muscled animals could be advised from these findings, while extra protein should also be considered.     

育肥后期锦江牛能量代谢规律及需要量的研究

本试验旨在研究育肥后期锦江去势公牛的能量代谢规律及需要量。在育肥前期不同能量水平饲喂的基础上(育肥前期5种肉牛日粮综合净能(NE_mf)依次为6.02、6.38、6.74、7.10、7.46 MJ/kg,每组10头牛,育肥116 d)继续育肥,育肥前期试验结束后,保持分组不变,挑选35头体型接近、体重(355.94±35.11) kg锦江牛继续育肥。按照中国《肉牛饲养标准》(NY/T 815-2004)中350 kg肉牛日增重1.2 kg/d所需净能的100%(A组)、106%(B组)、112%(C组)、118%(D组)、124%(E组)配制5种不同能量水平的试验日粮,5种日粮的NE_mf依次为6.21、6.58、6.95、7.33、7.70 MJ/kg。采用饲养试验和消化代谢试验测定育肥后期锦江去势公牛生长性能及能量代谢指标,并建立消化能和代谢能能量需要模型。预试期10 d,正试期128 d。结果表明:①D、E组育肥后期锦江去势公牛的总能采食量较其他组显著降低(P<0.05)。②B组肥后期锦江去势公牛能量利用效率最高,总能消化率、总能代谢率分别为90.59%和83.36%。③育肥后期锦江去势公牛日增重与消化能采食量和代谢能采食量存在高度线性正相关(R^2=0.997、R^2=0.993),其消化能、代谢能需要量的回归方程分别为:DE_m=0.770W^0.75+40.088×ADG;ME_m=0.645W^0.75+38.603×ADG(其中DE为消化能总需要量(MJ/d);ME为代谢能总需要量(MJ/d);W^0.75为单位代谢体重(kg);ADG为平均日增重(kg/d))。综上所述,育肥后期锦江牛的维持消化能总需要量(DE_m)和代谢能总需要量(ME_m)分别为0.770、0.645 MJ/(kg W^0.75·d),每千克增重的消化能和代谢能需要量分别为40.088、38.603 MJ。     

The energy metabolism of growing swine in the body weight range of 10-50 kg. 2. Nitrogen and energy metabolism

Investigations were carried out about nitrogen and energy metabolism feeding rations with 17-24 and 44-47% crude protein content on maintenance and growing level to castrated male hybrid pigs of line 150. In growing periods the N deposition amounted to 10 g/animal.d (15 kg live weight), 18 g/animal.d (30 kg LW) and 21 g/animal.d (greater than 40 kg LW) on lower protein feeding level. In experiments with higher protein feeding level the corresponding results were 17, 22 and 22 g N deposition/animal.d. The partial utilization of metabolizable energy for deposition amounted to 70% for the rations with 17-24% protein content and to 59% for the rations with 44-47% protein content, without correlation to the animals development and the alternation in the protein feeding level. The results of regression analysis about maintenance requirement were 814, 775 and 806 kJ metabolizable energy/kg LW0.62.d in trials feeding rations with 17-24% crude protein content as well as 764, 846 and 818 kJ metabolizable energy/kg LW0,62.d in trials feeding rations with 44-47% crude protein content. 1,5-1,8 MJ metabolizable energy were used per MJ protein energy deposition and 1,3-1,4 MJ per MJ fat energy deposition respectively. The energy deposition per kg live weight gain amounted to values between 9 (10 kg LW) and 18 MJ (60 kg LW).     

Energy intake of adult Great Danes]

Energy intake and weight development were recorded in 5 adult Great Danes kept in outdoor kennels during maintenance, pregnancy and lactation. In periods with constant weights the energy requirement was 0.96 MJ DE/kg BW0.75/d with a minimum in summer (0.82) and a maximum in winter (1.1). In pregnancy (weight gain depending on litter size 7-15 kg) energy intake (ad-lib.-feeding) was similar to maintenance. During lactation energy intake reached 1.6-1.7 MJ DE/kg BW0.75/d with a dry matter intake of 83 g/kg BW0.75/d. Pups of great litters had to be fed with milk substitute for normal weight development.     

Nutrient and energy retention in growing lambs. 1. Results of growth experiments

The results of a growth experiment are explained which was carried out as basis for the control slaughtering method and for N-balance experiments with hybrid fattening lambs of both sexes and of three different genetic constructions in comparison with male merino mutton lambs. The animals were kept individually on perforated floor and received pelleted fattening feed for lambs ad libitum. In the live weight range between 15 and 30 kg the male hybrid lambs reached significantly higher live weight gains than the merino mutton sheep, the former tending towards higher feed intake and lower energy expenditure. The male hybrid lambs showed a significantly higher live weight gain and a significantly lower energy expenditure than the female animals of equal live weight whereas there were no differences between the sexes in feed intake.     

Energy metabolism of growing broiler chickens kept in groups in relation to the environmental temperature. 1. Feed intake, heat production and energy utilization]

In 5 experiments with young broiler chickens kept in groups at environmental temperatures (ET) of 35, 30, 25, 20 and 15 degrees C feed intake, live weight development and heat production were ascertained between their 5th and 57th days of life. Feed intake and live weight were ascertained separately according to sex. With live weight being the same, feed intake increased with decreasing ET. There was no maximum feed intake up to the 57th day of life at between 20 and 25 degrees C. At the end of their 57th day of life the male and the female chickens had achieved the following live weights (in the sequence of decreasing ET 35-15 degrees C): 1342; 2014; 2829; 2946; 2374 and 1367; 1900; 2512; 2496 and 2200 g/animal resp. The highest live weight gain of the chickens was achieved at between 25 and 20 degrees C with 60-70 g/d for male and with 50-60 g/d for female animals. Heat production (HP) increased progressively with age. The highest HP was registered at 20 degrees C ET.     

Urea utilization of growing lambs. 2. Growth experiments

In two growth experiments with 30 lambs per group the influence of urea (1-2% of the ration) and straw (20-30% of the ration) on the performance was studied under conditions of intensive fattening. The content of native crude protein in the rations varied between 10.1 and 18.4% of the DM. In experiment 1 the lambs of the two groups which received mixed feed for fattening lambs without urea on average consumed with 1.02 and 1.11 kg DM resp., 696 and 757 EFUcattle resp. and 173 and 189 g crude protein resp. per animal and day. The groups with wheat or barley and urea consumed 1.06 and 0.96 kg DM resp., 714 and 627 EFUcattle resp. and 209 and 155 g crude protein resp. Their weight gain of 247 g/animal and day was 24% and that of 230 g resp. 26% higher than that in the control groups without urea. The lowest dry matter intake (0.91 and 0.82 kg resp.) was shown by the animals of the groups which received wheat or barley without urea. In the second experiment the control group achieved an average daily weight gain of 327 g at an expenditure of 2.24 kEFUcattle/kg weight gain. Despite a partly higher dry matter intake, the daily EFUcattle intake was between 12.2 and 24.5% lower in the straw groups and energy expenditure increased between 12.0 and 27.8%. Urea supplements of between 1 and 2% improved dry matter, EFUcattle and crude protein intake and thus weight gain.     

Growth studies in bulls of the black-and-white dairy cattle breed at a low feeding level. 1. Growth progress and fattening performance]

In 2 experiments with a total of 108 black-and-white dairy bulls feed intake, growth performance and energy expenditure for live weight and carcass production with low feeding intensity were ascertained. The performance level varied between 600 and 650 g; 24-month-old bulls achieved a final weight of 458 and 517 kg resp. Growth and retention performance were considerably influenced by the energy concentration of the ration and by daily energy intake.     

20～50 kg川藏黑猪的能量代谢与沉积规律研究

本试验旨在研究20~50 kg川藏黑猪的能量代谢与沉积规律。试验一:选择体重接近[(20.17±3.46)kg]的川藏黑猪配套系商品猪64头,预试7 d后屠宰4头猪测定胴体成分;其余试验猪按公母随机分成5个组,每个组4个重复,每个重复3头,各重复单圈饲养,自由采食消化能(DE)水平分别为13.79、13.37、12.96、12.54和12.12 MJ/kg的饲粮。试验猪体重达50 kg时,结束试验一,测定平均日采食量、平均日增重和料重比,并在各组选择1头猪屠宰测定胴体成分。试验二:选择试验一中15头体重接近[(48.34±4.07)kg]的公猪,随机分成5个组,每个组3个重复,每个重复1头,单独饲养于代谢笼,分别采食上述5种DE水平的饲粮,进行消化代谢试验。预试期3 d,正试期4 d。采用析因法建立能量需要量预测模型。结果表明:饲粮DE水平影响了20~50 kg川藏黑猪的平均日采食量、平均日增重和料重比;饲粮DE转化为代谢能(ME)效率(ME/DE)为97.26%~98.10%,ME用于沉积产品能的效率(DED/ME)平均值为41.71%;此阶段川藏黑猪维持需要ME平均值为0.49 MJ/W~(0.75)或0.85 MJ/W~(0.60)(按DE计为0.50 MJ/W~(0.75)或0.87 MJ/W~(0.60)),增重需要DE和ME平均值分别为18.91和18.47 MJ/kg。由此得出,20~50 kg阶段川藏黑猪能量需要模型为:DE(MJ/d)=0.504 W~(0.75)+18.91ΔW或DE(MJ/d)=0.867 W~(0.60)+18.91ΔW;ME(MJ/d)=0.492 W~(0.75)+18.47ΔW或ME(MJ/d)=