Modeling the edge effect in even-aged Monterrey pine (Pinus radiata D. Don) stands incorporating a competition index

The edge effect on DBH (diameter at breast height) in two adjacent Pinus radiata D. Don stands was modeled as a function of both the distance from the stand edge and Hegyi's competition index. The modeling was applied to edges with external-competition pressure and on free edges. In both edge types, the incorporation of the Hegyi's competition index in the modeling significantly reduced the residual variation. This modeling showed that the intensity of the intraspecific competition varies between stands. Particularly, the trees in younger stands are more sensitive to the effects of competition than in older stands. However, the intensity of the competition does not vary between edge aspects of stands. The results of the study related to the depth of edge effect can be useful for stratifying stands when getting more precise estimates of trees and stand parameters from sampling are required, and for defining the width of the buffer strip in experimental units used for long-term field assays.     

Effects of competition on growth of Amur linden plantation

IntroductionCompetitionorinterf6renceisimportantinfOrestgrowthandstandstructure(Weiner1982,1984).ForexampIe,thesizeofindividuaIcrownsanddiametersisaffectedbycompetition.Mbstint6rferencestudieshavefocusedonthemeanperformance0fplantsinreIationtodensity(Harper1977)andneighbouringdistance(Martinetal.1977;Weiner1984,Bigingeta[1992).Theyuseddistance-dependentcompetitionindices(Cl)todescribeinfluencefOrindividuaItreesthatincIudesaIIcompetitorsinthezoneofinfIuenceandprovidebettercorrelationswit…     

Crown radius of pedunculate oak (Quercus robur L.) depending on stem size,stand density and site productivity

Crown size is a good indicator of the growth potential of trees and is often used in forest management for outlining thinning guidelines or constructing forest growth models. The aim of this study was to analyse mean crown radius as a function of stem size, stand density and site productivity in even-aged stands of pedunculate oak (Quercus robur L.). Data included measurements of 620 trees from 53 plots in nine thinning experiments and one operational stand in Sweden, Denmark and Great Britain, representing a wide spectrum of thinning practices ranging from the strictly unthinned control to extremely heavy thinning with essentially solitary trees. Three sets of models were constructed based on different predictor variables, including indicators of individual stem size (diameter at breast height, DBH), stand density/thinning grade (quadratic mean diameter and stand basal area) and site productivity (stand top height). Preliminary results indicated a significant effect of DBH and (nominal) thinning grade on crown radius. The response pattern of the final models indicated an increasing crown radius with increasing DBH, with increasing thinning grade (decreasing stand density) and with decreasing site productivity. The models are valid for predicting the crown radius of pedunculate oak in even-aged forest stands.     

基于树冠因子的林木竞争指数研究

[目的]以湖南省黄丰桥国有林场杉木纯林为研究对象,进行树木竞争指数的研究,基于Hegyi简单竞争指数的形式,提出包含反映树冠特征因子的树冠竞争指数。[方法]将树冠竞争指数和Hegyi简单竞争指数分别与胸径增长量进行相关分析。[结果]Kendall's tau-b系数、Spearman's rho系数和相关系数R²均显示树冠竞争指数与林木生长量的相关性高于Hegyi简单竞争指数。[结论]该结果表明树冠竞争指数与林木生长量有更强的相关性,可以更好地体现树冠在树木生长中的作用,能作为模拟林分生长量的依据。     

A basal area increment model for individual trees in mixed continuous cover forests in Iranian Caspian forests

The aim of this study was to develop and test a new basal area growth model in mixed species continuous cover forests in northern Iran.Weanalyzed 421 core samples from 6 main species in the forest area to develop our growth model.In each plot,we measured variables such as total tree height(m),diameter at breast height(DBH)(cm)and basal area of larger trees as cumulative basal areas of trees(GCUM)ofDBH[5 cm.The empirical data were analyzed using regression analysis.There was a statistically significant nonlinear function between the annual basal area increment,as the dependent variable,and the basal area of the individual trees and competition as explanatory variables.Reference area from the largest trees,was circular plot with area of 0.1 ha.GCUM was estimated for trees of DBH>5 cm.Furthermore,we investigated the dependencies of diameter growth of different species on stand density at different levels of competition,and diameter development of individual trees through time.The results indicate that competition caused by larger neighborhood trees has a negative effect on growth.In addition,the maximum diameter increment is affected by competition level.Therefore,the maximum diameter increment of species occurs when the trees are about 35–40 cm in dense-forest(40 to 0 m^2 per ha)and when the trees are about 60 to 70 cm in very dense forest(60 to 0 m^2 per ha)which is more likely to Caspian natural forests with high level density due to uneven-aged composition of stands.     

Individual tree growth response to variable-density thinning in coastal Pacific Northwest forests

We examined 5-year basal area growth of nearly 2600 trees in stem-mapped plots at five locations differing in site characteristics, species composition, and management history on the Olympic Peninsula in Western Washington, USA. Our objectives were to determine if internal edges, the boundaries within the stand between components of the variable-density thinning, influenced individual tree growth, and whether incorporation of individual tree local competition indices in growth prediction models could account for treatment and edge effects. Treatment significantly affected tree growth at all sites, with trees in the thinned matrix displaying on average over 25% greater basal area growth than trees in unthinned patches. Proximity to canopy gaps created as part of the variable-density thinning increased basal area growth of trees in the thinned matrix by nearly 11%. In addition, growth of trees close to skid trails was 11% greater than trees located away from the trails. Past thinning history, and its effect on initial stocking rate, appeared to affect the magnitude of the edge effects. Blocks that had received earlier commercial thinnings, and thus had lower stocking at the onset of the study, displayed lower growth responses than previously unthinned blocks. Including local competition indices in the models generally reduced growth prediction error; however, the indices examined did not fully account for treatment or edge effects. Our results suggest that not accounting for internal edges in spatially complex stands could result in errors in projected growth of trees, although these edge effects are highly variable. Failure to account for the effects of internal edges could affect not just estimates of future stand yield, but also projections of future stand structure.     

桂西南马尾松人工林生长对不同强度采伐的动态响应

[目的]研究不同强度采伐下马尾松的生长动态,筛选适宜的采伐强度,为马尾松人工林近自然经营提供技术支撑。[方法]2007年10月在14年生马尾松人工林(保存密度1 100株·hm~(-2))内进行采伐试验,设置4个采伐强度,即保留密度分别为225、300、375、450株·hm~(-2),以不采伐为对照;其后,自2008年开始连续8 a,每2 a测定1次马尾松的胸径、树高、枝下高和冠面积等生长指标,并计算单株材积和林分蓄积量,应用方差分析和Duncan多重比较分析生长指标对不同采伐强度的动态响应。[结果]表明:采伐强度显著影响林分生长,其中,林分平均胸径、单株材积、冠面积的年均增长量随保留密度增大而减小,但均显著高于对照(P0.05)。采伐后第1 3年,马尾松冠面积增长量显著高于采伐后期,胸径则在采伐后第3 5年最高,而不同采伐强度对林分树高生长影响不明显。保留密度显著影响林分枝下高和蓄积量的动态变化,其年均增长量随密度增大而递增。5个处理间林分蓄积年均增长量的差异随林龄的增大而逐渐缩小。[结论]马尾松人工林生长对不同强度采伐的动态响应以树冠最敏感,冠面积首先陡然增大,进而引起胸径的快速生长。树高和枝下高在采伐后年均增量变化相对平稳。4个采伐强度均显著促进单株材积生长,而仅保留密度为225株·hm~(-2)的采伐对林分蓄积增长量影响显著。综合比较林分的单株材积和林分蓄积连年增长量,建议在桂西南15年生马尾松人工林近自然经营中宜选择300株·hm~(-2)的保留密度进行采伐。     

Snow damages on trees of an uneven age in mixed broadleaf forests:effects of topographical conditions and tree characteristics

Snow damage on natural stands is an important problem concerning mountainous forest management.In the present research,the frequency and intensity of heavy snow damage on natural stands and the relationship of damages with characteristics of trees,stand and topography were studied in mountainous forests of northern Iran.A systematic sampling design was applied to the study area(140 ha),with 122 circular plots.The grid dimension was100 m and each plot area was 1000 m^2.The four types of snow damage to trees include:crown damage(8.6%),stem breakage(5.4%),uprooting(3.2%),and bending(1.4%).The frequency of crown damage grew with an increase inthe diameter at breast height(DBH),while the frequency of stem breakage,uprooting and bending decreased with an increase the DBH.The frequency of crown damage,stem breakage,and total damages was related to tree species(p<0.01).Not all tree species were equally susceptible to snow damage.The amount of damage grew with increasing elevation and slope angle and decreased with increasing soil depth.Young trees(DBH<25 cm)were more susceptible to snow damage than other trees.Snow damage decreased by as the trees in the stand became more mixed in age.The susceptibility of trees to snow damage increased by height and slenderness coefficient.With adequate silvicultural operation,snow damage can be reduced to a minimum level in these natural forests.     

日本落叶松人工林密度调控技术研究

以日本落叶松中幼林抚育间伐样地监测数据,利用树冠竞争因子（CCF）、林分密度指数、单位断面积生长过程等指标,分析了抚育间伐对断面积和蓄积两方面的影响,结果表明：间伐林分的断面积和蓄积有相同的生长过程,即林分在一定间伐强度范围内（强度小于50％）间伐强度越大,年均生长率越大;用树冠竞争因子描述日本落叶松人工林断面积的生长过程,未间伐林分的立木度接近稳定状态,而间伐林分降低了林分的CCF值,随着林龄的增加,CCF值不断增加,逐渐向未间伐林分靠近。总的来说间伐林分在间伐初期要小于未间伐林分,但随着林龄的增大及林木竞争的加剧,间伐林分逐渐向未间伐林分靠近。     

毛竹混交林林分生长状况和结构特征研究

对福建省建阳交溪毛竹阔叶树混交林、毛竹杉木混交林、毛竹杉木阔叶树混交林及毛竹纯林等6种不同混交模式林分生长状况进行比较,结果表明:混交林毛竹个体生物量、毛竹胸径和竹高较大,不同模式林分间的胸径、竹高、冠幅等生长因子均达显著差异;竹阔混交林较有利于竹鞭孕芽发笋;各模式竹林结构综合评判为一般经营竹林;竹林竹鞭长、鞭径、鞭节生长以竹阔比例为5:5林分竹鞭较好。     

雷州尾叶桉树冠指标与抚育间伐关系的探讨

通过对雷州尾叶桉树冠有效受光面积、树冠比率、树冠投影面积进行调查研究,分别用树冠有效受光面积、树冠比率、投影面积与胸径建立一元幂函数回归方程进行研究与分析,结果表明树冠有效受光面积公式对指导培育中径材较好;树冠比率与树冠投影面积公式的回归效果基本一致,对指导培育小径材较为理想,并利用上述回归公式编制单株营养面积、公顷立木株数和公顷基面积表为尾叶桉人工林的抚育间伐设计与施工提供理论依据。     

Canopy development in young sugi (Cryptomeria japonica) stands in relation to changes with age in crown morphology and structure

In young sugi (Cryptomeria japonica D. Don) stands, crown shape (crown length/crown diameter) ratio, average branch inclination, and spatial density of foliage in the crown increased with stand age. Within crowns, foliage distribution increased from the apex downward and, until crown closure, reached a maximum near the crown base. After crown closure, the maximum occurred near the middle of the crown. In each stand, foliage distribution in the canopy showed almost the same vertical change over time as it did in individual crowns. The vertical distribution of foliage in the canopy moved upward with stand age, accompanied by an increase in canopy depth and leaf mass. The shape of the vertical distribution was almost symmetrical between the upper and lower halves in the closed stands, although slightly skewed downward. The logarithm of average spatial density decreased linearly as cumulative leaf mass increased with distance from the top of the canopy. The total cross-sectional area of the crowns exceeded the stand area from the middle of the canopy downward in the closed stands because of crown overlap. However, partly because of changes in crown morphology and structure, the increase in leaf mass with stand age did not always cause more severe crown competition.     

Determination of number of stems in coniferous forest stands by means of aerial photo‐interpretation

Two models for determination of the number of stems per hectare in forest stands (N) from attributes derived by aerial photo‐interpretation were developed. The models relied on the assumption that N could be determined by dividing the total stand volume per hectare with the volume of the “average tree”; defined by stand mean height and the diameter corresponding to mean basal area of a stand. Input variables of the models were stand mean height, crown closure and site quality. Additionally, model II required input of average stand volume per hectare and average mean diameter derived from stratified field sample plot inventories. Material for 143 coniferous stands was used for the testing of the models. The stands were recorded by intensive field measurements. Aerial photographs at the approximate scale of 1:15 000 were used for photo‐interpretation. The N value was underestimated in model I by 5.4–47.0%. The standard deviation for the differences was 15.2–26.2% for mature stands and 41.4–44.2% for young thinning phase stands. For model II, the mean difference between the predicted and observed N value was in the range ‐16.1% to 12.2%.     

造林技术措施对10年生马尾松幼林生长的影响

在处台江县革东镇设置了反映马尾松人工造林各主要技术环节的傈,在对试验林固定样地进行定期、定位测定的基础上,采用数量化理论１方法分析研究了不同造林技术措施对马尾松幼林生长的影响。结果表明,主要造林技术措施对马尾松幼林生长自大 的影响顺序为：立地选择,种要密度,幼林施工。其中立地选择、种源选择和造林密度是马尾松人工造林最关键的技术环节,幼林施肥对马尾松幼林分的生长有一定影响,整地方式则无明显影响,运用     

Individual-based measurement and analysis of root system development: case studies for <Emphasis Type="Italic">Larix gmelinii</Emphasis> trees growing on the permafrost region in Siberia

We present results of individual-based root system measurement and analysis applied for Larix gmelinii trees growing on the continuous permafrost region of central Siberia. The data of root excavation taken from the three stands were used for the analyses; young (26 years old), mature (105 years old), and uneven-aged over-mature stand (220 years old). In this article, we highlight two topics: (1) factors affecting spatio-temporal pattern of root system development, and (2) interactions between aboveground (i.e., crown) and belowground (i.e., root) competition. For the first topic, the detailed observation of lateral roots was applied to one sample tree of the overmature stand. The tree constructed a superficial (<30 cm in depth) and rather asymmetric root system, and each lateral root expanded mainly into elevated mounds rather than depressed troughs. This indicated that spatial development of an individual root system was largely affected by microtopography (i.e., earth hummocks). For these lateral roots, elongation growth curves were reconstructed using annual-ring data, and annual growth rates and patterns were compared among them. The comparison suggested that temporal root system development is associated with differences in carbon allocation among the lateral roots. For the second topic, we examined relationships between individual crown projection area (CA) and horizontal rooting area (RA) for the sample trees of each stand. RA was almost equal to CA in the young stand, while RA was much larger (three or four times) than CA in the mature and overmature stands. Two measures of stand-level space occupation, crown area index (aboveground: CAI; sum of CAs per unit land area) and rooting area index (belowground: RAI; sum of RAs), were estimated in each stand. The estimates of RAI (1.3–1.8 m² m₋₂) exceeded unity in all stands. In contrast, CAI exceeded unity (1.3 m² m₋₂) only in the young stand, and was much smaller (<0.3 m² m₋₂) in the two older stands. These between-stand differences in RAI–CAI relationships suggest that intertree competition for both aboveground and belowground spaces occurred in the young stand, but only belowground competition still occurred in the two older stands. Based on this finding, we hypothesized that competition below the ground may become predominant as a stand ages in L. gmelinii forests. Methodological limitations of our analysis are also discussed, especially for the analysis using the two indices of space occupation (CAI, RAI).     

内蒙古大青山前坡油松人工林生长特征研究

以内蒙古大青山前坡38 a生、17 a生油松人工林为研究对象,对林分生长特征、部分立地因子特征进行调查分析,研究造林立地因子对人工林生长的影响.结果表明:(1)38 a生油松人工林平均树高、胸径和冠幅分别为7.36 m、9.8 cm和261.1 cm;17 a生油松人工林平均树高、胸径和冠幅分别为2.86 m、8.1 ...     

马尾松格木幼林混交效果研究

【目的】通过分析不同混交模式下马尾松及格木生长量差异,探讨马尾松与格木混交的适宜模式,为马尾松格木混交林的营建提供理论基础。【方法】以广西凭祥5年生马尾松和格木人工林为研究对象,分析不同混交模式及混交比例马尾松和格木的胸径、树高、冠幅及材积的差异,探寻马尾松格木混交合理的混交模式和混交比例。【结果】1)不同混交模式马尾松胸径、树高、冠幅和材积均差异显著。处理a和CK3林分平均胸径最大,均为5.9 cm,CK3林分平均树高、上下冠幅和左右冠幅最大,分别为4.9、2.4和2.7 m。CK3的单株材积最大为0.0082 m3,处理c林分平均胸径、平均树高和单株材积均最小,分别为5.0 cm、4.1 m和0.0051 m3,同时上下冠幅和左右冠幅最小,分别为2.0 m和2.1 m。2)不同混交模式格木胸径、树高、冠幅和材积均差异显著。CK2林分平均胸径最大为5.1 cm,最小为处理c的4.1 cm。处理a和CK1林分平均树高最高,均为4.8 m,处理c树高最低为4.0 m。处理a、处理b和CK2上下冠幅最大,均为2.4 m,处理c上下冠幅最小为2.0 m,CK1左右冠幅最大为2.8 m,处理c左右冠幅最小为2.1 m。CK2格木单株材积最大为0.0058 m3,处理c单株材积最小为0.0035 m3。3)不同的混交模式和混交比例不影响径级数量的分布规律,各处理及对照中马尾松和格木的径级均呈正态分布。【结论】马尾松与格木幼林4种混交模式在胸径、树高、冠幅和材积均存在显著差异,马尾松与格木(75丛·hm^-2)各项生长指标相对较优,其次为马尾松+格木(4:1)行状混交,生长表现最差为马尾松+格木(210丛·hm^-2)。综合考虑,以培育马尾松大径材为经营目标,早期应选择马尾松与格木(75丛·hm^-2)模式进行混交。     

西南桦与高阿丁枫混交幼林生长及其混交匹配性分析

通过对普洱江城苦丁茶场及西双版纳普文试验林场西南桦与高阿丁枫不同混交模式的生长量指标进行调查分析,结果表明,(1)10年生前,西南桦的胸径较高阿丁枫速生,林木的平均胸径西南桦大于高阿丁枫;11年生后,林木的平均胸径高阿丁枫大于西南桦;(2)10年生时,普文林场混交林的西南桦和高阿丁枫胸径生长一致(11.7 cm),西南桦树高超出高阿丁枫树高3.0 m。说明混交林中西南桦为早期速生树种;幼林阶段,西南桦和高阿丁枫胸径于不同的时期呈现错峰生长。同时,西南桦和高阿丁枫的地理分布、立地的生态适宜条件和林学特征等分析结果表明这2个树种在适宜的混交模式下属混交匹配的树种。建议采用非均匀密度控制混交,以实现2个树种的良好生长和长期混交。     

GROWTH AND YIELD MODELS FOR DAHURIAN LARCH PLANTATIONS

Growth and yield models were developed for individual tress and stands based on336 temporary plots with 405 stem analysis trees of dahurian larch(Larix gmelinii(Rupr.)Rupr.)plantations throughout Daxing’anling mountains.Several equations were selected using nonlinearregression analysis.Results showed that the Richards equation was the best model for estimatingtree height,stand mean helght and stand dominant height from age; The Power equation was thebest model for prediction tree volume from DBH and tree height; The logarithmic stand volumeequation was good for predicting stand volume from age,mean height,basal area and other standvariables.These models can be used to construct the volume table, the site index table and other for-estry tables for dahurian larch plantations.     

坡位对杉木纯林生长的影响

通过对不同坡位7年生杉木纯林生长的调查与研究,结果显示:坡位对杉木纯林各生长指标均有显著影响;林分的平均胸径、树高、冠幅、枝下高、平均单株材积、单位面积木材蓄积量等生长指标一致表现为下坡位>中坡位>上坡位。坡位间以胸径和树高差异最为显著,下坡位林分平均胸径达到10.59 cm,高出中坡位的11.6%,高出上坡位的20.0%;下坡位林分平均树高为7.25 m,高出中坡位的16.7%,高出上坡位的25.4%。下坡位林分的单位面积木材蓄积量要远远高于中坡位和上坡位的,下坡位林分的单位面积木材蓄积量分别高出上坡位的75.1%,中坡位的44.8%。