Height variation of transplanted seedlings of Scots pine in Finnish nurseries

The height variation of Scots pine (Pinus sylvestris L.) seedling stock was studied using data from annual inventories of seedling crops. The heights of 240 batches of 2-year-old and 3-year-old transplanted seedling from the years 1973–1986 grown at three nurseries in southern Finland were examined. From each batch, 100 clusters of three successive seedlings were systematically located and measured. The mean heights of the seedling batches ranged from 7.8 to 27.0 cm, with a mean of 14.1 cm. Within batch standard deviations ranged from 1.7 to 7.8 cm, with a mean standard deviation of 3.7 cm. There was considerable variation in the mean heights of seedling batches both within a given year and between years. However, estimated variance components of seedling height showed that year, batch and cluster effect each accounted for about 20 percent of the total variance, while seedling effect explained the remaining 40 percent. The implications of results for single tree and batch grading are discussed.     

Temporal variation of microfibril angle in Eucalyptus nitens grown in different irrigation regimes

In 1990, a 2-ha plantation of Eucalyptus nitens (Deane and Maiden) Maiden was established in southeastern Tasmania and subjected to different irrigation regimes. Point dendrometers were installed in March 1995 to monitor radial stem movement every 15 min over several growing seasons. In this study, data from two growing seasons (1996-1998) were considered. From these measurements, daily increments of stem radius were determined. At the end of the second growing season, we extracted 12-mm cores and measured microfibril angles (MFA) of the wood at high resolution. Microfibril angles were rescaled on a time axis and mapped to daily and distance-based elements. Among treatments, irrigated trees in particular formed higher MFA early in the growing season (September-November) and lower MFA later in the growing season. Trees subjected to cyclic droughts showed clear relationships between MFA and soil water deficits, with MFA increasing in response to water stress release. Increases in MFA were preceded by accelerations in daily increment of stem radius. Among treatments, trees subjected to severe drought had the smallest MFA and generally low fluctuations in MFA. Irrigated trees were susceptible to changes in climate, whereas growth of the trees in the other treatments was limited by water availability. Use of path-analysis showed that temperature had an effect on stem radius increment but not on MFA; wind speed was the only factor that influenced MFA directly. Microfibril angle was correlated with stem shrinking and expansion phases; growth period length and growth rates were positively related to MFA.     

Long-term variation in Scots pine seed crop size and quality in northern Finland

Abstract

Quantitatively and qualitatively good Scots pine (Pinus sylvestris L.) seed years are rare in northern areas because of the short growing seasons. The seed crop size was studied in four natural pine forests in northern Finland during 1960–2004. Data on seed crop quality were collected during 1986–2004 and quality was determined by X-ray radiography. The long-term average annual seed crop of pine was 77 seeds m^?2, varying from 5 to 225 seeds m^?2. The expected average annual germination percentage of the seed crop was 61%, varying from 7 to 81%. General linear models were used to analyse the size and quality of the seed crop. Independent variables explained 52% of the variation in annual seed crop quality, and no proper model was found for determining the size of the seed crop; the coefficient of determination was only 5%. A combination of more than 100 seeds m^?2 and an expected germination percentage of over 50% was observed once during the years 1986–2004. In most years, natural regeneration in northern Finland is difficult and often limited by the seed crop's quality as well as quantity. These results confirm the importance of seed tree cutting in abundant seed crop years to enable the natural regeneration of pine.     

Fibrillar angle variation in red pine determined by Senarmont compensation

Summary Fibrillar angles were determined in the secondary wall layers of three rings across the stem of a mature red pine. The Senarmont compensation method was adopted for this study since it gives results for all three wall layers. It involves the determination of birefringence in a series of sections cut at various angles to the long axis of the tracheids. Maxima and minima in the resultant plots of birefringence versus section angle correspond to fibrillar orientations. In the early latewood of each ring, the S₂ layr was found to contain a single Z helix of small fibrillar angle and the S₁ dual S and Z helices of much flatter orientation. Results for S₃ were found to be so close to those for the corresponding S₂ layers that there was some doubt as to their validity. In each layer a slight decrease in fibrillar angle was observed between rings 4 (juvenile wood) and 12 (transition zone); a sharper decrease was seen between ring 12 and the mature ring 20. This trend corresponds inversely to tracheid length differences for the three rings. Correlations of tracheid wall thickness and overall cross-sectional dimensions with the fibrillar angle variations were less readily apparent.Contribution No. 59 from Empire State Paper Research Institute, State University College of Forestry, Syracuse, New York 13210. The authors gratefully acknowledge the discussions and advice of Drs. W. A. Côté, Jr. and R. E. Mark.     

Dielectric spectroscopy of Scots pine

Dielectric properties of Scots pine (42 trees) were compared with density, moisture content (MC), and resin acid content (RAC) (of heartwood). The samples were measured in frozen, green, conditioned and non-conditioned dry moisture states to evaluate the potential of dielectric spectroscopy in determining the wood characteristics at different stages of wood processing. Heartwood and sapwood parts of each sample were measured separately, and through-transmission measurement was conducted in longitudinal and tangential direction at frequencies from 1 MHz to 1 GHz. The MC and density correlated significantly with the dielectric parameters in both measurement directions but especially in longitudinal direction. The RAC of the heartwood correlated significantly with tanδ and ε″/(ε′ ? 1) of the green samples measured in the tangential direction at frequencies above 200 MHz. The correlation at 1 GHz was ?0.56 for green samples, ?0.66 for conditioned samples and ?0.61 for non-conditioned samples (P < 0.001, for all). The study suggests that the extractives also affect radio-frequency dielectric responses, which might be used for extractive analyses of pine heartwood.     

Theory of X-ray measurement of microfibril angle in wood

Summary The property of fibre symmetry as exhibited by wood cellulose can be used to derive an explicit relationship between the orientation of a cellulose microfibril and the orientation of the X-ray beam diffracted by any of its crystallographic planes. The solution applies to a microfibril of any orientation and so is well suited to evaluating the microfibril angle distribution in wood containing cells of any cross-sectional shape. The (002) and (040) reflections of cellulose have complementary properties that could be exploited to enable current problems associated with the use of each individually for evaluating the mean microfibril angle of the S2 layer to be overcome. It is expected that it will be possible to measure the microfibril angle distribution throughout the whole cell wall and also measure the average cell cross-section of a wood sample, by analysing (002) and (040) diffraction profiles in conjunction with each other.This work is supported by the NZ Foundation for Research, Science and Technology under contract # UOC 401     

Growth and phenology variation in progeny of Scots pine seed orchards and commercial seed stands

Tree improvement in Poland has been most advanced for Scots pine, but existing seed orchards have not been progeny-tested yet. We examined variation in growth traits—tree height at ages 4 and 8 years, and diameter at age 13 years—in the common garden experiment testing open-pollinated progenies of 31 seed orchards and 5 commercial seed stands (referred to as populations) at 5 locations. We also examined bud burst phenology at two to five sites at three growing seasons. At one experimental site during the 5th growing season, we measured shoot growth rhythm on all populations. Similar measurements of shoot growth were done on a subset of populations during the 6th growing season together with the analysis of needle growth and foliar chemistry. We found significant variation among populations in growth traits, but also significant population × site (G × E) interactions. We used the regression approach and ecovalence analysis to examine populations’ performance stability. Most populations had average responsiveness to environment, and a set of least-responsive poor-growing populations contributed the most to the G × E interaction. Variation in bud burst phenology was associated with geographical distribution of tested progenies. The early bud-bursting populations originated from the north-eastern to north-central Poland, and a group of late bud-bursting populations originated mainly from the south-eastern region. Correlations between bud burst and growth traits were weak to medium and varied by site, but early bud-bursting populations tended to show stronger growth on height and diameter. We found significant differences among populations in final leader length, shoot elongation time and relative growth rate (RGR). However, RGR and shoot elongation time explained less than 30% of variation in leader length and were weakly correlated with tree height. Populations varied in needle length, specific leaf area and foliar nitrogen concentration, but time trends in these traits did not vary among populations or predefined groups of populations. Therefore, the analysis of growth rhythm or needle traits did not help resolve variation in tree growth to support selection decisions. Contrary to our expectation, progeny of seed orchards did not perform significantly better than that of commercial seed stands. This finding, however, should not be extrapolated beyond our set of populations. Nonetheless, the local seed sources were not always the best. From a selection standpoint, our results help culling the worst populations rather than selecting the best ones. Therefore, testing individual family progeny and genetic roguing of existing seed orchards is highly recommended.     

Stand Density Control Diagrams for Scots pine and Austrian black pine plantations in Bulgaria

Stand Density Control Diagram (SDCD) is a stand-level mathematical model, which describes the relationships between yield, density and mortality throughout all stages of stand development. The SDCD is primarily used to derive density control schedules by management objectives. The main objectives of the present study are to define a modified model of SDCD for application to Scots pine (Pinus sylvestris L.) and Austrian black pine (Pinus nigra Arn.) plantations in Bulgaria, to examine the fitness of the model with representative experimental data sets from plantations of both species and to present a way of direct application of the SDCDs for practical purposes. The constructed SDCDs characterize the spatial-temporal dynamics of the pine plantations in a broad range of densities, forest sites and growth stages from 4 to 26 (28) m of dominant height class. The full density lines were fixed with self-thinning exponents α = 1.69 and α = 1.75 for Scots pine and Austrian black pine, respectively, and the trajectories of natural thinning for 23 initial densities (444-40000/ha) were determined. A direct way for application of the SDCDs to the plantation management was designed to estimate the optimal initial densities for the maximum attainable final yield and large-size wood production by self-thinning stands.     

Theory of X-ray measurement of microfibril angle in wood

Summary A diffraction intensity function for material bodies composed of arrays of crystalline fibres such as occurs with the cellulose of wood has been derived. It is implied in the analysis that the crystalline fibres making up the body have fibre symmetry- that there is a tendency for groups of fibres to have one set of crystal axes parallel while in the orthogonal direction the axes assume a low degree of order. It is further assumed that the patterns of the angular arrangement of the fibre groups relative to one axis of the body is independent of the direction about that axis. These conditions are believed to be compatible with the cellulosic structure found in wood. Thus it becomes possible to calculate the expected diffraction intensity profiles of realistic (and therefore complex) models of wood. This has aided the interpretation of the reflections from the (040) crystal planes of cellulose which are contaminated by low level reflections from other crystal planes, and it has been found that it might be possible by conjoint analysis of the paratropic (002) reflections and the diatropic (040) reflections to measure the complete cell wall planar microfibril angle distribution and the shape of the cell wall cross-section.This work is supported by the New Zealand Foundation for Research Science and Technology under contract # UOC 401     

Determination of microfibril angle distribution by X-ray diffraction

X-ray diffraction is a well-established method for the determination of the mean microfibril angle (MFA). When the sample is a slice of wood variations in the fibre orientation, the shape of the cells, and the measurement geometry affect the intensity curve. A general form for diffraction conditions in terms of angles describing the fibre orientation and the shape of the cell was derived. Intensity curves were calculated by using Monte Carlo method and compared with experimental ones. Both peak fitting and variance methods were used for determining the mean MFA from the intensity curves. Norway spruce was used as an example. Results indicate that deviations in the fibre orientation, the spiral grain, do not affect the mean MFA considerably when using the symmetrical transmission geometry. When using the perpendicular transmission geometry large deviations in spiral grain or tips tend to increase the MFA determined with the variance method and decrease the MFA determined with the fitting method. The shape of the cell should be considered when using the reflection 200 and the fitting method. The variance method is insensitive to the shape of the cell.     

Boron retranslocation in Scots pine and Norway spruce

We previously traced 10B-enriched boric acid from shoots to roots to demonstrate the translocation of boron (B) in Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) seedlings. To gain a more detailed understanding of B translocation, we sought: (1) to demonstrate B retranslocation directly, by showing that foliar-applied 10B is located in the new growth after dormancy; and (2) to assess whether shoot-applied B affects growth in the long term. We applied 10B-enriched boric acid to needles of Scots pine and Norway spruce seedlings. After a dormancy period and 9 weeks of growth, small but significant increases in the 10B isotope were found in the new stem and needles of both species. In Scots pine, the total B concentration of the new stem was also increased. Both species contained polyols, particularly pinitol and inositol. Boron-polyol complexes may provide a mechanism for mobilizing B in these species. To determine the long-term effects of applied B, seedlings were grown for two growing seasons after the application of 10B to shoots. In Norway spruce, the proportion of 10B in the root systems and current needles of the harvest year was slightly higher than in the controls, and in Scots pine root systems, marginally so. The B treatment had no effect on growth of Norway spruce seedlings. In Scots pine seedlings, the B treatment caused a 33% increase in total dry mass and significantly increased the number of side branches.     

Determining the transition from juvenile to mature wood microfibril angle in lodgepole pine: a comparison of six different two-segment models

? Context

The transition of microfibril angle (MFA) values from juvenile to mature wood marks the change from variable, low-quality wood to stronger and more consistent wood that can produce higher value products.

? Aims

We evaluate the utility of different statistical models that predict how much of a log is higher quality mature wood based on MFA.

? Methods

MFA was measured from pith to bark at breast height in six lodgepole pine stands in western Canada. Six different forms of two-segment regression models were assessed to determine the point of transition (TP) in MFA from juvenile to mature wood.

? Results

All six models provided useful and significant TP estimates. In the first segment (juvenile phase), the quadratic form produced the most conservative TPs, the linear form the least conservative, and the exponential form was intermediate. A linear second segment (mature phase) was only a minor improvement over a constant. There were significant differences in MFA TP among some sites. Analyses of the relationships between TP and tree variables, e.g., DBH, height, were inconclusive.

? Conclusions

Any of the six two-segment models can be used objectively to estimate MFA transition points; the choice of model will allow mill managers to manage risk in product out-turn.     

Empirical models for predicting the production of wild mushrooms in Scots pine (Pinus sylvestris L.) forests in the Central Pyrenees

Mushroom picking has become a widespread autumn recreational activity in the Central Pyrenees and other regions of Spain. Predictive models that relate mushroom production or fungal species richness with forest stand and site characteristics are not available. This study used mushroom production data from 24 Scots pine plots over 3 years to develop a predictive model that could facilitate forest management decisions when comparing silvicultural options in terms of mushroom production. Mixed modelling was used to model the dependence of mushroom production on stand and site factors. The results showed that productions were greatest when stand basal area was approximately 20 m² ha^?1. Increasing elevation and northern aspect increased total mushroom production as well as the production of edible and marketed mushrooms. Increasing slope decreased productions. Marketed Lactarius spp., the most important group collected in the region, showed similar relationships. The annual variation in mushroom production correlated with autumn rainfall. Mushroom species richness was highest when the total production was highest.     

Allometric equations for estimating the foliage biomass of Scots pine

The research described in this paper was performed in the Niepolomice Forest (Southern Poland) in 2001 as part of the Forest Environmental Monitoring and Management System (FOREMMS; 5FP IST) project. The material for the present study consisted of the measurement results of the biomass of Scots pine shoots with needles and needles alone carried out on 113 felled sample trees. The purpose of this study was to construct empirical equations for estimating the foliage biomass of Scots pine from easy to measure parameters. To achieve this aim, the dependence of the foliage biomass of Scots pine on stem diameter, height, age, crown length, basal area increment of the trees was analyzed. Using the biometric characteristics such as: tree diameter at breast height (dbh), basal area increment, age, height, and crown length empirical equations for estimating the foliage biomass of Scots pine reasonably precisely have been established. The created empirical equation gives accurate foliage biomass estimates. The explained variability varies between 65 and 85%, it depends on the number of variables applied in the equation. The equations presented in this paper were created with a view to their possible use in ecological studies where biomass quantity may be used, for example, in modeling carbon circulation in the forest ecosystem. From the point of view of forestry practice, these equations may help to assess biomass production in Scots pine stands.     

Genetic variation and divergence in Scots pine (Pinus sylvestris L.) within its natural range in Italy

Twenty-one populations of Scots pine sampled over the entire Italian range of the species were analysed for genetic variation scored at nine nuclear SSR markers. The main aim of the work was to find genetic features useful for conservation management, namely allelic composition, gene diversity and differentiation. High levels of intra-population variability were scored. The only population sampled in the Apennines gave the lowest values, confirming the genetic erosion undergone in the Scots pine remnants in this area. A low level of genetic variability was also scored for populations from the Po valley and hills of Piedmont. Most genetic diversity was found within populations, while only a small amount occurred among them (F _ST?=?0.058). Both Bayesian clustering and sPCA analysis showed a East–West subdivision, notwithstanding the unclear position of populations from the Po valley. The population from the Apennines was always clearly separated from the others. The results are discussed in terms of post-glacial recolonisation, as well as for defining genetically homogeneous regions for Scots pine in Italy. The management of genetic resources could benefit from the identification of such ‘gene zones’, thereby avoiding the use of non-local reproductive material for plantations, which can represent one of the most important reasons for failure of reforestation. In addition, the assessment of the biogeographic genetic structure by neutral markers is a prerequisite for disentangling the influence of selectively neutral and non-neutral processes on the distribution of adaptive genetic variability.     

The variation of microfibril angle in South African grown Pinus patula and its influence on the stiffness of structural lumber

Reduction in the rotation ages of softwood saw-log plantations in South Africa is causing increased proportions of low stiffness sawn lumber at final harvest. It has been shown for some species that the microfibril angle (MFA) of the S2 layer of tracheids is strongly related to the modulus of elasticity (MOE) of wood, even more so than wood density, especially in wood formed during juvenile growth. The objectives of this study were to describe the variation in MFA in young Pinus patula trees and to determine the relationship between MFA and the dynamic MOE of sawn P. patula lumber. Thirty 16- to 20-year-old trees from six compartments from the Mpumalanga escarpment were processed into discs and lumber. The MFA, density and ring width were measured at two height levels using Silviscan 3. The average annual ring MFA varied between 7° and 29°; the pattern of variation depended mainly on height level and the ring number from the pith. The MFA in P. patula followed the same within-tree variation trends as in New Zealand-grown Pinus radiata but the average MFA was lower in absolute terms and differences between height levels were less pronounced. The MFA and density exhibited highly significant Pearson correlations of 0.73 and 0.70, respectively, with board dynamic MOE. A multiple regression model, which included MFA, density and ring width, explained 71% of the variation in the dynamic MOE of boards. A sensitivity analysis on the model showed that MFA and density had approximately similar influences on predicting the dynamic MOE of Pinus patula boards.     

Predicting lumber grade and by-product yields for Scots pine trees

The purpose of this study was to develop models for estimating yields of lumber grades and by-products of individual Scots pine (Pinus sylvestris L.) trees using stem and crown dimensions as explanatory variables. Two separate data sets were used: (1) one simulated by the process-based growth model, PipeQual, which provides information about stem form and branch properties. The model was used to predict the 3D structure of Scots pine stems from thinning regimes of varying intensity and rotation periods and (2) an empirical data set with detailed 3D measurements of stem structure. The stems were sawn using the WoodCim sawing simulator and the yields and grades of the individual sawn pieces, as well as by-products, were recorded. The sawn timber was classified on A, B, C and D-grades for side and centre boards separately (Nordic Timber grading). By-products were pulpwood, sawmill chips, sawdust and bark.     

Modelling stand structure in young Scots pine dominated stands

The purpose of this study was to construct models for predicting the structure of young Scots pine (Pinus sylvestris L.) stands. The two-parameter Weibull function characterized the height distribution of the stands. In young stands height was preferred to dbh as a random variable because of its continuous feature. Tree diameters were predicted using a multiplicative model, fitted as a linearized mixed-effect model. The modelling data consisted of repeatedly measured Scots pine dominated juvenile stands, carried out on a sub-sample of the 7th National Forest Inventory. The data covered a dominant height range from 0.2 up to 17 m. Two independent data sets were used to validate the models. The Weibull function was fitted using the maximum likelihood method. Four methods for predicting the distributions were compared: (1) parameter prediction models (PPM) consisting of seemingly unrelated regression equations, (2) a generalized linear model (GLM) which was a one-stage distribution and model fitting procedure, (3) a hybrid method including PPM for the shape parameter together with moment-based parameter recovery for the scale parameter, and (4) inclusion of moment-based parameter recovery for the scale parameter in the estimated GLM. Goodness-of-fit were tested in terms of Kolmogorov–Smirnov and error index statistics. Parameter recovery showed no improvement when used with PPM, but it improved GLM and gave the overall best performance for this new method. The constructed diameter–height model showed quite flexible and unbiased behaviour. Models are recommended as practical tools for Finnish forest management planning purposes.