Comparison of Herbicides for Reducing Annual Grass Emergence in Two Great Basin Soils

Reducing seed germination and seedling emergence of downy brome (Bromus tectorum L.) improves the success of revegetating degraded shrubland ecosystems. While pre-emergence herbicides can potentially reduce these two processes, their impact on germination and emergence of downy brome and revegetation species in semiarid ecosystems is poorly understood and has not been comprehensively studied in soils with potentially contrasting herbicide bioavailability (i.e., residual plant activity). We designed a greenhouse experiment to evaluate the effects two pre-emergence acetolactate synthase–inhibiting herbicides (rimsulfuron and imazapic) on germination and emergence of downy brome and two revegetation grass species (crested wheatgrass [Agropyron cristatum {L.} Gaertn.] and bottlebrush squirreltail [Elymus elymoides {Raf.} Swezey]) that were grown in representative soils from salt desert and sagebrush shrublands. Pre-emergence herbicides significantly (P &spilt; 0.05) reduced seedling emergence and biomass production of downy brome and crested wheatgrass and increased mortality more so in sagebrush compared to salt desert soil, suggesting that these common Great Basin soils fundamentally differ in herbicide bioavailability. Also, germination and emergence of the two highly responsive species (crested wheatgrass and downy brome) were clearly more impacted by rimsulfuron than imazapic. We discuss these results in terms of how the specific soil physiochemical properties influence herbicide adsorption and leaching. Our results shed new light on the relative performance of these two promising herbicides and the importance of considering soil properties when applying pre-emergence herbicides to reduce germination and emergence of invasive annual grasses and create suitable seedbed conditions for revegetation.     

Public Perceptions of Sagebrush Ecosystem Management in the Great Basin

Intact sagebrush communities in the Great Basin are rapidly disappearing because of invasion of nonnative plants, large wildfires, and encroachment of pinyon and juniper woodlands. Land management options, including the use of prescribed fire, grazing, herbicides, or mechanical treatments, can reduce the potential for wildfire and restore plant communities. Public acceptance of management actions, and trust in agencies to carry out those actions, is a critical component of developing and implementing successful long-term land management plans. This study examines citizens’ opinions and perceptions about rangeland management in the Great Basin. In fall 2006 we conducted a mail survey of randomly selected households in three urban and three rural regions of the Great Basin, receiving 1 345 valid responses for a 45% response rate. Overall, respondents perceived that the environment is moderately healthy; however, they do recognize threats to sagebrush ecosystems. Public acceptance is relatively high for the use of prescribed fire, grazing, felling woodland trees, and mowing shrubs, but low for herbicide treatment and chaining. Although respondents indicated high levels of acceptance for some management actions, they expressed relatively low levels of trust in land management agencies to implement these actions.     

Vegetation Characteristics of Mountain and Wyoming Big Sagebrush Plant Communities in the Northern Great Basin

Dominant plant species are often used as indicators of site potential in forest and rangelands. However, subspecies of dominant vegetation often indicate different site characteristics and, therefore, may be more useful indicators of plant community potential and provide more precise information for management. Big sagebrush (Artemisia tridentata Nutt.) occurs across large expanses of the western United States. Common subspecies of big sagebrush have considerable variation in the types of sites they occupy, but information that quantifies differences in their vegetation characteristics is lacking. Consequently, wildlife and land management guidelines frequently do not differentiate between subspecies of big sagebrush. To quantify vegetation characteristics between two common subspecies of big sagebrush, we sampled 106 intact big sagebrush plant communities. Half of the sampled plant communities were Wyoming big sagebrush (A. tridentata subsp. wyomingensis [Beetle & A. Young] S. L. Welsh) plant communities, and the other half were mountain big sagebrush (A. tridentata subsp. vaseyana [Rydb.] Beetle) plant communities. In general, mountain big sagebrush plant communities were more diverse and had greater vegetation cover, density, and biomass production than Wyoming big sagebrush plant communities. Sagebrush cover was, on average, 2.4-fold higher in mountain big sagebrush plant communities. Perennial forb density and cover were 3.8- and 5.6-fold greater in mountain compared to Wyoming big sagebrush plant communities. Total herbaceous biomass production was approximately twofold greater in mountain than Wyoming big sagebrush plant communities. The results of this study suggest that management guidelines for grazing, wildlife habitat, and other uses should recognize widespread subspecies as indicators of differences in site potentials.     

Linking Plant Spatial Patterns and Ecological Processes in Grazed Great Basin Plant Communities

Observational studies of plant spatial patterns are common, but are often criticized for lacking a temporal component and for their inability to disentangle the effect of multiple community-structuring processes on plant spatial patterns. We addressed these criticisms in an observational study of Great Basin shrub-steppe communities that have been converted to a managed grazing system of planted crested wheatgrass (Agropyron cristatum [L.] Gaertn.) stands. We hypothesized that intraspecific interference and livestock grazing were important community-structuring processes that would leave unique spatiotemporal signatures. We used a survey-grade global positioning system to quantify crested wheatgrass spatial patterns along a chronosequence of stands that differed only in time since planting (9–57 yr), as well as in a 57-yr-old grazing exclosure to examine pattern formation in the absence of grazing. Three replicate survey plots were established in each stand, and a total of 6 197 grasses were marked with a spatial error of ≤ 2 cm. The data were analyzed using L-statistics in program R, and hypothesis testing was conducted using Monte Carlo simulation procedures. We detected fine-scale regularity, frequently considered a sign of interference via resource competition, in all stands including the exclosure. Coarser-scale aggregation, which we attributed to the effects of prolonged grazing disturbance, was only detected in the oldest grazed stand. Our results suggest that interference acts over finer spatial and temporal scales than grazing in structuring these stands, reinforcing the importance of interference in semiarid communities. Analysis of exclosure data suggests that, in the absence of grazing, crested wheatgrass stands organize into a statistically regular pattern when primarily influenced by interference. In the presence of prolonged grazing, crested wheatgrass stands become more heterogeneous over time, likely a result of seedling mortality via disturbance by cattle.     

Restoration of Sagebrush in Crested Wheatgrass Communities: Longer-Term Evaluation in Northern Great Basin

Crested wheatgrass (Agropyron cristatum [L] Gaertm. and Agropyron desertorum [Fisch.] Schult.), an introduced bunchgrass, has been seeded on millions of hectares of sagebrush steppe. It can establish near-monocultures; therefore, reestablishing native vegetation in these communities is often a restoration goal. Efforts to restore native vegetation assemblages by controlling crested wheatgrass and seeding diverse species mixes have largely failed. Restoring sagebrush, largely through planting seedlings, has shown promise in short-term studies but has not been evaluated over longer timeframes. We investigated the reestablishment of Wyoming big sagebrush (Artemisia tridentata spp. wyomingensis [Beetle & A. Young] S.L. Welsh) in crested wheatgrass communities, where it had been broadcast seeded (seeded) or planted as seedlings (planted) across varying levels of crested wheatgrass control with a herbicide (glyphosate) for up to 9 yr post seeding/planting. Planting sagebrush seedlings in crested wheatgrass stands resulted in full recovery of sagebrush density and increasing sagebrush cover over time. Broadcast seeding failed to establish any sagebrush, except at the highest levels of crested wheatgrass control. Reducing crested wheatgrass did not influence density, cover, or size of sagebrush in the planted treatment, and therefore, crested wheatgrass control is probably unnecessary when using sagebrush seedlings. Herbaceous cover and density were generally less in the planted treatment, probably as a result of increased competition from sagebrush. This trade-off between sagebrush and herbaceous vegetation should be considered when developing plans for restoring sagebrush steppe. Our results suggest that planting sagebrush seedlings can increase the compositional and structural diversity in near-monocultures of crested wheatgrass and thereby improve habitat for sagebrush-associated wildlife. Planting native shrub seedlings may be a method to increase diversity in other monotypic stands of introduced grasses.     

Performance of Quantitative Vegetation Sampling Methods Across Gradients of Cover in Great Basin Plant Communities

Resource managers and scientists need efficient, reliable methods for quantifying vegetation to conduct basic research, evaluate land management actions, and monitor trends in habitat conditions. We examined three methods for quantifying vegetation in 1-ha plots among different plant communities in the northern Great Basin: photography-based grid-point intercept (GPI), line-point intercept (LPI), and point-quarter (PQ). We also evaluated each method for within-plot subsampling adequacy and effort requirements relative to information gain. We found that, for most functional groups, percent cover measurements collected with the use of LPI, GPI, and PQ methods were strongly correlated. These correlations were even stronger when we used data from the upper canopy only (i.e., top “hit” of pin flags) in LPI to estimate cover. PQ was best at quantifying cover of sparse plants such as shrubs in early successional habitats. As cover of a given functional group decreased within plots, the variance of the cover estimate increased substantially, which required more subsamples per plot (i.e., transect lines, quadrats) to achieve reliable precision. For GPI, we found that that six–nine quadrats per hectare were sufficient to characterize the vegetation in most of the plant communities sampled. All three methods reasonably characterized the vegetation in our plots, and each has advantages depending on characteristics of the vegetation, such as cover or heterogeneity, study goals, precision of measurements required, and efficiency needed.     

松嫩草原几种主要禾草植物群落水分生态的比较研究

采用自行设计的蒸散仪和ＣａｍｐｂｅｌｌＣＲ１０ｄａｔａｌｏｇｇｅｒ自动气象设备,研究了松嫩草原羊草、星星草、朝鲜碱茅和虎尾草４种主要禾草植物水分生态状况。结果表明,在生长季晴天条件下,４种群落的蒸腾速度基本为双峰曲线,但同一时期各各落间及不同时期同一群落的蒸腾速度差异显著。在影响群落蒸腾速度的诸多因素中,太阳辐射强度为主导因了,其与群落蒸腾速度的相关关系最密切。在生长季,４种群落中,只有虎尾草群落     

Postfire Seeding and Plant Community Recovery in the Great Basin

As wildland fire frequency increases around the globe, a better understanding of the patterns of plant community recovery in burned landscapes is needed to improve rehabilitation efforts. We measured establishment of seeded species, colonization of Bromus tectorum and other nonnative annual plants, and recovery of nonseeded native species in topographically distinct areas within five fires that burned Great Basin shrub-steppe communities in Elko County, Nevada. Plant density, frequency, and cover data were collected annually for 4 yr postfire. Vegetation composition varied among flat areas and north- and south-facing aspects, and changed over the course of the sampling period; recovery varied among sites. In general, B. tectorum densities were higher on south aspects, particularly 3 and 4 yr after fire, when densities increased dramatically relative to prefire conditions. Nonseeded native perennial grasses, forbs, and shrubs were abundant in three of the five fire sites, and were more likely to be present on north aspects and flat areas. Over time, nonseeded perennial grass densities remained relatively constant, and nonseeded forbs and shrubs increased. Seeded species were most likely to establish in flat areas, and the density of seeded perennial grasses, forbs, and shrubs decreased over time. Frequency and density measurements were highly correlated, especially for perennial species. Our results emphasize the value of considering site aspect and the potential for native regrowth when planning and monitoring restorations. For example, effective rehabilitation of south aspects may require the development of new restoration methods, whereas north aspects and flat areas in sites with a strong native component were not improved by the addition of seeded species, and may require weed control treatments, rather than reseeding, to improve recovery. Tailoring revegetation objectives, seed mixes, seeding rates, and monitoring efforts to conditions that vary within sites may lead to more cost effective and successful restoration.     

Basalt Milkvetch Responses to Novel Restoration Treatments in the Great Basin

The restoration of native forbs in the Great Basin and similar dryland ecosystems remains a great challenge for land managers. Variable soil water, precocious germination and emergence, and the presence of soil fungal pathogens often reduce plant establishment. Novel restoration treatments that increase soil water during early forb life stages, delay germination and emergence to coincide with favorable spring conditions, and reduce mortality from soil fungal pathogens may benefit native forb restoration. We compared the efficacy of three novel treatments—snow fencing, plant protection fabric, and seed coatings—to improve establishment of a forb species native to the Great Basin, basalt milkvetch (Astragalus filipes Torr. ex A. Gray). We replicated treatments plus a no-treatment control at three sites in the Great Basin in a randomized complete block design over 2 yr. To evaluate the efficacy of our restoration treatments, we measured germination, seedling emergence, establishment, and second-yr survival. The effects of snow fencing and plant protection fabric varied by site and life stage and were likely influenced by the wet spring conditions across sites, particularly the two northern sites. Snow fencing increased establishment and second-yr survival at one site. Plant protection fabric did not benefit any life stage at any site. Seed coatings increased seedling emergence at all sites but did not affect establishment or survival. Overall, second-yr survival remained low across all sites and treatments. Our study indicates that our treatments can positively affect basalt milkvetch emergence, establishment, and survival, but additional management actions are needed to improve long-term restoration success of native forbs.     

Native Vegetation Composition in Crested Wheatgrass in Northwestern Great Basin

Crested wheatgrass, an introduced perennial bunchgrass, has been seeded extensively on the rangelands of western North America. There is a perception that this species is very competitive and that it forms monoculture or low diversity stands where successfully seeded. However, there is limited information on species composition in sites previously seeded to crested wheatgrass. We measured native vegetation and environmental characteristics in areas seeded with crested wheatgrass across the northwestern Great Basin. Plant community composition within these crested wheatgrass stands was variable, from seedings that were near monocultures of crested wheatgrass to those that contained more diverse assemblages of native vegetation, especially shrubs. Environmental factors explained a range of functional group variability from 0% of annual grass density to 56% of large native bunchgrass density. Soil texture appeared to be an important environmental characteristic in explaining vegetation cover and density. Native vegetation was, for all functional groups, positively correlated with soils lower in sand content. Our results suggest environmental differences explain some of the variability of native vegetation in crested wheatgrass stands, and this information will be useful in assessing the potential for native vegetation to co-occupy sites seeded with crested wheatgrass. This research also suggests that crested wheatgrass seedings do not always remain in near monoculture vegetation states as seedings substantially varied in native vegetation composition and abundance with some seeded areas having a more diverse assemblage of native vegetation. In half the sites, there were five or more perennial herbaceous species and 63% of sites contained Wyoming big sagebrush. Although not exclusively true, species most commonly encountered in crested wheatgrass seedings are those that are able to minimize competition with crested wheatgrass via temporal (i.e., Sandberg bluegrass, annual forbs, annual grasses) or spatial (i.e., shrubs) differentiation in resource use.     

青海湖湖盆南部植物群落二维极点排序

利用极点排序对青海湖湖盆南部的植物群落进行了分析。结果表明，在研究地区不同海拔影响植物群落分布的主要环境因子不同。根据影响植落分布的环境因子的变化情况，可将研究地区划分为３个地段，即较低地段，中间地段和较高地段。     

Short-Term Response of Sage-Grouse Nesting to Conifer Removal in the Northern Great Basin

Conifer woodlands expanding into sage-steppe (Artemisia spp.) are a threat to sagebrush obligate species including the imperiled greater sage-grouse (Centrocercus urophasianus). Conifer removal is accelerating rapidly despite a lack of empirical evidence to assess outcomes to grouse. Using a before-after-control-impact design, we evaluated short-term effects of conifer removal on nesting habitat use by monitoring 262 sage-grouse nests in the northern Great Basin during 2010–2014. Tree removal made available for nesting an additional 28% of the treatment landscape by expanding habitat an estimated 9603 ha (3201 ha [± 480 SE] annually). Relative probability of nesting in newly restored sites increased by 22% annually, and females were 43% more likely to nest within 1000 m of treatments. From 2011 (pretreatment) to 2014 (3 yr after treatments began), 29% of the marked population (9.5% [± 1.2 SE] annually) had shifted its nesting activities into mountain big sagebrush habitats that were cleared of encroaching conifer. Grouping treatments likely contributed to beneficial outcomes for grouse as individual removal projects averaged just 87 ha in size but cumulatively covered a fifth of the study area. Collaboratively identifying future priority watersheds and implementing treatments across public and private ownerships is vital to effectively restore the sage-steppe ecosystem for nesting sage-grouse.     

加强基层兽医工作机构国有资产管理的思考

<正>县级畜牧兽医局、动物疫病预防控制中心、动物卫生监督所和乡镇畜牧兽医站是我国基层兽医工作机构,担负着兽医行政管理、动物疫病防控、动物卫生行政执法和具体组织实施动物防疫公益性服务活动等社会事务,是我国兽医公共卫生安全管理的基础和基本工作单位。     

Encounters with Pinyon-Juniper Influence Riskier Movements in Greater Sage-Grouse Across the Great Basin

Fine-scale spatiotemporal studies can better identify relationships between individual survival and habitat fragmentation so that mechanistic interpretations can be made at the population level. Recent advances in Global Positioning System (GPS) technology and statistical models capable of deconstructing high-frequency location data have facilitated interpretation of animal movement within a behaviorally mechanistic framework. Habitat fragmentation due to singleleaf pinyon (Pinus monophylla; hereafter pinyon) and Utah juniper (Juniperus osteosperma; hereafter juniper) encroachment into sagebrush (Artemisia spp.) communities is a commonly implicated perturbation that can adversely influence greater sage-grouse (Centrocercus urophasianus; hereafter sage-grouse) demographic rates. Using an extensive GPS data set (233 birds and 282,954 locations) across 12 study sites within the Great Basin, we conducted a behavioral change point analysis and constructed Brownian bridge movement models from each behaviorally homogenous section. We found the probability of encountering pinyon-juniper among adults was two and three times greater than that of yearlings and juveniles, respectively. However, the movement rate in response to the probability of encountering pinyon-juniper trees was 1.5 times greater for juveniles. Parameter estimates indicated a 6.1% increase in the probability of encountering pinyon-juniper coupled with a 6.2 km/hour increase in movement speed resulted in a 56%, 42% and 16% increase in risk of daily mortality, for juveniles, yearlings, and adults, respectively. The effect of pinyon-juniper encounters on survival was dependent on movement rate and differed among age class. Under fast speed movements (i.e., flight), mortality risk increased as encountering pinyon-juniper increased across all age classes. In contrast, slower speeds (i.e., average) yielded similar adverse effects for juveniles and yearlings but not for adults. This analytical framework supports a behavioral mechanism that explains reduced survival related to pinyon-juniper within sagebrush environments, whereby encountering pinyon-juniper stimulates riskier movements that likely increase vulnerability to visually acute predators.