Effect of different grazing management systems on the herbage mass and pasture height of a Nardus stricta grassland in western Scotland, United Kingdom

The aims of this study were to examine the effect of three grazing treatments (year‐round stocking rates of 0·8 ewes ha^?1, 0·5 ewes ha^?1 and 0·5 ewes ha^?1 plus grazing cattle in summer), imposed for 4 years, on the herbage mass and surface height of a Nardus stricta‐dominated grassland in western Scotland and to obtain estimates of annual productivity of this grassland. Nardus stricta‐dominated grassland comprised proportionately 0·20 of the grazing area. Stocking rate of sheep had no significant effect on the herbage mass of the grassland in the first 2 years of the experiment, although mean summer pasture heights were significantly higher under the lower stocking rate of sheep. The pasture on the treatment with cattle grazing in summer had a significantly lower herbage mass and lower surface height than the two sheep‐only grazing treatments. Year‐to‐year variation in the herbage mass and surface height of herbage in summer was greater than the effect of treatments. Despite changes in surface height, the structural diversity of the grasslands was not increased by the treatments. The annual production of vascular plant material ranged from 417 g DM m^?2 in 1994 to 628 g DM m^?2 in 1996.     

Impact of deferred grazing and fertilizer on herbage production,soil seed reserve and nutritive value of native pastures in steep hill country of southern Australia

Developing sustainable grazing management systems based on perennial species is critical to preventing land degradation in marginal land classes. A field study was conducted from 2002 to 2006 to identify the impacts of deferred grazing (no defoliation of pastures for a period generally from spring to autumn) and fertilizer application on herbage accumulation, soil seed reserve and nutritive value in a hill pasture in western Victoria, Australia. Three deferred grazing strategies were used: short‐term deferred grazing (no defoliation between October and January), long‐term deferred grazing (no defoliation from October to the autumn break) and optimized deferred grazing (withholding time from grazing commenced between annual grass stem elongation and seed head emergence and concluded in February/March). These treatments were applied with two fertilizer levels (with or without fertilizer at 50 kg phosphorus ha^?1 and 2000 kg lime ha^?1 applied in year 1 only) in a factorial arrangement and two additional treatments: continuous grazing (CG) and no grazing (NG) in year 1. The deferred grazing treatments on average produced herbage dry matter of 4773 kg ha^?1, the NG produced 4583 kg ha^?1 and the CG produced 3183 kg ha^?1 in year 4 (2005–06) of the experiment. Deferred grazing treatments with and without fertilizer application produced an average of 5135 and 4411 kg DM ha^?1 respectively. Averaged over 4 years, deferred grazing increased the germinable seed pool of perennial grasses by 200% and annual grasses by 50% (except optimized deferred grazing that considerably decreased the annual grass seed pool) compared with the CG. The best of the deferred grazing strategies increased the digestibility of pastures by 7% compared with the CG. The results demonstrated that deferred grazing from spring to autumn followed by rotational grazing could be an effective tool to increase herbage production and soil seed pool and improve the digestibility of native pastures in the steep hill country of southern Australia.     

The conservation management of mesotrophic (meadow) grassland in Northern England. 2. Effects of grazing, cutting date, fertilizer and seed application on the vegetation of an agriculturally improved sward

The plant species number and composition, iind yield of herbage biomass of an agriculturally improved hay meadow were assessed after 4 years under various combinations of grazing, fertilizer applicution. cutting date and seed addition treatments in a replicated split-plot design. Grazing treatments consisted of either autumn grazing with cattle and sheep, spring grazing with sheep or both regimes. Fertilizer application treatments consisted of either 25 kg ha^?1 N plus 12–5 kg ha^?1 P and K or no fertilizer. Cutting date treatments consisted of cuts on either 14 June, 2i July or 1 September. Seed addition treatments consisted of either no addition or sowing with a range of meadow species in the autumn. Data analysis was by correspondence analysis and analysis of variance. Species number decreased with fenilizer use and when the cutting date was 1 September. A range of species was affected by the main treatments and there were some first-order interactions, mainly between cutting date and fertilizer application. Rhinanthus minor was particularly favoured by the seed addition treatment. Species attnbutes in the regenerative and established phase were related to treatments and their effect on species composition. The National Vegetation Classification communities were associated with particular treatment regimes. The 21 July cutting date favoured ‘improved’ over ‘unimproved-traditional’ swards, with spring grazing favouring ‘unimproved-traditional’ swards. Lowest yields of herbage biomass were associated with autumn and spring grazing, the 14 June cutting date and no fenilizer treatments. The fenil-izer, 1 September cutting date and auiumn grazing treatments gave the highest yields. The implications of these results are discussed in terms of the conservation management required to return agriculturally improved mesotrophic grassland to a species composition similarto that of traditionally managed grassland.     

Effects of date of autumn closing and timing of winter grazing on herbage production in winter and spring

Growth of grass herbage in Ireland is highly seasonal with little or no net growth from November to February. As a result, feed demand exceeds grass supply during late autumn, winter and early spring. At low stocking rates [≤2 livestock units (LU) ha^?1], there is potential to defer some of the herbage grown in autumn to support winter grazing. This study examined the effects of four autumn‐closing dates and four winter‐grazing dates in successive years on the accumulation of herbage mass and on tiller density in winter and subsequent herbage production at two sites in Ireland, one in the south and one in the north‐east. Closing swards from grazing in early and mid‐September (north‐east and south of Ireland respectively) provided swards with >2 t DM ha^?1 and a proportion of green leaf >0·65–0·70 of the herbage mass above 4 cm, with a crude protein (CP) concentration of >230 g kg^?1 DM and dry matter digestibility (DMD) of >0·700. The effects of autumn‐closing date and winter‐grazing date on herbage production in the subsequent year varied between the two sites. There was no significant effect of autumn‐closing date in the north‐eastern site whereas in the south earlier autumn closing reduced the herbage mass in late March by up to 0·34 t DM ha^?1 and delaying winter grazing reduced the herbage mass in late March by up to 0·85 t DM ha^?1. The effects of later grazing dates in winter on herbage mass continued into the summer at the southern site, reducing the herbage mass for the period from late March to July by up to 2 t DM ha^?1. The effects of imposing treatments in successive years did not follow a consistent pattern and year‐to‐year variation was most likely linked to meteorological conditions.     

Effects of grazing by lambs in autumn on a red clover-perennial ryegrass sward

In 1976, 1977 and 1978 a red clover-perennial ryegrass sward was cut twice for silage, and in the autumn of 1976 and 1977 it was either grazed at low and high stocking rates, i.e. seventeen and thirty-four lambs per ha respectively, or was cut with a forage harvester. The effects of grazing on yield in the following year were examined. Herbage growth in the grazing period was slow and did not differ significantly between the treatments. The yield of silage dry matter taken in May and July was highest in ungrazed plots (9·8 and 8·1 t ha^-1 in 1977 and 1978 respectively) and lowest in plots stocked at the high rate (5·9 and 5·7 t ha^-1 in 1977 and 1978 respectively). The aftermath yield for grazing in 1977 was slightly but significantly greater on treatments grazed in the previous year compared with ungrazed treatments. Red clover content decreased markedly during grazing, the high stocking rate treatment containing 2·4% clover and the ungrazed treatment 57·3% clover. The high stocking rate treatment also had the lowest red clover content in the first silage cut. Red clover content in grazed plots increased to a level similar to that in ungrazed plots by the start of the grazing period in the subsequent harvest year. Animal performance was higher at the low than at the high stocking rate but herbage consumption per head did not differ significantly between the two grazing treatments. Possible reasons for the adverse effect of grazing on the red clover are defoliation and treading. It is concluded that such experiments can form the basis of an economic assessment of red clover and help the farmer decide whether or not he should integrate the crop into his system.     

Spring and autumn animal treading effects on pre‐grazing herbage mass and tiller density on two contrasting pasture types in Ireland

A perennial ryegrass (Lolium perenne L.)‐dominated sward on a well‐drained soil (Experiment 1) and a creeping bent (Agrostis stolonifera L.)‐dominated sward on a poorly drained soil (Experiment 2) were subjected to four treading treatments: control (C, no damage), light damage (L), moderate damage (M) or severe damage (S) to quantify the effects on herbage dry‐matter (DM) production and tiller density. In Experiment 1, treading damage was imposed in spring. In Experiment 2, one‐third of the site was damaged in autumn, one‐third in spring and one‐third in both spring and autumn. Both sites were rotationally grazed after treading treatments. Pre‐grazing herbage mass was measured eight times in Experiment 1 and seven times in Experiment 2 on each plot, and tiller density was assessed four times in each experiment. In Experiment 1, pre‐grazing herbage mass was reduced by 30% in S plots at the first harvest after damage, but cumulative pre‐grazing herbage DM production was not different between treatments (12·7 t DM ha^?1). In Experiment 2, annual cumulative pre‐grazing herbage mass was reduced by between 14 and 49%, depending on intensity of treading damage event and season when damage occurred. Tiller density was not affected by treatment in either experiment. A perennial ryegrass‐dominated sward on a well‐drained soil was resilient to heavy treading damage. A creeping bent‐dominated sward on poorly drained soil requires a more careful grazing management approach to avoid major losses in cumulative pre‐grazing herbage mass production during wet weather grazing events.     

The long-term effects of a range of pasture treatments applied to three semi-natural hill grassland communities. 1. Pasture production and botanical composition

Five pasture treatments: (1) controlled grazing, (2) controlled grazing + lime, (3) controlled grazing + lime + phosphate, (4) controlled grazing + lime + phosphate + oversown white clover and (5) controlled grazing + lime + phosphate + oversown white clover + oversown perennial ryegrass were applied to three semi-natural hill grassland communities. The communities were those dominated by Agrostis/Festuca, Molinia and Nardus. The proportion of Nardus at the Nardus-dominant site was substantially reduced by herbicide before treatments were applied. All treatments were grazed simultaneously by mature wether sheep on three occasions each year. There were two grazing periods each of 4 weeks duration between mid-May and mid-August with a further grazing period of 3 weeks starting mid-October. During each grazing period stock numbers were set so that a residual herbage mass of 560 kg DM ha ^?1 remained at the end of the grazing period. Measurements of net herbage accumulation (NHA) were made annually over a period of 13 years at each site. The green:dead ratio of grasses, species composition of the pasture and the pH of the soil were measured at intervals during the experiment. Estimates of mean annual NHA ranged from 3860 kg DM ha^?1 for treatment 1 (controlled grazing) to 5170 kg DM ha^?1 for treatment 5 (oversown white clover and perennial ryegrass). The application of lime and phosphate increased annual NHA by 300–350 kg DM ha^?1 with a further increase of around 400 kg DM ha^?1 when white clover was sown. Increases in NHA between year 1 and year 13 ranged from 30% for treatment 1 to around 55% for treatment 5. Although there was no difference in the mean NHA between sites, the herbage from the Agrostis/Festuca site contained a higher proportion of green grass and white clover than that from the other sites. The highest levels of green grass and white clover were found on the oversown treatments at each site. The grazing pressure exerted produced relatively little change in the botanical composition at the Agrostis/Festuca site. At the Molinia-dominant site the Molinia was largely replaced by Nardus during the first 6 years and Nardus also increased in cover at the Nardus site. Application of lime and phosphate generally increased the proportion of Poa pratensis, Festuca rubra and Agrotis tenuis but did not halt the spread of Nardus at the Molinia and Nardus sites. White clover and perennial ryegrass were successfully introduced by oversowing and proportions remained high throughout the 13 years. The cover by bryophyte spp. increased at all sites with the greatest increases occurring in each case on the less comprehensive pasture treatments. These results suggest that on Agrostis/Festuca-dominant pastures herbage biomass production can be increased with relatively low-cost pasture treatments while maintaining Species diversity. However, Nardus and Molinia dominant pastures are likely to require more comprehensive pasture treatments involving sown grasses and white clover to provide herbage of acceptable quality and avoid an increase in Nardus and bryophytes in the sward. With a regime of episodic summer grazing and the addition of fertilizers oversown pastures can be maintained over long periods of time.     

Effect of grazing severity on grass utilization and milk production of rotationally grazed dairy cows

Two experiments were carried out to examine the effects of grazing severity on the performance of January/February calved British Friesian dairy cows. In Experiment 1, three groups of cows were rotationally grazed across twenty-four one-day paddocks with high (H), medium (M) or low (L) herbage allowances. Cows on treatment M were offered a daily herbage allowance designed to achieve a residual sward height of 50 mm, assessed by a rising-plate sward stick. The daily herbage allowance (g organic matter (kg live-weight)^-1 on treatments L and H were 0.30 below and above that for M, respectively, to give residual sward heights of 42 and 59 mm. In Experiment 2, three groups of cows were grazed across twenty-four one-day paddocks to obtain residual sward heights of 50 mm (severe), 60 mm (moderate) and 80 mm (lax). Average milk yields on the L, M and H treatments in Experiment 1 were 11.8, 14.6 and 14.5 kg d,^-1 and in Experiment 2 they were 13.7, 16.0 and 17.0 kg d^-1 on the severe, moderate and lax treatments, respectively. The results indicate that the critical herbage height below which milk production per cow declines may vary with the production potential of the animal. There were no significant treatment effects on milk composition. Milk output ha^-1 and utilized metabolizable energy ha^-1 were greatest with the low herbage allowance in Experiment I and the moderate treatment in Experiment 2. Net herbage accumulation on the severe treatment in Experiment 2 was 30% lower than that on the lax treatment, as a result of treading damage in early season. It is concluded that, in a rotational grazing system, a reasonable compromise between sward utilization and animal performance can be achieved by grazing January/February calved cows to a residual sward height of 60 mm as assessed by a rising-plate sward stick. This is equivalent to a sward surface height of about 80 mm.     

The effect of initial spring grazing date and subsequent stocking rate on the grazing management, grass dry matter intake and milk production of dairy cows in summer

The objective of this study was to investigate the effects of an early (February; F) or delayed (April; A) primary spring grazing date and two stocking rates, high (H) and medium (M), on the grazing management, dry matter (DM) intake of grass herbage and milk production of spring‐calving dairy cows grazing a perennial ryegrass sward in the subsequent summer. Sixty‐four Holstein‐Friesian dairy cows (mean of 58 d in milk) were assigned to one of four grazing treatments (n = 16) which were imposed from 12 April to 3 July 2004. Cows on the early spring‐grazing treatment were grazed at 5·5 cows ha^?1 (treatment FH) and 4·5 cows ha^?1 (treatment FM) while cows on the late‐grazing treatment were grazed at 6·4 cows ha^?1 (treatment AH) and 5·5 cows ha^?1 (treatment AM). The organic matter digestibility and crude protein concentration of the grass herbage were higher on the early‐grazing treatment than on the late‐grazing treatment. The cows on the FM treatment had significantly (P < 0·001) higher milk (24·5 kg), solids‐corrected milk (22·5 kg), fat (P < 0·01, 918 g) and protein (831 g) yields than the other three treatments. Cows on the FM treatment had a higher (P < 0·001) DM intake of grass herbage by 2·3 kg DM per cow per day than cows on the AH treatment, which had a DM intake significantly lower than all other treatments (15·2 kg DM per cow per day). The results of the present study showed that grazing in early spring has a positive effect on herbage quality in subsequent grazing rotations. The study also concluded that early spring‐grazed swards stocked at a medium stocking rate (4·5 cows ha^?1; FM) resulted in the highest DM intake of grass herbage and milk production.     

The effect of strategic use of fertilizer nitrogen in spring and/or autumn on the productivity of a perennial ryegrass/white clover sward

The productivity of a mixed sward, comprising perennial ryegrass cvs Barlano and Bastion and white clover cvs Donna and Aran, was measured under sixteen fertilizer N treatments. These involved 0.25, 50 and 75 kg N ha^-1 in spring only, in autumn only and in all combinations of spring N and autumn N. A simulated grazing regime of six cuts annually at 3- to 6-week intervals was imposed. Increasing rates of total N application increased total herbage DM regardless of application pattern. Yield response was greater with N applied in the spring, and total herbage DM was higher with high spring N-low autumn N than the reverse. Mean yield responses at the first harvest to 25, 50 and 75 kg ha^-1 N in spring were 13.6, 10.8 and 11.6 kg DM per kg N. Corresponding responses at the final harvest to N rates in the autumn were 7.2, 5.8 and 6.8 kg DM per kg N. Responses were similar at these times for treatments receiving combined spring and autumn N. Over all treatments, mean annual production of total herbage was between 7.08 t ha^-1 DM with no N and 8.19 t ha^-1 with 75 kg ha^-1 N in both spring and autumn. Owing to drought, mean production in year 2 fell by 32% compared with year 1. White clover production fell progressively with increasing N application. Treatments with spring-applied N gave the most marked decrease. White clover was more markedly depressed than the associated grass by the drought in the second year. The mean reductions in white clover content were 0.17, 0.07 and 0.12 percentage units per kg applied N for spring N, autumn N and combinations. Autumn N use depressed white clover less than spring N but the yield response of grass was less. It is concluded that any applied N adversely affects white clover performance to some degree. Where management factors are unfavourable to white clover even strategic N use may not be wise. Instead, it is suggested that a ‘dual-sward’ approach be adopted in practice, namely, grass/white clover swards with no N. and complementary grass swards receiving optimum applied N to give better production at times when grass/white clover swards are relatively less productive.     

The effects of autumn closing date on sward leaf area index and herbage mass during the winter period

The provision of grass for early spring grazing in Ireland is critical for spring calving grass‐based milk production systems. This experiment investigated the effect of a range of autumn closing dates (CD), on herbage mass (kg DM ha^?1), leaf area index (LAI) and tiller density (m^?2) during winter and early spring. Thirty‐six grazing paddocks, closed from 23 September to 1 December 2007, were grouped to create five mean CD treatments – 29 September, 13 October, 27 October, 10 November, 24 November. Herbage mass, tiller density and LAI were measured every 3 weeks from 28 November 2007 to 20 February 2008; additionally, herbage mass was measured prior to initial spring grazing and tiller density was measured intermittently until September 2008. Delaying CD until November significantly (P < 0·05) reduced herbage mass (by approximately 500 kg DM ha^?1) and LAI (by approximately 0·86 units) in mid‐February. On average, 35% of herbage mass present on swards on 20 February was grown between 28 November and 30 January. LAI was positively correlated with herbage mass (R² = 0·78). Herbage mass increased by approximately 1000 kg DM ha^?1 as spring grazing was delayed from February to April. Tiller density increased from November to February, although it did fluctuate, and it was greatest in April (9930 m^?2). This experiment concludes that in the south of Ireland adequate herbage mass for grazing in early spring can be achieved by delaying closing to early mid‐October; swards required for grazing after mid‐March can be closed during November.     

Soil chemical properties,plant species composition,herbage quality,production and nutrient uptake of an alluvial meadow after 45 years of N,P and K application

Little is known about the long-term effects of mineral N, P and K application on the nutritional status of mown alluvial grasslands. We asked how long-term fertilizer application affected soil chemical properties, plant species composition, herbage production, nutrient concentrations in soils and plants and balance of nutrients. Six treatments (control, PK, N50PK, N100PK, N150PK and N200PK) were investigated at the Černíkovice Experiment (Czech Republic) established in 1966 on an Alopecurus pratensis meadow, using annual application rates of 50, 100, 150 and 200 kg N, 40 kg P and 100 kg K ha⁻¹. Data were collected and analysed for 2007, 2008 and 2009. Although fertilizers had been applied over 45 years, differences in soil chemical properties between fertilization treatments were small. The legumes Lathyrus pratensis and Trifolium repens responded highly positively to PK application, and tall grasses, A. pratensis in particular, to NPK application. Herbage quality was high in terms of content of major nutrients, and its chemical properties varied considerably between treatments, cuts and years. Mean annual herbage yield ranged from 6·1 in the control to 9·7 t ha⁻¹ in the N200PK treatment. Herbage production was N-limited in 2007 and 2009, but not in 2008. Seasonal N agronomical efficiency ranged from 4·2 to 22·9 kg of DM herbage per kg of applied N. The herbage N:P and N:K ratios did not reflect the actual response of herbage production to N application. A negative balance between N applied and N removed in harvested herbage was recorded in all treatments. We concluded that in highly productive alluvial grasslands, mineral-rich soils can respond weakly to N, P and K application, fertilizer application modifies plant species composition and herbage production is not N-limited in all years. Nutrient ratios must be interpreted with caution for the estimation of nutrient limitation.     

Effect of replacing time available for grazing with time available for eating maize silage and soyabean meal on milk yield and feeding behaviour in dairy cows

Low rates of herbage dry matter (DM) intake impose limits on total daily DM intake in grazing dairy cows. The objective of this study was to increase total daily DM intake and milk production by restricting daily time available for grazing (TAG) and replacing it with time available for eating a maize silage/soyabean meal (TAMS) diet indoors. The treatments (TAG + TAMS) were 20 + 0, 19 + 1, 10 + 10 and 5 + 15 h. Measurements were made of milk production, intake and feeding behaviour. The interactions of TAG + TAMS treatments with sward height (SH) and concentrate level (CL) were also examined. Two experiments, each lasting 42 days, were carried out in spring ( Experiment 1 ) and autumn ( Experiment 2 ) using forty‐eight and twenty‐four Holstein‐Friesian cows respectively. Treatments were arranged in a factorial design with TAG + TAMS treatments, SH ( Experiment 1 only) and CL as the independent variables and a TAG + TAMS of 20 h. Reducing TAG and increasing TAMS significantly reduced estimated herbage DM intake and significantly increased maize silage/soyabean meal intake in both experiments, but there were no significant main effects of TAG + TAMS treatments on milk yield (mean, 27·4 and 25·5 kg d^?1 for Experiments 1 and 2 respectively), and yield of milk constituents. Increasing SH ( Experiment 1 ) and CL ( Experiments 1 and 2 ) significantly increased milk yield. In Experiment 1 , there was a significant interaction between TAG + TAMS treatments and SH with the taller sward height of 8–10 cm and the 20 + 0 treatment having the highest milk yield (29·7 kg d^?1) and the 5 + 15 treatment the lowest (27·2 kg d^?1), whereas at the lower sward height of 4–6 cm, milk yield was lowest on the 20 + 0 treatment (25·5 kg d^?1) with the other three treatments being higher (mean, 26·9 kg d^?1). Replacing TAG with TAMS significantly increased liveweight gain in Experiment 1 but not in Experiment 2 . Estimated rates of intake of herbage were lower in the autumn experiment ( Experiment 2 , 9·6 g DM min ^?1) than in the spring experiment ( Experiment 1 , 29·4 g DM min ^?1) but rates of intake of maize silage were higher in the autumn (112·4 g DM min^?1) than in the spring (72·5 g DM min^?1). In conclusion, in spring the response to replacing TAG with TAMS was dependent on sward conditions with the highest milk fat plus protein yield being on the 20 + 0 treatment at the high sward height and on the 19 + 1 treatment at the low sward height. The high liveweight gain of the 5 + 15 treatment could be an important means of restoring body condition in grazing lactating cows. In autumn, intakes of herbage were low in spite of its high estimated nutritive value with all treatments having a similar level of performance.     

Defoliation patterns and their implications for the management of vegetative tropical pastures to control intake and diet quality by cattle

This study assessed the use of pasture attributes to control daily intake and diet quality during progressive defoliation on pastures of Axonopus catarinensis. Three consecutive 12‐day grazing treatments of progressive defoliation were conducted with Brahman cross‐steers. Daily forage intake and defoliation dynamics were assessed using a pasture‐based method. The treatments differed in initial sward height (33, 44 and 61 cm) and herbage mass (1030, 1740 and 2240 kg ha^?1). The post‐grazing residual sward height, at which forage intake decreased, appeared to increase with the initial sward height (12·3, 14·6 and 15·5 cm). Steers grazed up to four distinctive grazing strata in all treatments. The depth and herbage mass content of the top grazing stratum were at least five times higher than the lower grazing strata in all treatments. This explains why forage intake decreased when the top grazing stratum was removed in approximately 93% of the pasture area in all treatments, equivalent to approximately 7% of the pasture area remaining ungrazed. We conclude that the residual ungrazed area of the pasture, rather than residual sward height, can be used to develop grazing management strategies to control forage intake and diet quality in a wide range of pasture conditions.     

Modelling pasture management practices for soil organic carbon gain in livestock systems

Our ability to develop strategies to mitigate climate change includes an understanding of, and our capacity to predict soil organic carbon (SOC) dynamics in livestock systems. Here we assess the capability of the Sustainable Grazing System (SGS) Pasture Model for predicting pasture growth (elongated wheatgrass, Thinopyrum ponticum) and SOC accumulation in different environments and under a range of pasture management practices in hydrohalomorphic soils located in South-eastern Buenos Aires Province, Argentina. After Model calibration, aerial net primary productivity (ANPP) and TSOC content under two grazing intensities (7.5 and 11 cm post-grazing target heights) and two N fertilization levels (0 and 100 kg N ha⁻¹ yr⁻¹) were simulated over a 10 year-period. The SGS Pasture Model predicted 87% of the observed ANPP, with observed and predicted ANPPs averaging 1.46 and 1.42 Mg ha⁻¹ yr⁻¹, respectively. There were differences in simulated ANPP between fertilized and unfertilized treatments both at high and low grazing intensities for the last year of the period. Total SOC contents from the modelling showed differences between high (83.7 to 84.2 Mg ha⁻¹) and low (86.8 to 87.5 Mg ha⁻¹) grazing intensities, with treatments receiving N also showing higher carbon stocks. The positive effect of reduced grazing intensity on soil carbon was explained by an increased input of aerial and subterranean dry matter into the soil. Sensitivity analysis showed that SGS is a robust model, capable of performing effectively under a variety of conditions. Hence, it can be used for exploring management practices to mitigate the impact of livestock systems on emissions and SOC stocks.     

Productivity,grazing utilization,forage quality and primary production controls of species‐rich alpine grasslands with Nardus stricta in northern Spain

Species‐rich alpine grasslands with Nardus stricta are important communities for both animal production and environmental conservation in Europe. We selected two contrasting types of Nardus grasslands (mesic and wet) within a rangeland of northern Spain and measured annual above‐ground net primary productivity (ANPP), botanical components, forage utilization and their respective seasonal patterns, during a 5‐year period. We analysed their chemical properties and recorded soil moisture and temperature in order to construct models able to explain grassland productivity. Mean annual ANPP of mesic Nardus grassland was about half (216 g DM m^?2 year^?1; ±29·8 s.e.) that of the wet grassland (406 g DM m^?2 year^?1; ±54·3 s.e.), with significant intra‐ and interannual variability. Mesic grassland, with a more important contribution of forbs and legumes over graminoids in its botanical composition, was the preferred forage source of grazing livestock and showed better chemical properties in spring and early summer. In summer and autumn, wet grassland had a higher utilization owing to its ability to maintain high biomass production. This was partially explained by soil moisture, a limiting factor of mesic grassland productivity. Our results provide new and relevant information on key aspects of species‐rich alpine Nardus grasslands, potentially useful for the definition of management options for these habitats of priority conservation.     

Setting management limits for the production and utilization of herbage for out-of-season grazing

Three experiments were carried out on perennial ryegrass‐dominant swards to provide a basis for recommendations for the limits to (a) building up and timing of utilization of a herbage ‘bank’ for out‐of‐season grazing and (b) duration and intensity of early spring grazing in the United Kingdom and Ireland. In experiment 1, the effect of regrowth interval (from 7 September, 20 October, 17 November or 15 December) in autumn on herbage accumulation, leaf turnover and on subsequent spring growth was investigated. Swards regrown from early September reached maximum herbage mass (about 3 t ha^–1 DM) and leaf lamina content in mid‐November, by which time senescence rate exceeded rate of production of new leaves. New leaf production and senescence rates were greater in swards remaining uncut until December than in those cut in October or November. Time of defoliation up to December had no effect on spring herbage mass in the subsequent spring. Defoliating in March reduced herbage mass in late May by less than 20%. Experiment 2 investigated the progress in herbage growth and senescence in swards regrowing from different times in late summer and autumn to produce herbage for utilization beyond the normal grazing season. Treatments in a randomized block design with three replicates were regrowths from 19 July, 8 August, 30 August and 20 September. Based on a lower ceiling of leaf and total herbage mass being reached with progressively later regrowths, beyond which leaf senescence generally exceeded leaf production and herbage mass declined, it was concluded that currently recommended rotation lengths for this period should extend from 3 weeks in late July to 8 weeks for swards previously grazed in mid‐September. In both experiments, leaf senescence commenced earlier (by one leaf‐age category) than previously published estimates and so brought forward the time at which senescence rates balanced leaf growth rates. In experiment 3, designed to evaluate the effect of daily grazing period and intensity in early spring on herbage regrowth, dairy cows grazed successive plots (replicates) for 2 or 4 h each day at two intensities (target residual heights of 5 or 7 cm) in March to mid‐April. Regrowth rate was similar in all treatments including the ungrazed control, despite soil moisture content being relatively high on occasions. Tiller density was significantly reduced in May by grazing plots in early or mid‐April. It is concluded that in autumn there are limits to which rotation lengths should be extended to produce herbage for out‐of‐season grazing owing to attainment of ceiling yields. Although utilization in early spring may reduce herbage availability in spring, out‐of‐season utilization need not reduce herbage growth rates in early spring.     

Selective defoliation by sheep according to slope and plant species in the hill country of New Zealand

The objective of the study was to evaluate the grazing behaviour by sheep in hill country paddocks in New Zealand which had received two long‐term fertilization and stocking rate treatments [high fertility–high stocking rate (HH); low fertility–low stocking rate (LL)]. Herbage accumulation and selective grazing were evaluated within low slope (LS), medium slope (MS) and high slope (HS) categories. Transects lines were placed and tillers of Agrostis capillaris and Lolium perenne in the LS category; A. capillaris, Anthoxanthum odoratum and L. perenne in the MS category; and A. capillaris and A. odoratum in the HS category were marked. The leaf length of each marked tiller was measured and used to determine selective grazing over 3 weeks during each season. The highest herbage accumulation rates were during spring and the lowest in summer and winter. The LS category showed the highest herbage accumulation rates and HS the lowest. Except for autumn, the marked tillers were more frequently grazed in the HH than in the LL paddock. During summer, autumn and spring, grazing frequency in the slope categories was in the order LS > MS > HS. During winter sheep did not discriminate between slope category. During summer, autumn and spring, sheep did not selectively graze the species studied but this was not the case during winter. Overall, sheep selectively grazed L. perenne. In all the seasons L. perenne consistently had the longest leaves but within species there was no consistent relationship between leaf length and probability of being grazed. Selective grazing changed through the year according to herbage accumulation rate. Sheep concentrated grazing in the category LS when herbage accumulation rate was high, but they did not discriminate between slope categories in winter when herbage accumulation rate was low.     

The long-term effects of a range of pasture treatments applied to three semi-natural hill grassland communities.

Five pasture treatments were applied to three semi-natural hill grassland communities. The pasture treatments were: (1) controlled grazing, (2) controlled grazing + lime, (3) controlled grazing + lime + phosphate, (4) controlled grazing + lime + phosphate + oversown white clover and (5) controlled grazing + lime + phosphate + oversown white clover + oversown perennial ryegrass. The communities were dominated by Agrostis/Festuca (site 1), Molinia (site 2) and Nardus (site 3). The Nardus at site 3 was substantially reduced by herbicide before treatments were applied. All treatments were grazed simultaneously by mature wether sheep on three occasions each year to a residual herbage mass of 560 kg DM ha^?1. There were two grazing periods, each of 4 weeks duration, beginning in mid-May and mid-July respectively and a further grazing period of 3 weeks duration beginning in mid-October. The number of sheep grazing days and liveweight gain of sheep were recorded during each grazing period for 13 years at each site. Organic matter intake (OMI) and digestibility (OMD) of ingested herbage were measured in years 2, 5 and 13. Mean daily OMI per head ranged from 1290 g for treatment 1 to over 1400 g for treatments 4 and 5. OMI values were higher for the Agrostis/Festuca site (mean OMI 1450 g) than for the Molinia and Nardus sites (1310 g and 1370 g respectively) largely owing to differences in the values for treatments 1, 2 and 3. OMI values for each treatment decreased by around 300 g between May and October. Mean OMD values from treatments 1, 2 and 3 were higher for the Agrostis/Festuca site (0·66) than for the Molinia and Nardus sites (0·63 and 0·64 respectively). The OMD values for treatments 1, 2 and 3 at the Molinia and Nardus sites declined by between 0·05 and 0·02 between years 2 and 13 and also showed the greatest decrease between May and October (0·08). OMD values for treatments 4 and 5 (0·67 and 0·69 respectively) were higher than for treatments 1, 2 and 3 at all sites and levels were maintained over 13 years. Individual liveweight gains of sheep increased from treatment 1 (?18 g d^?1) to treatment 5 (82 g d^?1), but they decreased over time on all treatments. The Agrostis/Festuca site gave 17% more annual sheep grazing days than the Molinia site and 33% more than the Nardus site. The mean number of annual sheep grazing days for each treatment ranged from 2250 for treatment 1 to 3640 for treatment 5. Annual sheep grazing days increased over 13 years by between 35 and 45% for treatment 1 and 55 and 70% for treatment 5. The results are described in relation to the changes in pasture composition and herbage accumulation.     

Effect of cultivar and seed rate of perennial ryegrass and strategic fertilizer nitrogen on the productivity of grass/white clover swards

Four cultivars of perennial ryegrass (intermediate diploid cv. Talbot and tetraploid cv. Barlatra, and late diploid cv. Parcour and tetraploid cv. Petra) were each sown at 10,20 and 30 kg ha^-1, all with 3 kg ha^-1 of white clover cv. Donna. Herbage productivity was measured over 3 harvest years, 1982–84. under two annual rates of fertilizer N (0 and 150 kg ha^-1); the 150 kg ha^-1 rate was split equally between March and August applications. Fertilizer N increased total herbage DM production; the 3-year means for the 0 and 150 kg ha^-1 N rates were 8·04 and 8·91 t ha^-1, respectively. In successive years, total herbage responses to N (kg DM (kg N applied)^-1) were 6·6, 35 and 72 (overall mean, 58). Mean white clover DM production over the 3 years was reduced from 4·48 t ha^-1 at nil N to 2·82 t ha^-1 at the 150 kg ha^-1 rate, a fall of 37%. Grass seed rate did not influence total herbage production or white clover performance. The two intermediate perennial ryegrass cultivars had a marginal advantage in total herbage production over the two late cultivars, but white clover content and production were higher with tetraploids than diploids. It is concluded that the value of increased herbage production from strategic use of fertilizer N has to be weighed against its depressive effect on white clover performance; application of 75 kg ha ha^-1 N in both spring and autumn was excessively high if maintenance of a good white clover content in the sward is an objective. There is considerable flexibility in the grass: clover seed ratio in seeds mixtures. Modern highly-productive perennial ryegrass varieties do not differ substantially in compatibility with white clover but tetraploids permit better clover performance than diploids.