Genetic and environmental control of crown development in Picea sitchensis and its relation to stem wood production

Differences in stem wood production were found among four clones of Picea sitchensis (Bong.) Carr. at both an agricultural and a forest site. Clonal rankings were not consistent between the sites. Four variables were identified that may influence stem wood production, foliage amount, total aboveground production per unit foliage, proportional allocation of production to new foliage and to branch wood thickening. Comparison of clonal performance between sites showed that stem wood production could be influenced by differences in each of these variables. The four variables were themselves determined by components of crown structure, these were (i) production of new branches from the main stem, (ii) the numbers and lengths of branchlets supported on branches from the main stem, (iii) foliage production per unit branchlet length and foliage longevity, and (iv) branch wood thickening per unit branch length. These components varied both among clones and between sites. Branch production from the mainstem varied among clones, and for some clones it varied substantially between sites. In some clones branch production was positively correlated with mainstem height increment. There were differences among clones in the way that branchlet production varied between the agricultural and forest sites. In the absence of needle fall, foliage weight/branch length varied threefold among clones, but for each clone, varied little between sites. Branch wood weight/branch length was generally greater when foliage weight/branch length was large, but total branch wood increment/tree was strongly influenced by total branch length. The dynamics of crown development and its relation to stem wood production are discussed with reference to (i) the importance of needle longevity, (ii) the importance of tallness and narrowness of crowns and (iii) the importance of branch thickening, a process that competes with stem growth in the utilization of photosynthate. It is concluded that although each of these factors may be important, none singly accounts for the observed differences in stem wood production among genotypes and the way these change in response to the environment.     

A technique to identify annual growth rings in Eucalyptus grandis usingannual measurements of diameter at breast height and gamma raydensitometry

Many eucalypt species do not show distinct growth rings because cambial activity does not show a strong response to seasonal variation in climate. Eucalyptus grandis, one of the most important commercial hardwood species in South Africa, is one such example of a species that does not have well-defined growth rings. The light and dark bands visible on the cross-section of the wood of E. grandis do not always correspond with the growing season, which makes it difficult to resolve wood property data into annual increments. A method was developed to separate growth rings on wood density profiles of E. grandis into annual increments using bark-to-pith density profiles and annual measurements of diameter at breast height (DBH) from permanent sample plot (PSP) datasets. Using the PSP data, it was possible to assess the annual pattern of stem diameter growth at a compartment level by calculating the radial increment (RI) per year and expressing that value as a percentage of the radius at the end of the increment for that year. Mean radial increment percentage (%MRI) was calculated for each year and used to predict annual RI at an individual tree level. Predicted RI values for each tree were expressed as cumulative distances from the bark end and superimposed onto their respective density profiles. Predicted RI corresponded well with latewood density peaks and these separation points were considered a reliable guide to divide the density profile into annual increments closer to the bark end and into broader age classes closer to the pith. By assessing the pattern of variation in radial density within the context of the growth history of a compartment by means of annual PSP data, it was possible to confirm that growth rings on density profiles of E. grandis closer to the bark end can serve as a reliable representation of annual growth.     

毛白杨优良无性系(新品种)材性测定研究

按生长性状预选出毛白杨１３年生１０个无性系用于材质测定。试材取自北方大兴县测定林,每无性系在３个重复中各抽样１株,伐倒取样,分析表明,毛白杨无性系间纤维长度,宽度和长宽比差异极显著,重复率依次为０．７７７,０．５３７和０．６７６;木材全年密度,早材密度和晚材密度差异极显著,重复率依次为０．５３６,０．５１４和０．４２９。     

Radial stem growth in response to microclimate and soil moisture in a drought-prone mixed coniferous forest at an inner Alpine site

Dendroclimatological studies in a dry inner Alpine environment (750 m a.s.l.) revealed different growth response of co-occurring coniferous species to climate, which is assumed to be caused by a temporal shift in wood formation among species. The main focus of this study therefore was to monitor intra-annual dynamics of radial increment growth of mature deciduous and evergreen coniferous species (Pinus sylvestris, Larix decidua and Picea abies) during two consecutive years with contrasting climatic conditions. Radial stem growth was continuously followed by band dendrometers and modelled using Gompertz functions to determine time of maximum growth. Histological analyses of tree ring formation allowed determination of temporal dynamics of cambial activity and xylem cell development. Daily fluctuations in stem radius and radial stem increments were extracted from dendrometer traces, and correlations with environmental variables were performed. While a shift in temporal dynamics of radial growth onset and cessation was detected among co-occurring species, intra-annual radial growth peaked synchronously in late May 2011 and early June 2012. Moist atmospheric conditions, i.e. high relative air humidity, low vapour pressure deficit and low air temperature during the main growing period, favoured radial stem increment of all species. Soil water content and soil temperature were not significantly related to radial growth. Although a temporal shift in onset and cessation of wood formation was detected among species, synchronous culmination of radial growth indicates homogenous exogenous and/or endogenous control. The close coupling of radial growth to atmospheric conditions points to the importance of stem water status for intra-annual growth of drought-prone conifers.     

Effect of complete competition control and annual fertilization on stem growth and canopy relations for a chronosequence of loblolly pine plantations in the lower coastal plain of Georgia

Stem growth, developmental patterns and canopy relations were measured in a chronosequence of intensively managed loblolly pine stands. The study was located on two distinct sites in the lower coastal plain of Georgia, USA and contained a factorial arrangement of complete control of interspecific competition (W) and annual nitrogen fertilization (F). The W treatment increased growth rate for several years, while the F treatments led to sustained growth increases. The combination of the W and F treatments resulted in more than 180 Mg ha⁻¹ stem biomass production at age 15 which is more than double the production of control treatment. Stem biomass production is continuing to increase through age 15 as indicated by the current annual increment in stem biomass continuing to exceed the mean annual increment in stem biomass. The F treatment decreased wood quality by decreasing whole tree latewood specific gravity from 0.565 to 0.535 and by lengthening the transition from juvenile to mature wood from 4 to 5 years. Increased rates of stem growth in response to cultural treatments were largely mediated by increased leaf area, with strong functional relationships between leaf area index and current annual increment. However, growth efficiency (stem production per unit of leaf area) decreased with stand age. These results indicate that nutrient amendments are necessary for sustaining high rates of stand development on relatively nutrient poor lower coastal plain soils.     

Growth and thinning effects during a rotation period of hybrid aspen in southern Sweden

Hybrid aspen is an interesting tree species for wood production in northern Europe. In this study we examined growth dynamics over the whole rotation period. Height and diameter development, as well as annual growth of stem volume and stem biomass, were repeatedly recorded in 14 planted and 2 root sucker stands, aged up to 26 years, in southern Sweden. A main aim was to study the productivity level for hybrid aspen forestry with an expected rotation period of about 25 years. The study verified earlier prognoses, showing a mean annual increment (MAI) of 19.5 m³ of stem wood ha^?1 yr^?1 after 25 years. This corresponds to a dry weight of stem biomass of 7.2 tons dry matter (DM) ha^?1 yr^?1. Maximum MAI was still not reached after 25 years, although the growth curve was flattening out. If branch biomass is included, MAI is estimated to about 9 tons DM ha^?1 yr^?1 and further improvements in growth is expected by using the best genotypically selected clones available. Results from three different thinning regimens showed that thinning intensity provided significantly larger diameter growth, while no significant yield effects were seen among the thinning regimens.     

尖峰岭绿楠树生长过程的研究

对海南尖峰岭地区３０年生的热带乡土树种———绿楠人工林进行了样地调查,并进行了样木横剖树干解析,研究其生长规律及与环境的关系。研究结果表明：（１）绿楠是比较速生的树种,地径、胸径、树高和材积的年生长都呈多峰态曲线;（２）胸径年生长高峰值出现在第１１年,而树高年生长量高峰期晚于地径和胸径生长高峰值出现的年份;（３）在生长初期（８ａ前）,材积生长极缓慢,年生长量＜００００９ｍ３,其后开始较快增长,到第１７年出现最大值０００８６８ｍ３;（４）从平均生长量看,地径变化比较平缓;胸径在１４ａ前呈稳态上升趋势,１４ａ后呈平稳发展状态并略有下降;树高在１９ａ前基本呈稳态上升趋势,１９ａ后保持在０５０ｍ／ａ左右,并呈缓慢地下降趋势;材积一直是随着年龄的增加而增加;（５）不同坡位的水肥等条件不同,因而影响了绿楠树的生长,离水沟最近的下坡地的树高和胸径比上坡分别大３７５％和２９１％;（６）绿楠树地径、胸径和树高的连年生长变化趋势基本上与降雨蒸发比（Ｒ／Ｅ）变化趋势相一致,说明水分状况是影响绿楠生长的一个重要环境因子。     

Seasonal patterns of radial root growth and starch dynamics in plantation-grown Sitka spruce trees of different ages

Seasonal patterns of radial root growth within 1 m of tree stems were examined in Scottish plantations of Sitka spruce trees aged 9, 15 and 20 years. Results were compared with parallel measurements of shoot extension, radial growth of stems and amounts of starch stored in tissues external to root wood. Youngest trees produced the largest annual increments in root cross-sectional area and numbers of new cells along radial files of tracheids. Irrespective of tree age, new cells were present in roots before bud burst and the onset of radial growth occurred progressively later with increasing distances from the stems. At ages 15 and 20, both stem cross-sectional area and radial root growth up to 0.5 m from the stem base had a minor peak of activity preceding and a major peak following shoot elongation. Further than 0.5 m from the stem, root growth was frequently restricted to the period following shoot extension. Starch storage in the roots reached a maximum in April and May, which was greatest for 9-year-old trees and least for 20-year-old trees. At all ages, radial root growth in early spring occurred concurrently with increased starch storage. Later in the season starch reserves declined rapidly during the period of shoot elongation and root growth occurred whilst reserves were low. At all ages for positions on the root at the base of the stem and 0.25 m from it, starch depletion, at its maximum rate during June, accounted for less than the measured increment of root wood growth at that point. This indicates a substantial translocation of substrates to these zones during growth. At the same time, the reduction in starch concentrations at more distal points from the stem far exceeded that required for local root thickening.     

红豆树等6种珍贵用材树种的生长特性和材性分析

利用浙江龙泉和庆元两地在较好立地上营造的21 36年生红豆树、江南油杉、伯乐树、闽楠、刨花楠和乐东拟单性木兰6种珍贵用材树种的片林,以研究其生长、干形和木材基本密度的变化规律。研究结果表明,红豆树是一个树高和胸径生长量大、径生长速生持续期长、木材基本密度中等及其径向均匀性较高的珍贵用材树种,平均年轮宽度达到0.8 1.2 cm,36年生时按宽度和面积计算心材比例分别为60.57%和37.47%。因红豆树分叉干特性明显,在栽植的第2年就应及时修枝抹芽和施肥,以培育树干通直、心材比例高的优质干材。江南油杉和闽楠等其它5种珍贵用材树种皆较少分叉干,其树干通直。闽楠以材质优良而闻名,木材密度中等,虽然其生长速度较慢,但生长势很强,年生长量稳定,加之树冠窄小,适宜长周期大径材的培育;伯乐树生长速度中等,木材密度略低,但其径向均匀性较高,宜作为优质的工艺材来培育;树冠窄小的乐东拟单性木兰生长速度中等,但其木材密度大、径向均匀性高,是优良的珍贵用材树种;江南油杉是个早期生长快而后期生长慢,木材密度中等的针叶树种,可通过加强经营管理延长速生期和提高木材密度的径向均匀性;刨花楠虽然生长速度中等,但其木材密度却较低,浸水有粘液,不宜作为珍贵用材树种培育。     

马尾松纸浆材重要经济性状配合力及杂种优势分析

报道了马尾松不同种源优良无性系 6× 6全双列交配的 7a生遗传测定林的分析结果。研究发现 ,树高、胸径和材积生长、干物质积累及木材基本密度等纸浆材主要经济性状的一般配合力 (GCA)和特殊配合力 (SCA)效应极为显著 ,但显性基因效应起主导地位 ,加性基因效应次之。基因上位效应则因性状而异 ,仅对木材基本密度作用显著 ,而对树高、胸径、材积生长和干物质积累作用较小或可忽略。马尾松生长和材质材性指标的遗传控制模式不同于玉米、杉木等研究结果 ,这可能与交配亲本来源于不同地理种源及经过一般配合力间接选择有关。马尾松生长性状普遍存在自交衰退现象 ,一般配合力小的亲本自交衰退程度有增加的倾向。马尾松不同交配组合间的杂种优势差异巨大 ,正反交杂种优势在总体上大致相似。研究虽然发现一般配合力较高的亲本间杂交在材积增加和干物质积累等性状上会产生强杂种优势的倾向 ,却无明显的规律可寻。以干物质指数作为主要筛选指标 ,初选出 8个优良杂交组合     

桉树多树种无性系在川南地区生长比较

为选择出适合在川南地区造林的桉树无性系,对17个桉树新无性系在川南地区的宜宾市进行了造林对比试验。结果表明：以 SC-GU2、SC-GU1和 GR-6三个无性系生长表现好,4年生的胸径、树高、单株材积和蓄积均高于对照 DH32-29,其中单株材积分别比 DH32-29高出9.51%、4.53%和4.94%;其次是 G5,在川南地区也生长较好,生长量仅次于 DH32-29。4个桉树无性系 SC-GU2、SC-GU1、GR-6、G5的胸径、树高、单株材积和蓄积均均极显著高于5个赤桉无性系(H1,、H2,、H3、 H4和 H10),年均胸径、树高、蓄积分别达3 cm、4 m、31.72 m3·hm-2以上,表现了速生丰产的特性,可作为下一批优良桉树无性系在川南地区推广造林。     

Weather factors controlling growth of Oriental beech are on the turn over the growing season

A better understanding of the ecophysiological basis of wood formation by monitoring radial growth over the whole vegetation period may help to explain possible discrepancies between long-term average climate–growth relationships and short-term climatic impacts on tree growth. To understand how growth–climate associations of Oriental beech vary throughout a vegetation period, we studied seasonal growth patterns of high-elevation beech trees growing in the north of Iran by collecting wood anatomical micro-cores in 10 to bi-weekly intervals and measuring stem increment with high-resolution electronic dendrometers. Wood formation was for two consecutive years with contrasting inter-annual climate conditions (2011 and 2012). We divided the growing period into three equal time phases and related daily climate variability to the cambial growth in each phase. The pattern of climate–growth relationships varies over a complete growing season and between years: in both study years, trees responded homogenously at the beginning and at the end of the growing season, but showed opposing influence of relative humidity during spring and early summer (June and July). Temperature as the main driver of xylogenesis had a stimulating effect on growth at the beginning of the growing season, but had negative effects on radial increment during late June and July mainly due to the excess of ambient temperatures over an optimum threshold. Higher temperature in late June and July 2011 compared with those of 2012 led to a significantly narrower tree ring in 2011, despite the similar sums/means of annual climate parameters in these two years. Since regional climate change scenarios expect higher temperature and reduced moisture conditions in future, radial growth of beech in the north of Iran may be adversely affected.     

Production of hybrid aspen on agricultural land during one rotation in southern Sweden

High potential productivity together with short rotation periods have made hybrid aspen an interesting option for wood production on former arable land in Nordic countries.In this study,some of the oldest active experimental plots with hybrid aspen in Sweden were remeasured at 23-30 years of age.A main aim was to assess age and productivity at the time of maximum mean annual volume increment.In addition,the influence of commercial thinning on stand development and differences in genetic gain among clones were investigated.Data from five experiments in southern Sweden were used,including three genetic trials,one demonstration stand with a clone mixture and one stand regenerated from root suckers.The three genetic trials were treated as single plot experiments,subject to a standard thinning program.In the remaining experiments,different thinning strategies were tested in a balanced block design.Volume growth had culminated or was close to maximum at age25-30 years.Mean annual stem volume increment at culmination was 20-22 m~3 ha~(-1) a~(-1).Dominant height reached30-35 m at 28-30 years of age.Mean diameter at breast height was 27-29 cm after 29-30 years in the genetic trials.Clonal ranking based on diameter at age 7-9 years was positively correlated with the ranking at the final measurement in the genetic trials,20 years later.This indicates that clones can be selected for superior growth based on results from young trials.More intense thinning programs increased the mean diameter compared to light thinning.The study indicates that one or two early and relatively heavy thinnings can promote the development of crop trees,without jeopardizing total volume production during a rotation of 25-30 years.     

南亚热带杉木林分蓄积量生长立地与密度效应

目的 揭示南亚热带不同密度和立地条件下杉木人工林蓄积量发育规律。 方法 以广西大青山36 a生杉木密度试验林为研究对象,分析A（1 667株·hm-2）、B（3 333株·hm-2）、C（5 000株·hm-2）、D（6 667株·hm-2）、E（10 000株·hm-2）5种初植密度杉木人工林总蓄积、活立木蓄积动态变化。 结果 初植密度与立地对林分活立木蓄积生长的影响与总蓄积一致。中幼龄阶段林分总蓄积与初植密度和立地正相关,初植密度愈高,立地指数愈大,林分蓄积生长量愈大;但15 a后中等密度林分总蓄积最大。林分蓄积连年生长高峰随密度增大而愈早;平均生长达高峰后下降,密度越大下降越早、下降趋势越明显;林分蓄积连年生长量、平均生长量大多分别在10、14 a达到高峰,林分蓄积生长率降到3%时林龄为19~25 a。 结论 随初植密度增大,立地指数对林分总蓄积的影响程度减弱。若以蓄积生长量最大为培育目标,16指数级以C密度为最适宜造林密度。南亚热带杉木林首次间伐、最后间伐、主伐时间分别为10、14、25 a;主伐林龄不宜早于19 a,且初植密度愈低主伐林龄更晚,A、B密度林分的主伐林龄可推迟到25 a。     

承德地区天然白桦林生长规律研究

以河北省承德坝上高原地区天然白桦林为研究对象,根据白桦天然群落样地调查资料,选择平均木进行树干解析,以了解其生长规律。研究结果表明：35年生白桦林并没有达到数量成熟期,其树高的连年生长量和平均生长量整体呈平缓上升趋势,胸径的连年生长量和平均生长量的趋势有些动荡,但是总体还是在上升,材积的连年生长量和平均生长量的上升趋势比较明显。     

桂西南马尾松人工林生长对不同强度采伐的动态响应

[目的]研究不同强度采伐下马尾松的生长动态,筛选适宜的采伐强度,为马尾松人工林近自然经营提供技术支撑。[方法]2007年10月在14年生马尾松人工林(保存密度1 100株·hm~(-2))内进行采伐试验,设置4个采伐强度,即保留密度分别为225、300、375、450株·hm~(-2),以不采伐为对照;其后,自2008年开始连续8 a,每2 a测定1次马尾松的胸径、树高、枝下高和冠面积等生长指标,并计算单株材积和林分蓄积量,应用方差分析和Duncan多重比较分析生长指标对不同采伐强度的动态响应。[结果]表明:采伐强度显著影响林分生长,其中,林分平均胸径、单株材积、冠面积的年均增长量随保留密度增大而减小,但均显著高于对照(P0.05)。采伐后第1 3年,马尾松冠面积增长量显著高于采伐后期,胸径则在采伐后第3 5年最高,而不同采伐强度对林分树高生长影响不明显。保留密度显著影响林分枝下高和蓄积量的动态变化,其年均增长量随密度增大而递增。5个处理间林分蓄积年均增长量的差异随林龄的增大而逐渐缩小。[结论]马尾松人工林生长对不同强度采伐的动态响应以树冠最敏感,冠面积首先陡然增大,进而引起胸径的快速生长。树高和枝下高在采伐后年均增量变化相对平稳。4个采伐强度均显著促进单株材积生长,而仅保留密度为225株·hm~(-2)的采伐对林分蓄积增长量影响显著。综合比较林分的单株材积和林分蓄积连年增长量,建议在桂西南15年生马尾松人工林近自然经营中宜选择300株·hm~(-2)的保留密度进行采伐。     

Productivity and thinning effects in hybrid aspen (Populus tremula L. x P. tremuloides Michx.) stands in southern Sweden

The productivity of hybrid aspen stands in southern Sweden,originating from clone selections performed during the 1980s,was investigated. Thirteen former research and demonstrationsites were included. Repeated measurements of tree diametersand heights were taken, from which yields were estimated. Theresults indicate that the mean annual increment, obtained withoutany artificial addition of fertilizers or irrigation, will exceed20 m³ of stem wood ha^–1 a^–1 during a 20–25-yearrotation period. This corresponds to a total average woody biomassproduction, including branches, of over 8 tonnes dry matterha^–1 a^–1. These production levels show that hybridaspen is a competitive alternative for short rotation forestryin Sweden, and that the clone selection programme has greatlyenhanced productivity, compared with published growth ratesof the material previously used. For comparison, two standsthat regenerated via root suckers from material used in thebreeding programme up to the 1960s were included. They showedhigher growth than the 13 planted stands, mainly due to higherstem density and faster height development. This indicates thateven further increases in hybrid aspen yield are possible instands derived from suckers of the most recently selected clones.Thinning studies, including three different weights of thinning,were carried out on five of the sites. Five years after thestart of the treatments, stands given no thinning showed significantlyhigher current annual increment, but significantly smaller meanstem diameter than stands in which thinning was applied. However,the faster diameter development (and thus higher volumes ofvaluable wood assortments) associated with heavier thinningsmay compensate economically under current market conditionsfor the loss in total volume production.     

The laser-sandblasting method: A new method for the qualitative annual ring analysis of conifers

Summary For the assessment of many different aspects of growth processes in trees millions of annual rings are being measured all over the world every year. Other studies dealing with the prediction or explanation of wood properties also require numerous density measurements. While the density of a wood sample is relatively easy to determine by stereogravimetric methods, density relationships or variations within a piece of wood and within individual annual rings can only be detected by time-consuming techniques, e.g. radiodensitometry. A new, simplified method for ring analysis was developed with the aid of the sandblasting and laser sensor method. This process permits the automatic recording of annual radial increment and simultaneously provides results on the mean and intra-annual density of the wood specimens. Sandblasting of cross-sections of stems or of radial strips produces hardness profiles. Due to the hardness variations in these profiles individual annual rings can be more or less easily identified macroscopically, depending on the wood species. Furthermore, density variations are also macroscopically visible on account of the relationship between abrasion depth and density: Sandblasting can be said to provide a three-dimensional representation of the density relationships in wood. Following the numerical recording of these surface profiles with the aid of a laser sensor and upon conversion of abrasion depth profiles into density profiles radial increment and density parameters for stem radii, segments of radii or even individual annual rings can be determined. A comparison with the conventional radiodensitometric ring analysis method shows very good correlations with the results achieved in this study. The advantage of the laser-sandblasting method is that large samples can be investigated without great preparatory effort and in a single processing step.     

马尾松无性系种子园半同胞子代变异分析和家系选择

以福建省漳平五一林场马尾松无性系种子园自由授粉子代半同胞家系为研究对象,分析其树高、胸径、材积、冠幅、通直度、高径比、冠径比、木材基本密度、管胞长度和宽度、管胞长宽比等性状,发现家系间除管胞长宽比性状差异显著,其余性状均达差异极显著,暗示着马尾松种子园半同胞家系间存在较为丰富的变异,具选育潜力。8年生林分内各性状的家系遗传力以树高性状为最高(0·691),其次是木材基本密度(0·640)。各性状间的相关性分析结果表明,生长量性状的改良可间接改善通直度,且生长量性状与木材基本密度存在显著正相关关系,木材基本密度与管胞长度和宽度不相关。采用10%入选率,发现5年生林分和8年生林分入选家系一致,材积遗传增益分别达19·74%和19·23%,且5年生林分和8年生林分生长量性状的遗传相关极显著,认为马尾松制浆造纸材短轮伐期的初选年龄可确定为5年生;根据性状遗传力的相对大小和典型相关分析的结果,认为马尾松家系选择可适当注重树高生长量指标。以树高和材积为选择指标,按照10%入选率,兼顾材性和种子园内无性系的开花结实情况,筛选出制浆造纸材短轮伐期优良家系12个。     

毛白杨多功能新品种选育研究

观测北京、河北、河南３地点的１３年生毛白杨无性系试验林,无性系间的树高、胸径、木材密度、纤维长度等性状的差异均极显著;无性系的遗传稳定性、生长适应性、主干通直系数、树体形态系数、抗病虫指数等性状存在一定差别。材积生长量无性系×地点交互作用极显著,无性系间生长型不同（“广适型”、“高产型”、“低产型”）。无性系的材积生长量、木材密度、主干通直系数、树体系数和抗病虫害指数的重复率分别为０．５８０、０．５３６、０．３９０、０．４１１和０．５００,性状经济权重分别为０．３５、０．２５、０．１０、０．１０和０．２０,建立多功能型无性系的选择指数,其综合育种增益比对照品种≥１５％为中选优良无性系新品种。为北京、河北、河南分别选育出８个、７个和６个优良无性系,平均综合增益分别提高２３．９％、２４．９％和２１．０％;３个地点都中选的３７、３５１、９８３２和１０１２等优良无性系的平均综合育种增益提高２５．１％。所选无性系皆为雄性,显著提高了城乡绿化效应。