Carbon allocation and nitrogen acquisition in a developing Populus deltoides plantation

We established Populus deltoides Bartr. stands differing in nitrogen (N) availability and tested if: (1) N-induced carbon (C) allocation could be explained by developmental allocation controls; and (2) N uptake per unit root mass, i.e., specific N-uptake rate, increased with N availability. Closely spaced (1 x 1 m) stands were treated with 50, 100 and 200 kg N ha(-1) year(-1) of time-release balanced fertilizer (50N, 100N and 200N) and compared with unfertilized controls (0N). Measurements were made during two complete growing seasons from May 1998 through October 1999. Repeated nondestructive measurements were carried out to determine stem height and diameter, leaf area and fine-root dynamics. In October of both years, above- and belowground biomass was harvested, including soil cores for fine-root biomass. Leaves were harvested in July 1999. Harvested tissues were analyzed for C and N content. Nondestructive stem diameter and and fine-root dynamic measurements were combined with destructive harvest data to estimate whole-tree biomass and N content at the end of the year, and to estimate specific N-uptake rates during the 1999 growing season. Shoot growth response was greater in fertilized trees than in control trees; however, the 100N and 200N treatments did not enhance growth more than the 50N treatment. Root biomass proportions decreased over time and with increasing fertilizer treatment. Fertilizer-induced changes in allocation were explained by accelerated development. Specific N-uptake rates increased during the growing season and were higher for fertilized trees than for control trees.     

Regulation of ponderosa pine foliar physiology and insect resistance mechanisms by basal area treatments

We compared foliar physiology and several measures of tree resistance to insect attack among ponderosa pine (Pinus ponderosa var. scopulorum Engelm.) trees growing in thinned stands. Measurements were made in a second-growth ponderosa pine forest in northern Arizona where the basal area treatments (6.9, 18.4, 27.6, 78.2 m(2) ha(-1)) have been experimentally maintained by frequent thinnings for 32 years before our measurements began in 1994. Most of the physiological characteristics measured were affected by the basal area treatments. As stand basal area increased from 6.9 to 78.2 m(2) ha(-1), predawn water potential, midday water potential, net photosynthetic rate, resin production, phloem thickness, and foliar toughness decreased. Foliar nitrogen concentration was greatest in trees in the intermediate basal area treatments. Our results show that the physiological condition of second-growth ponderosa pine can be manipulated by silvicultural control of stand basal area, and support the hypothesis that high stand basal area increases tree stress and decreases tree resistance to insect attack.     

Stand density and fertilization effects on aboveground allocation patterns and stem form of Pinus sylvestris in young stands

The increasing demand for forest biomass for energy generation could be partially met by growing denser stands and use of fertilizer. Before this is done at large scale, more knowledge of the effects of stand density and fertilization on aboveground allocation patterns and stem form is needed. Therefore, effects of pre-commercial thinning (PCT) to 3000 stems ha^?1, an unthinned dense control (C), and PCT combined with two levels of fertilization (100 kg ha^?1 of nitrogen applied either during the establishment of the field experiment (F1) or annually (F2)) were examined in 23- to 26-year-old Scots pine (Pinus sylvestris L.) stands six years after the establishment of the field experiment. In total, 114 sample trees were harvested using destructive biomass sampling. The growth allocation and stem form of trees with diameter at breast height (DBH; 1.3 m height) >5.0 cm were not affected by either the PCT or fertilization. Small trees (DBH < 5 cm) in denser, unthinned control plots had more slender stems (lower DBH/height ratios) and allocated less growth to branches and foliage than trees in PCT plots. Fertilization had little effect on the stem form and growth allocation of the smallest trees. Therefore, effects of stem density and fertilization on stem form and growth allocation to foliage were only found for small suppressed trees, and the treatments had very little influence on dominant and codominant trees.     

Effects of atmospheric CO(2) on longleaf pine: productivity and allocation as influenced by nitrogen and water

Longleaf pine (Pinus palustris Mill.) seedlings were exposed to two concentrations of atmospheric CO(2) (365 or 720 micro mol mol(-1)) in combination with two N treatments (40 or 400 kg N ha(-1) year(-1)) and two irrigation treatments (target values of -0.5 or -1.5 MPa xylem pressure potential) in open-top chambers from March 1993 through November 1994. Irrigation treatments were imposed after seedling establishment (i.e., 19 weeks after planting). Seedlings were harvested at 4, 8, 12, and 20 months. Elevated CO(2) increased biomass production only in the high-N treatment, and the relative growth enhancement was greater for the root system than for the shoot system. In water-stressed trees, elevated CO(2) increased root biomass only at the final harvest. Root:shoot ratios were usually increased by both the elevated CO(2) and low-N treatments. In the elevated CO(2) treatment, water-stressed trees had a higher root:shoot ratio than well-watered trees as a result of a drought-induced increase in the proportion of plant biomass in roots. Well-watered seedlings consistently grew larger than water-stressed seedlings only in the high-N treatment. We conclude that available soil N was the controlling resource for the growth response to elevated CO(2) in this study. Although some growth enhancement was observed in water-stressed trees in the elevated CO(2) treatment, this response was contingent on available soil N.     

Changes in the relationship between tree size and aboveground respiration in field-grown hinoki cypress (Chamaecyparis obtusa) trees over three years

Respiration measurements of aerial parts of 18-year-old hinoki cypress (Chamaecyparis obtusa (Sieb. et Zucc.) Endl.) trees were made under field conditions over three years to study changing relationships with tree age between respiration and phytomass, phytomass increment, and leaf mass. The relationship between annual respiration (r(a)) and phytomass (w(T)) was approximated by a proportional function (r(a) = aw(T)), where the proportional constant (a) decreased year by year. The effect of time on the relationship between annual respiration and phytomass of each sample tree was fitted by a power function. Respiration of the tree suppressed by the canopy decreased year by year, but respiration of the other trees increased slightly with age. The relationship between annual respiration and leaf mass was also approximated by a generalized power function. Excluding the suppressed tree, the relationship between annual respiration (r(a)) and the annual increment of aboveground phytomass (Deltaw(T)) was described by a proportional function (r(a) = 2.27Deltaw(T)), where the proportional constant, 2.27, was independent of sample tree and year, indicating that about 2.3 times of the annual aboveground phytomass increment equivalent was respired annually. For any tree, the time constant relationships between annual respiration and leaf mass and phytomass increment for different-sized trees were similar to the corresponding time continuum relationships. In contrast, the time continuum relationship between annual respiration and phytomass differed from the time constant relationship, indicating that respiration of less active woody tissue contributed significantly to aboveground respiration. Based on the relationship between tree size and annual respiration, annual aboveground stand respiration was estimated to be 25.0, 26.9, and 25.8 Mg(dm) ha(-1) year(-1) for the three consecutive years, respectively, and the corresponding aboveground stand biomass was 60.0, 69.0, and 76.8 Mg(dm) ha(-1).     

Effects of thinning on soil and tree water relations, transpiration and growth in an oak forest (Quercus petraea (Matt.) Liebl.)

To quantify the effects of crown thinning on the water balance and growth of the stand and to analyze the ecophysiological modifications induced by canopy opening on individual tree water relations, we conducted a thinning experiment in a 43-year-old Quercus petraea stand by removing trees from the upper canopy level. Soil water content, rainfall interception, sap flow, leaf water potential and stomatal conductance were monitored for two seasons following thinning. Seasonal time courses of leaf area index (LAI) and girth increment were also measured. Predawn leaf water potential was significantly higher in trees in the thinned stand than in the closed stand, as a consequence of higher relative extractable water in the soil. The improvement in water availability in the thinned stand resulted from decreases in both interception and transpiration. From Year 1 to Year 2, an increase in transpiration was observed in the thinned stand without any modification in LAI, whereas changes in transpiration in the closed stand were accompanied by variations in LAI. The different behaviors of the closed and open canopies were interpreted in terms of coupling to the atmosphere. Thinning increased inter-tree variability in sap flow density, which was closely related to a leaf area competition index. Stomatal conductance varied little inside the crown and differences in stomatal conductance between the treatments appeared only during a water shortage and affected mainly the closed stand. Thinning enhanced tree growth as a result of a longer growing period due to the absence of summer drought and higher rates of growth. Suppressed and dominant trees benefited more from thinning than trees in the codominant classes.     

龙脑型阴香造林密度与施肥试验研究

开展龙脑型阴香 (Cinnamomum burmannii chvar. Borneol) 造林初植密度和施肥试验,3 a 生试 验林分析结果表明,不同密度和不同施肥处理对龙脑型阴香的生长（树高、地径和冠幅）有着显著的影 响,而且在主效因子（密度）、副效因子（施肥）以及两者的交互作用上均达到极显著差异（P<0.01）, 说明采用不同密度以及不同的施肥处理可调节林分生长,选择合适的处理组合可提高林分生长量。初步 结论认为营建龙脑型阴香原料林其初植密度以株行距 1.5 m×2.0 m 为宜,即 3 330 株 /hm2 且以复合肥 (N:P:K=15:15:15) 作基肥和追肥（即试验的 D2F1 组合）,可在幼林期间可以获得较高的生长量。     

Response of LAI-2000 estimates to changes in plant surface area index in a Scots pine stand

We assessed the accuracy with which the LAI-2000 plant canopy analyzer measured changes in leaf area index (LAI) and plant area index (PAI) in a 25-year-old Scots pine (Pinus sylvestris L.) stand. Stand density was 2100 stems ha(-1) and mean tree height was 8.7 m. Needle and branch areas of the stand were reduced progressively to zero by the stepwise removal of branches on all trees growing in a circular plot with a radius of 25 m. An LAI-2000 estimate was taken after each step reduction. The needle and branch surface areas removed at each step were estimated from direct measurements and were compared with the changes in the LAI-2000 estimates. Initially (before removal of branches), directly measured PAI was 5.2 (needles = 86%, branches = 8% and stems = 6%). The LAI-2000 estimate of total surface area was 66% of direct PAI and 77% of direct LAI. There was a nonlinear relationship between the LAI-2000 estimate and directly measured PAI, such that their ratio (equivalent to the clumping factor) increased from 0.66 to 1.05 with decreasing PAI. At the last measurement, when only stems were left, the LAI-2000 estimate agreed well with the direct measurement of PAI. The LAI-2000 underestimated the direct measurement of LAI at the first three steps when LAI was > 2 and the proportion of woody area was small (< 20%). However, because the LAI-2000 estimate included stem and branch areas, it overestimated the direct measurement of LAI at the last three measurements when the proportion of woody area was large (> 20%).     

Response of a rising 2-year-old natural Piedmont upland stand to fertilization

Hardwood stands in the southern U.S. are often regenerated naturally following clearcutting, with little or no silvicultural intervention in the early stages of stand development. Fertilizer was applied to a young naturally regenerating stand in order to evaluate the effectiveness of nutrient addition as a silvicultural tool in recently clearcut stands and to better understand the ecological relationships between site fertility and stand development. The study was installed on a rising 2-year-old naturally regenerated mixed pine-hardwood stand in the Hill Demonstration Forest in the Piedmont region (Durham County) of North Carolina. Fertilizer treatments were control, N, N + P, and N + P + K (at 200 kg N per ha, 50 kg P per ha and 100 kg K per ha). On a whole stand basis, increased growth rates were observed for trees on N + P and N + P + K plots. Fertilizer treatments did not affect total stand density, but the density of loblolly pine significantly decreased in N + P plots compared to the Control. The density of stems of stump origin also increased in plots receiving N + P. Density of sweetgum (Liquidambar styraciflua) significantly increased and hickory (Carya spp.) density decreased with the application of N + P. Dominant hardwood species with the exception of yellow-poplar (Liriodendron tulipifera) responded with an increase in height to only N + P fertilizer treatments. Yellow-poplar increased in groundline diameter, height and mean tree volume index with the application of N + P. Loblolly pine (Pinus taeda) responded to N + P + K fertilizer treatment with an increase in height, diameter and volume index over the Control. There was no growth response among treatments for the 16 largest trees per species per plot, with the exception of red oak (Quercus rubra) which responded with an increase in height in N + P plots. Projected growth responses were modeled on a whole stand level and indicate that the use of N + P fertilizer may be an effective silvicultural instrument to increase growth, accelerate stand development, and shorten rotation length.     

Dependence of the aboveground respiration of hinoki cypress (Chamaecyparis obtusa) on tree size

Nighttime respiration was measured at monthly intervals over one year on the aboveground parts of five sample trees in an 8-year-old hinoki cypress (Chamaecyparis obtusa (Sieb. et Zucc.) Endl.) stand, by an enclosed standing-tree method. The respiration rate rose rapidly from early spring to a maximum in June, and decreased abruptly in July and then gradually toward autumn and winter. The seasonal change in the respiration rate was synchronized with stem volume increment rather than with monthly mean air temperature. The respiration rate, r, of individual trees increased with increasing tree dimensions, such as stem volume, v(S), and stem girth at the base of the live crown, G(B). The dependence of respiration rate on tree size was successfully represented by a power function. The r - v(S) dependence was rather stronger than the r - G(B) (2) dependence, especially toward the end of the growing season (from July to September). The observed respiration rate was almost the same as the respiration rate corrected for the monthly mean air temperature. The annual respiration of individual trees was directly proportional to their phytomass or to its increment. Although the annual respiration of individual trees decreased proportionally to the square root of the leaf mass, it decreased abruptly in the range close to the smallest sample tree. Combining the monthly relationship between respiration rate and stem volume with the tree size distribution in the stand, the stand aboveground annual respiration was estimated to be 20.4 Mg CO(2) ha(-1) year(-1) (= 12.5 Mg dry mass ha(-1) year(-1)) for an aboveground biomass of 17.4 Mg ha(-1) with an annual increment of 6.51 Mg ha(-1) year(-1), i.e., the stand aboveground annual respiration amounted to the equivalent of 72% of the biomass or to almost twice the biomass increment.     

Organic fertilization leads to increased peach root production and lifespan

We evaluated the effects of mineral and organic fertilizers on peach root dynamics in the growing season from 2003 to 2006 in a nectarine (Prunus persica L.) orchard, planted in 2001 and located in the Po valley, northeastern Italy. Very few studies have conducted long-term investigations of root dynamics of fruit crops. Our main objective was to determine whether organic fertilizers affect root dynamics differently than mineral fertilizers. The experiment was a completely randomized block design with four replicates of three treatments: unfertilized, mineral fertilized and composted with municipal waste. Mineral fertilizers included P (100?kg?ha(-1)?year(-1)) and K (200?kg?ha(-1)?year(-1)) applied only at planting and N (70-130?kg?ha(-1)?year(-1)) split into two applications, one at 40 days after full bloom (60%) and the other in September (40%) each year. The compost fertilization represented a yearly rate of 10 metric tons (t) dry weight?ha(-1), which approximates (in kg?ha(-1)?year(-1)) 240 N, 100 P and 200 K, split similarly to that described for the mineral fertilization of N. Both root growth and survival were evaluated at 20-day intervals during the growing season by the minirhizotron technique. Compost increased the production of new roots compared with the other treatments (P?相似文献   

Maintenance and growth respiration of the aboveground parts of young field-grown hinoki cypress (Chamaecyparis obtusa)

Aboveground respiration of five 8-year-old trees of field-grown hinoki cypress (Chamaecyparis obtusa (Sieb. et Zucc.) Endl.) was nondestructively measured at monthly intervals over 1 year with an enclosed standing tree method. The relationship between monthly specific respiration rate and monthly mean relative growth rate at the individual tree level was described by a linear equation. During the dormant season, respiration was used mainly for maintenance purposes, whereas during the growing season, more than 40% of the respiration was used for growth purposes, i.e., 60 to 70% in May. We conclude that annual maintenance and growth respiration of a tree are directly proportional to the aboveground phytomass and its annual increment, respectively. The maintenance coefficient was estimated to be 0.504 +/- 0.039 (SE) kg kg(-1) year(-1), indicating that the amount respired for maintaining already existing phytomass was equivalent to about half of the existing phytomass. The growth coefficient was estimated to be 0.772 +/- 0.043 (SE) kg kg(-1), indicating that the amount respired for constructing new phytomass was equivalent to about three-fourths of the annual phytomass increment. The annual stand maintenance and growth respiration were, respectively, 8.8 Mg ha(-1) year(-1) for an aboveground biomass of 17.4 Mg ha(-1) and 5.0 Mg ha(-1) year(-1) for an annual stand aboveground biomass increment of 6.5 Mg ha(-1) year(-1). About two-thirds of the total respiration was used to maintain already existing biomass, and about one-third was used to construct new biomass.     

Importance of weed control, fertilization, irrigation, and genetics in slash and loblolly pine early growth on poorly drained spodosols

Research reported here has evaluated interactions of factors limiting the biological potential of slash pine (Pinus elliottii var. elliottii Engelm.) and loblolly pine (P. Taeda L.). Water and nutrients were manipulated by irrigation, weed control, and intensive fertilization. Genetic factors were incorporated as (1) improved loblolly seedlings from a commercial clonal seed orchard and (2) slash seedlings from four fast-growing, open-pollinated seed-orchard clones. Three replications of a 2 × 2 × 2 factorial experiment were established on a Typic Haplaquod soil in northern Florida using four cultural treatments: no treatment (F0H0); weed control (F0H1); fertilizer (F1F0); and combination of weed control and fertilizer (F1H1) for each species. During the first year there were large, sustained responses to all cultural practices. Loblolly and slash pine showed similar growth responses and had a lengthened growing season (60–100 days). Slash pine growth was superior to that of loblolly. Growth responses observed during the first year continued. After 4 years, irrigation had no measurable effect on tree growth. Both pine species responded equally to F1H0 and F0H1, but slash retained a slight edge. Extra nutrients]available via fertilizer, reduced competition, or additional rooting space, increased slash-pine volume index from an average of 2.0 m³ ha⁻¹ (F0H0) to 11.90 and 13.59 m³ ha⁻¹ (F0H1 and F1H0, respectively). The F1H1 treatment produced the largest slash-pine response, diameter at breast height (D_bh) averaged 9.84 cm, height reached 5.02 m, and volume index rose to about 22.94 m³ ha⁻¹. With teh F1H1 treatment, average loblolly-pine height and volume exceeded that of the average slash pine. At age 4, the growth response due to increased nutrient supply was 13–29 times that of the best genetic response within F0H0 trees of the four slash-pine progenies. The best-growing slash-pine family (6–56) exceeded loblolly-pine growth under all four treatments and approximated the vigor shown by slash pine in the southern hemisphere.     

A basal area increment model for individual trees in mixed continuous cover forests in Iranian Caspian forests

The aim of this study was to develop and test a new basal area growth model in mixed species continuous cover forests in northern Iran.Weanalyzed 421 core samples from 6 main species in the forest area to develop our growth model.In each plot,we measured variables such as total tree height(m),diameter at breast height(DBH)(cm)and basal area of larger trees as cumulative basal areas of trees(GCUM)ofDBH[5 cm.The empirical data were analyzed using regression analysis.There was a statistically significant nonlinear function between the annual basal area increment,as the dependent variable,and the basal area of the individual trees and competition as explanatory variables.Reference area from the largest trees,was circular plot with area of 0.1 ha.GCUM was estimated for trees of DBH>5 cm.Furthermore,we investigated the dependencies of diameter growth of different species on stand density at different levels of competition,and diameter development of individual trees through time.The results indicate that competition caused by larger neighborhood trees has a negative effect on growth.In addition,the maximum diameter increment is affected by competition level.Therefore,the maximum diameter increment of species occurs when the trees are about 35–40 cm in dense-forest(40 to 0 m^2 per ha)and when the trees are about 60 to 70 cm in very dense forest(60 to 0 m^2 per ha)which is more likely to Caspian natural forests with high level density due to uneven-aged composition of stands.     

固定角规点上估测林分生长的期初特征值法

复测固定角规点时，令保留木和进界木的期初断面积分别为ｇ１和ｇ０，它们的期末断面积为ｇ２，若把断面积比（ｇ１／ｇ２）和（ｇ０／ｇ２）分别称为保留木和进界木期初断面积特征值δ１和δ０，而它们的期末特征值仍为δ＝１，则林分每公顷断面积生长量△Ｇ＝Ｆ（∑Ｚ２ｉ＝１δｉ－∑Ｚ１ｊ＝１δ１ｊ－∑Ｚ２－Ｚ１ｋ＝１δｏｋ）。当保留木和进界木的期初形高分别为ｈｆ１和ｈｆ０，它们的期末形高为ｈｆ２时，则林分每公顷的蓄积生长量△Ｍ＝Ｆ（∑Ｚ２ｉ＝１ｈｆ２ｉ－∑Ｚ１ｊ＝１（δ１ｈｆ１）ｊ－∑Ｚ２－Ｚ１ｋ＝１（δ０ｈｆ０）ｋ。期初特征值法的特点是，既能保持直接法的优点———与林分蓄积现状具有一致性，便于连续清查，又能避免其缺点———抽样效率低。能真实地反映林分生长的连续性、渐进性。模拟实验证明，期初特征值法的抽样精度为９５％左右，平均系统误差为±５％左右。抽样效率是直接法的１０—３０倍。     

Spatial and Temporal Patterns in Stand Structure,Biomass, Growth,and Mortality in a Monospecific Nothofagus solandrivar. cliffortioides (Hook,f.) Poole Forest in New Zealand

Abstract

In 250 20 m X 20 m permanent plots in the Craigie-burn Range, Canterbury, New Zealand, 1970 stem density was 2,191/ha, basal area was 52.4 m²/ha, and stem biomass was 178.1 Mg/ha. Net production of stemwood (1974-1987) was 2.0 Mg/ha/yr; mortality was 3.5 Mg/ha/yr. By 1987 density had decreased by 30%, basal area by 12%, and stem biomass by 13%.

Stands with many short trees of small mean dbh were common at high elevation, whereas stands with fewer, taller trees with large mean dbh were common at low elevation. Stemwood production and mortality rate were higher in tall stands. Mortality was well distributed among plots, indicating small, frequent canopy openings; stand turnover calculations were 66 year (based on 2.2% annual biomass loss) to 83 year (based on 1.2% annual stemwood production). Larger canopy openings were also evident, but were more infrequent, so stand turnover times due to 'catastrophic' disturbances were in the range of 350-4000 yr. Consequently, the small, high-frequency disturbances blurred effects of larger disturbances on stand structure and also constrained the fluctuation in forest biomass.     

Availability of residual phosphorus fertilizer for loblolly pine

Fertilizing pine plantations with phosphorus has been a common practice on the Coastal Plain since the 1960s. A decision to fertilize a second or subsequent rotation must take into consideration the availability of residual phosphorus fertilizer. We tested the efficacy of phosphorus applied in a first rotation loblolly pine stand to support the growth of the subsequent second rotation stand. Phosphorus rates applied in 1967 at the time of planting of the first rotation were 0, 28 and 56 kg/ha. Retreatments included the application of 45 kg/ha P at midrotation in 1978, and the application of 45 kg/ha P at the beginning of the second rotation in 1991. The field trial had a split-block design with the three initial P rates crossed by five retreatments, and replicated four times. The first rotation 28 kg/ha P treatment had 91% and the 56 kg/ha P treatment 101% of the volume at age 5 of all the treatments fertilized with P in the second rotation. Foliar P levels, in the second rotation, were elevated at age 2 relative to ages 3 and 5, presumably due to effective competition control. Twenty-nine percent, 40% and 75% of the trees fertilized with 28 kg/ha P in 1967, 56 kg/ha P in 1967, and 45 kg/ha P in 1991, respectively, had foliar P concentrations of 0.10% or above at age 3. Based on foliar analyses, we recommend refertilizing stands fertilized with 45 kg/ha P in the first rotation by age 3 in the second rotation.     

Leaf nutrient variation in mature carob (Ceratonia siliqua) trees in response to irrigation and fertilization

Seasonal variations in leaf nitrogen, phosphorus and potassium concentrations were studied in a mature carob (Ceratonia siliqua L. cv "Mulata") orchard subjected to a 4-year irrigation and fertilization experiment. Three irrigation regimes (0, 50 and 100%), based on the evaporation values obtained from a class A pan, were tested in combination with two nitrogen (N) supply regimes in which 21 kg ha(-1) year(-1) (low-N) and 63 kg ha(-1) year(-1) (high-N) were supplied as ammonium nitrate. Leaf nitrogen concentration increased throughout the experiment, independently of treatments. There were no significant differences in leaf N concentration between trees in the high-N and low-N treatments. Irrigation regimes had no effect on leaf mineral concentration but influenced the amount of leaves shed and slightly modified the pattern of leaf shedding that occurred during the summer drought period. Nutritional balances between N and P and N and K were both closely and significantly correlated. Potassium was translocated from leaves to fruits during spring, independently of treatments. Severe water stress periods occurring during spring or autumn induced shedding of leaves leading to nutrient mobilization. Nutrient retranslocation during these drought periods may represent an adaptive mechanism. Nitrogen retranslocation was higher for trees in the high-N treatments than for trees in the low-N treatments, whereas phosphorus retranslocation was independent of the irrigation and fertilization treatments.     

Foliar Responses of Understorey Abies lasiocarpa to Different Degrees of Release Cutting in a Betula papyrifera and Conifer Mixed Species Stand

Foliar responses of subalpine fir [Abies lasiocarpa (Hook.) Nutt.] to thinning were studied in a 35-yr-old mixed stand of paper birch (Betula papyrifera Marsh.) and conifers. The stand regenerated naturally after a wildfire with a canopy dominated by paper birch (average height 9.8 m) and an understorey dominated by subalpine fir (average height 1.6 m). The stand was thinned to four densities of birch: 0, 600 and 1200 stems ha-¹ and control (unthinned at 2300-6400 stems ha-¹) in the autumn of 1995. The understorey conifers, mainly subalpine fir, were thinned to 1200 stems ha-¹. The study used a completely randomized split-plot design. Three sample trees were systematically selected from each treatment replicate and each tree stratum (upper, intermediate and lower understorey). One-year-old and older age class needles were collected from one south-facing branch within the fifth whorl from the tree top. Thinning of paper birch significantly (p <0.001) increased leaf area and dry weight per 100 needles for intermediate and short trees except in the 0 birch treatment. Understorey subalpine fir trees in 600 stems ha¹ birch (T3) had the largest leaf area and leaf dry weight per 100 1-yr-old needles. Specific leaf area (SLA) decreased from unthinned (T1) to 0 birch (T4). Lower understorey trees had the largest SLA. One-year-old needles had significantly higher N, P and K concentrations in all the thinning treatments. These responses are consistent with the shade tolerance of subalpine fir. The results suggest that when managing a paper birch-conifers mixed-wood forest it may be of benefit to understorey conifers to leave a birch canopy as a nursing crop.     

Long-term growth and yield effects of respacing natural regeneration of Sitka spruce in Britain

Stands of Sitka spruce (Picea sitchensis (Bong.) Carr.) originating from natural regeneration can be extremely dense with high mortality, poor growth form and low volume production. Although costly, respacing (pre-commercial thinning) can reduce resource competition for the remaining trees, altering the final crop produced. Three experiments were established to examine the effect of respacing 1–2 m tall trees to different target densities. The impact on stand growth and yield was measured 11–17 years after respacing, and the longer-term impact was modelled to year 50. Unrespaced control treatments self-thinned at a similar and constant rate at two sites. At the third, extremely high initial density resulted in a higher rate of self-thinning and lower density after 11 years. Wider respacing treatments resulted in larger mean tree diameter, but there was no significant effect of respacing on stand volume 11–17 years after respacing; greater diameter growth did not compensate for low tree number. Results indicate a volume penalty associated with delaying respacing until trees were 4 m tall, but this treatment was unreplicated. Modelled stand volume in year 50 was higher for the 2.6 m × 2.6 m respacing treatment than for the 1.8 m × 1.8 m, 2.1 m × 2.1 m or 3.3 m × 3.3 m treatments. However, open-grown conditions may reduce timber quality compared to closer spacing treatments. The results are presented using a stand density management diagram for Sitka spruce growing in Canada and support recent suggestions that British stands have a shallower size–density relationship than Canadian stands.