The relationship between yield and assimilate supply in lychee (Litchi chinensis Sonn.)

Summary

Experiments were conducted on lychee (Litchi chinensis Sonn.) in subtropical Australia (lat. 27° –29°S) to evaluate the role of assimilates on fruit retention. All the leaves of the last flush, all the leaves of the previous flush (about eight leaves per terminal shoot), or all the old leaves were removed from trees. Medium (3–5.cm diameter) or large branches (5–10.cm diameter) were girdled and defoliated after fruit set, and fruit retention compared with ungirdled and undefoliated branches. Other branches were girdled and defoliated between anthesis and fruit harvest. Finally, 20, 50 or 80% of the flowering panicles were defruited on large trees. Defoliated trees had 35 to 45% lower yields than the controls. This was despite the treatment with all the old leaves removed having a much lower leaf area index than the other defoliation treatments (1.7 vs. 2.3 and 2.8). Leaves next to the inflorescences are more important for yield than the older leaves. Fruit retention was very low on girdled branches that had been defoliated, especially when the leaves were removed in the first 20.d after anthesis. This suggests that the yields of girdled branches were determined by the availability of assimilates soon after fruit set. In contrast, the number of fruit retained on ungirdled branches was unrelated to the number of leaves, with defoliation having no effect on yield. Fruit on these branches were supported by resources from elsewhere in the tree. Thinned trees had similar yields to those of unthinned plots (65–82.kg tree^–1). Thinning apparently increased fruit retention in the remaining clusters, under a higher leaf:fruit ratio. There were large differences in the concentrations of starch in the tree, and seasonal changes, with starch declining from flowering to fruit harvest. In contrast, there were only small responses to the treatments, suggesting that the fruit were mainly dependent on current photosynthesis. Photosynthesis in the leaves behind the fruit clusters was more important than photosynthesis in the older shaded leaves.     

果树缺铁性黄化植株诊断方法的研究进展

该文主要评述了果树缺铁性黄化诊断中叶分析的不足和花分析的可行性。通过对国内外文献分析可知,叶分析存在滞后性和矛盾性两方面不足,而花分析能更好地了解果树铁营养状况,比叶分析具有一定的优越性。今后应加强花诊断指标和采样部位的研究。     

Citrus fruit enlargement by means of summer girdling

The method was based on summer girdling two-thirds of the tree’s main branches. The girdle width was c. 2.5 mm, carried out in early July and repeated about three weeks later. Such girdles, when applied to healthy vigorous trees with many leaves, may cause a fruit size increase of 15% or more. The year after girdling, treated branches tended to bear more and somewhat smaller fruits than untreated branches. A harvesting scheme to obtain maximum returns, for cultivars with long picking seasons, is also described.     

纽荷尔脐橙果实对源叶光合作用的影响

在纽荷尔脐橙果实膨大期,选取1年生双果枝、单果枝和营养枝,晴天测定源叶净光合速率(Pn)、气孔导度(Cond)、蒸腾速率(Trmmol)、胞间CO2浓度(Ci)和叶面温度(Tleaf)等生理指标,探讨挂果对源叶光合特性的影响。结果表明,挂果水平对源叶Pn、Cond、Trmmol有显著影响,对Ci和Tleaf无明显影响;上午以挂果多的枝条的源叶Pn、Cond、Trmmol峰值更低,下午则呈相反的趋势。不论枝条挂果多少,源叶Pn日变化呈典型的"双峰"曲线;Pn与Tleaf呈抛物线相关,当叶温达36℃左右时叶片光合作用最强。     

The Effect of Partial Defoliation at Different Times in the Season on Fruit Drop and Shoot Growth in Lord Lambourne Apple Trees

Six-branched trees were deblossomed on alternate branches and various defoliation treatments given. For the period of 7 to 30 days from the time of treatment, defoliations increased the compound-interest rates of fruit drop. After this period these rates of drop became equal to those of untreated trees and the relative differences in crop numbers established by this time persisted until harvest though the actual numbers still fell. The earlier in the season that the defoliations were applied the greater were the relative differences. The removal of spur leaves from branches bearing fruit caused greater drop than the removal of spur leaves from neighbouring deblossomed branches. The removal of leaves from extension and bourse shoots of fruiting or deblossomed branches had no significant effect upon fruit drop or shoot growth.

The removal of all flowers from alternate branches at mouse-ear caused more shoots to grow from these branches than from branches allowed to fruit. Removal, at pink bud, of spur leaves from either fruiting or non-fruiting branches increased the number of shoots on both branches, but removal deferred till later in the season had no effect.     

磁化水灌溉对冬枣生长及品质的影响

以‘沾化冬枣1号’和‘沾化冬枣2号’为试材,利用磁化处理的淡水和地下浅表层微咸水灌溉,通过对枣吊、叶片及果实的生长和养分含量测定分析,探究磁化水灌溉对枣生长及品质的影响。结果表明：（1）磁化水灌溉处理能显著提高叶片叶绿素含量和鲜质量,果实有机酸、维生素C、花青素含量,并且大幅度提高果实的耐贮性。（2）沾化冬枣1号经磁化（进口磁化器A400p）淡水灌溉处理,相比淡水对照枣吊长度与直径提高了11.375%和15.810%,单叶面积提高了12.418%,单果质量、含水量、果实横径和纵径分别提高了23.779%、2.377%、3.650%和5.071%;沾化冬枣2号经磁化（进口磁化器A400p）淡水灌溉处理,枣吊长度与直径提高了23.602%和13.710%,单叶面积与叶片厚度提高了23.622%和13.825%,单果质量和还原糖含量提高了12.526%和12.110%。（3）利用磁化（自主研发磁化器DS-948-1）地下浅表层微咸水灌溉处理沾化冬枣2号,与微咸水对照相比,果实含水量和还原糖含量分别提高了4.386%和9.158%。（4）灌溉水经磁化处理后,冬枣叶片矿质元素N、P、Cu含量提高（P < 0.05）。磁化水灌溉处理有效促进了沾化冬枣的枝叶和果实生长发育,提高了果实的品质和耐贮性。     

Effect of part-tree flower thinning on fruiting,vegetative growth and leaf photosynthesis in ‘Cox’s Orange Pippin’ apple

Flower clusters were removed at full bloom from ten year old ‘Cox’s Orange Pippin’ trees on M.9 rootstocks, over the whole tree, on alternate branches or on whole sides. Mean fruit weight per tree at harvest was linearly dependent on leaf area per fruit and on light interception per fruit, both relationships accounting for over 90% of the variance. These relationships did not differ between treatments, implying either a mobile pool of carbohydrate or photosynthetic adjustment within the tree to crop load. Measurements of leaf photosynthesis in July and September showed no statistically significant differences in photosynthetic rate of spur or extension shoot leaves on bearing or non-bearing branches. Although the treatments caused no overall effects on shoot growth or leaf area per tree, sides of trees without fruit had greater leaf area and shoot growth than did sides bearing fruit. Fruit mineral composition and percentage dry weight were not affected by treatment except where the treatments significantly altered fruit size. In the following spring, although the treatments did not affect the total number of flower buds produced, branches that were deflowered in the previous spring carried significantly more flower buds than did branches which had cropped.     

螺旋环剥对幼龄‘桂味’荔枝果期光合和蒸腾作用的影响

以6年生幼龄‘桂味’荔枝(Litchi chinensis Sonn.‘Guiwei’)为试材,于5月果实发育期,从螺旋环剥处理和对照树上分别选取来自同一基枝的有果枝和无果枝,观察枝梢生长的情况并进行叶片光合和蒸腾作用的研究。结果表明:螺旋环剥有利坐果;螺旋环剥缩小有果叶与无果叶在光合效率上的差别,显著降低了叶片的最大光合速率(Amax)、表观量子效率(AQY)和羧化效率(CE),提高了光补偿点(LCP),削弱了光合效率;净光合速率(Pn)与蒸腾速率(Tr)极显著相关,螺旋环剥显著降低了叶片的Tr,Pn也下降但差异不显著,蒸腾作用减弱的程度比光合作用减弱的程度大;对照叶片的Pn和Tr日变化为单峰曲线型,14:00时达最高峰,螺旋环剥叶片的Pn和Tr在13:00时有明显的午休现象,气孔限制是午休的主要因素;相同处理的有果枝叶片,其日均Pn和Tr都高于无果枝叶片,说明果实的存在可提高‘桂味’荔枝的Pn和Tr。     

Productivity of olive trees with different water status and crop load

SUMMARY

A field experiment was conducted over two growing seasons to determine the combined effect of crop load and irrigation on yield components of olive trees (Olea europaea L. ‘Leccino’) planted at 6 m 3.8 m in a sandy-clay soil. Different crop loads were established by manual thinning of fruits. Drip irrigation was managed to maintain pre-dawn leaf water-potentials (PLWP) within the following ranges: (i) higher than –1.1 MPa (FI; fully irrigated); (ii) between –1.0 and –3.3 MPa (DI; deficit irrigated); or (iii) below –1.2 MPa, but not lower than –4.2 MPa (SI; severe deficit irrigated). The irrigation period lasted from 6 – 16 weeks after full bloom (AFB) in 2003, and from 5 – 19 weeks AFB in 2004. In 2003, full bloom was on 26 May; in 2004, it was on 3 June. Neither irrigation regime nor crop load affected flowering or flower quality the following Spring. The combined fruit yields [on a fresh weight (FW) basis] over both years in SI and DI trees were 49.0% and 81.6% of FI trees, respectively. The oil yields of SI and DI trees were 52.5% and 81.2% of FI trees, respectively. Fruit FWs in FI trees were greater than those of DI or SI trees at 8 weeks AFB. At harvest, FI trees bore the largest fruits, and SI trees the smallest fruits. The FWs of individual fruits at harvest in the FI and DI treatments decreased as crop load increased, but no such relationship was apparent for SI trees. The oil content of the mesocarp increased as PLWP increased from approx. –3.5 MPa to –1.5 MPa. The oil content of FI trees at harvest decreased from 53.1% to 45.7% dry weight as fresh fruit yield increased from 5 – 25 kg dm^–2 trunk cross-sectional area. However, crop load did not have any effect on the oil content of the mesocarp in DI trees. Fruit maturation was delayed by irrigation. Maturation index also decreased (indicating delayed maturation) as the crop load on FI or DI trees increased, but did not vary with crop level in SI trees.     

‘糯米糍’荔枝碳素营养储备动态与坐果的关系

 以10～12年生的‘糯米糍’荔枝(Litchi chinensis Sonn. ) 为材料, 研究了高产树(60～65 kg·株^{- 1} ) 和低产树(5～10 kg·株^{- 1} ) 不同部位(叶片、各级枝条和主干) 碳素营养储备(淀粉) 的差异及季节动态; 分析了不同果实发育阶段树体碳素营养水平与坐果率的关系。结果表明, 果实成熟时高产树各部位的淀粉含量均低于低产树, 而可溶性糖含量高于低产树。果实采收后, 低产树早于高产树抽发新梢。入冬季前(11月底前) , 低产树和高产树分别抽发了3次和2次秋梢。7～11月秋梢生长发育期间, 高产树和低产树均无显著淀粉积累, 11月中旬以后枝梢生长停滞期间, 各部位, 尤其是4 cm直径以内的枝条大量积累淀粉, 在花穗发育前达到高峰。之后, 随花穗发育、开花及坐果而持续降低。高产树和低产树各部位淀粉高峰并无明显差异, 表明坐果量对树体碳素营养储备的累积并无明显的长期影响。叶片、主枝和主干积累的淀粉含量较低, 总体相对稳定; 而4 cm直径以下的枝梢淀粉含量变化剧烈, 说明这些枝梢是更为活跃的碳素储备库。本研究还表明, 坐果早期(花后3周内) 枝条(2 cm直径) 的淀粉含量与该枝条上最终坐果率呈显著正相关, 而果实发育中期(花后8周) 枝条淀粉含量与坐果率无关, 说明早期果实发 育一定程度依赖树体碳素营养储备, 而中后期果实发育几乎不依赖树体储备。     

Relations Between Growth of Leaves,Fruit and Shoot of Glasshouse Tomato Plants

The weekly rates of shoot extension and total growth in leaf length of the glasshouse tomato variety Potentate increased from germination until soon after the start of ovary swelling. It is postulated that when, or soon after, the first flower primordia were initiated, an internal reaction was stimulated that eventually resulted in the cessation of the progressive increase in the rates of shoot extension and growth in leaf length. Increase in the rate of vegetative growth ceased soon after the start of ovary swelling, but this occurred even when no ovaries were allowed to swell.

The diameter of the shoot was unaffected by removal of leaves but increased when no fruits were allowed to develop. Conversely, shoot extension growth was unaffected by removal of fruit but decreased when alternate immature leaves were removed, although the removal of mature leaves had no effect on shoot extension growth.

Removal of leaves was associated with a reduction in total fruit volume and an increase in total leaf length.

Removal of immature fruit was associated with an increase in total leaf length. Removal of alternate immature fruits had little effect on total fruit . volume ; the difference between normally fruiting plants and plants from which alternate fruits had been removed was less than 0?5%.     

脱落酸对长白忍冬果实和叶绿原酸含量的影响

以长白忍冬为试验材料,在展叶期及果实转色期分别用脱落酸(ABA 200 mg/L)对其叶片以及果实进行处理.结果表明:ABA处理显著增加了长白忍冬果实中绿原酸含量,高于对照15.04％,对叶片中绿原酸含量增加无显著差异.     

植物器官体积变化连续测微法指导果树自动化灌溉合理指标的研究

对盆栽葡萄（VitisviniferaL．）、苹果（MaluspumilaMill．）、山楂（CrataeguspinnatifidsBunge）和枣（ZiziphusjnyubaMill．）4种果树在不同水分营养状况下茎于微变化过程中的日最大收缩量（MDS）、日净增长量（DG）、当日完全复原所需时间（RT）等3个重要指标进行研究，结果表明：3个指标都比叶片日出前水势对土壤水分营养状况的反应灵敏得多；随着土壤可利用水分的不断减少，山楂和枣的MDS急剧增加，而葡萄和苹果的MDS减小，不同类型果树之间的反应不一致；在土壤水分状况较好时，4种果树的DG的变化平缓，变化幅度较小；在良好的土壤水分条件下，4种果树的RT随着土壤相对持水量的下降而迅速延长，并在土壤水分含量降低到一临界值以下时维持在24h。因此，在应用植物器官体积变化连续测微法指导果树灌溉时，MDS和DG并不适于单独作为所有果树的自动化灌溉指标，而RT作为果树自动化灌溉指标较为理想。     

The Influence of Shoot Competition on Fruit Retention and Cropping of Apple Trees

Fruit set was increased by removing all shoots 5 days after full bloom and at weekly intervals thereafter from trees of Sunset and Laxton’s Fortune, but removing shoots from Fortune trees 25 days after full bloom produced no beneficial effect on fruit retention. All treatments resulted in a heavier rate of fruit shedding during the ‘June drop’ period than occurred from control trees, and at harvest the trees without shoots had fewer fruits, and lower yields, than the controls. In a comparison of shoot removal and shoot tip removal starting 15 days after full bloom on Fortune trees, both treatments improved set, but whereas shoot removal caused a heavier ‘June drop’ compared with untreated trees, shoot tip removal increased the number of fruits retained to harvest and produced a yield increase. The difference between the two treatments in their influence on fruit retention, during and after the ‘June drop’, is accounted for by the beneficial effect of a relatively small number of leaves on each tipped shoot. Studies on the pattern of distribution of photosynthates, using ¹⁴CO₂ and autoradiography, produced results supporting the concept of competition between fruits and shoots and also showed changes in the pattern of assimilate movement brought about by shoot tip removal. It is concluded that competition between fruits and shoots, occurring during blossoming and the following 2–3 weeks, may limit fruit set, but the presence of shoot leaves is beneficial to fruit retention in the later part of the season, particularly during the ‘June drop’ period.     

Salinity effects on strawberry plants grown in rockwool. I. Growth and leaf water relations

Comparison of four everbearing or day-neutral cultivars of strawberry (Rapella, Ostara, Fern and Selva) grown under glasshouse conditions in rockwool showed Rapella to accumulate the greatest shoot dry weight and leaf number together with the production of few stolons but many relatively large fruits per inflorescence. Subsequent studies with cv. Rapella found that increasing the salinity of the irrigation supply from 2.5 to 5.7 and to 8.5 mS cm"1, by the addition of NaCl, progressively reduced leaf number emerged and final leaf area by up to 36% and 48% respectively, after 100 d of treatment. Shoot dry weight, excluding fruits, was reduced by up to 47%. The number of crowns (lateral branches) was reduced from 11 in the lowest salinity to 8 and 7 at the higher salinity levels. Leaf water potential, \)/w, was reduced by up to 0.23 MPa in young leaves and 0.36 MPa in older leaves. Leaf osmotic potential, xj/^, was decreased by up to 0.14 MPa in younger leaves following salinity treatment at 8.0 mS cm-1. Consequently leaf turgor was reduced more in older than in younger leaves. Inflorescence number was reduced from 27 to 20 and 17. Within the remaining inflorescences however, flower numbers, fruit numbers and duration of fruit ripening were not affected. A marked decrease in the fresh weight of the fruit was detected in both higher salinity treatments but fruit dry weight was only slightly reduced.     

Mobilization of nitrogen in the olive bearing shoots after foliar application of urea

Mature ‘on’ and ‘off’ ‘Manzanillo’ olives trees with three healthy branches of 10–12 cm in diameter were selected to determine changes in nitrogen levels in the bearing shoots after foliar application of urea. Each selected branch of each tree received one of the following treatments: (i) control without urea application; (ii) foliar application of urea to all the current-season leaves; and (iii) foliar application of urea to all the one-year-old leaves. Urea was applied in May, four days after full bloom in the ‘on’ year trees. Each treated leaf was immersed in a test tube containing a 4% urea solution and 0.1% Tween 20 for 5 s. Bearing shoots, composed of both one-year stems and leaves and current-season stems and leaves, were collected at intervals from the beginning of the experiment until 64 days after urea application. Nitrogen was determined in stems and leaves from each part of the bearing shoot, and in fruits during the ‘on’ year. Nitrogen uptake from the leaves was rapidly mobilized from the older to the current-season leaves of the bearing shoot, and thereafter to other storage organs of the tree or to the fruit, which is the largest nitrogen sink in the bearing shoot. No translocation of nitrogen from the current-season leaves to older leaves was observed. The rapid translocation of nitrogen from the younger leaves to other storage organs of the tree could explain the insensitivity of leaf analysis to detect excess nitrogen, since mature leaves from current-season shoots must be sampled to determine the nutritional status of the tree. The failure of leaf analysis to detect excess nitrogen may be a cause of nitrogen over-fertilization in olive orchards.     

Root growth following defruiting improves peach tree water status

Summary

Tree growth and water status throughout the growing season and after fruit removal were studied in container-grown peach trees. Trees with fruit (F) and defruited (DF) trees were sampled destructively at bud break (8 March), 1 month after fruit removal (3 June), at harvest (6 August), and before leaf fall (15 October) to determine the mass of leaves, current season shoots, branches, trunk, and the entire root system. Tree water status was determined from the mid-day stem water potential (SWP) the day before each sampling date. Root growth in DF trees was greater than that observed in F trees, while the above-ground biomass was similar in DF and F trees. DF trees therefore had lower leaf:root biomass ratios than F trees throughout the fruit growing season. Environmental factors did not fully explain the seasonal variations in SWP, but there was a significant correlation between leaf:root biomass ratios and SWP. Reductions in leaf:root biomass ratios were accompanied by increases in SWP and, ultimately, DF trees had higher SWP values than F trees in mid-Summer. Improvements in tree water status following fruit removal can be explained, in part, by additional root growth.     

Drying half of the root-zone from mid fruit growth to maturity in ‘Hass’ avocado (Persea americana Mill.) trees for one season reduced fruit production in two years

We tested the effect of extended drying of half the root system on fruit yield and fruit Ca concentration, an indirect measure of fruit quality, in avocado (Persea americana Mill. cv Hass). In a field experiment on a sandy soil, withholding irrigation and plastic sheeting was used to dry the root-zone beneath the whole canopy (DD) or half the canopy (WD), compared with well-watered trees (WW). The irrigation water contained added nutrients and was slightly saline. Yield, shoot growth, leaf conductance, leaf and fruit water status and mineral concentrations of leaves and fruit were studied. The responses of treated trees were assessed in the following season during which normal irrigation practices were restored. With respect to yield, the WD treatment behaved the same as the DD treatment. It reduced yield by more than half and proportionately more than the reduction in water supply thus reducing irrigation efficiency. Re-watering did not restore yield of WD or DD-trees in the next season. The WD and DD treatments had no effect on the concentration of Ca in the fruit mesocarp and so are unlikely to affect fruit quality. The main impact of reduced water supply on the trees was fruit abscission and this was linked to dry soil around the roots rather than the water status of the leaves or fruits. We conclude that extended drying of half of the root-zone in one season reduced irrigation efficiency for two seasons by promoting the abscission of developing fruit to the same extent as occurred when the whole root system was exposed to extended drying.     

李树叶片细菌性穿孔病的发生规律及防治研究

细菌性穿孔病可导致李树叶片大量脱落,树势衰弱,产量降低,品质下降.研究发现,病害的初侵染源来自枝干病斑,生长季节枝、叶、果互相交叉传染.种植感病品种、树势强旺、雨日多、雨量大为该病的流行因素.选用抗病品种、消除初侵染源、加强栽培管理、生长期用药控制是防治该病的有效措施.     

核桃果实发育初期的水分状况

对在不同土壤水分条件下核桃枝条、叶片和果实水分含量动态与生长发育关系的研究结果表明,土壤供水不足时,树体水分亏缺,影响枝条生长和果实发育,降低坐果率.对有关水分生理指标测试表明,不同品种核桃叶水势(φW)日变化虽有差别,但相似,呈双峰型,叶φw愈低,晚上回升愈缓慢.叶φw日变化还受枝条上着生果实的影响,未着生果实枝条叶φw高于有果枝的.叶φw降低抑制光合产物的形成和累积.核桃蒸腾作用的日变化也呈双峰曲线,并与环境因子有密切关系.叶组织水分亏缺时,抗脱水能力强,具有一定的抗旱性能,晚熟品种优于早熟品种.