A COMPARISON OF THE HERBAGE YIELD OF ITALIAN RYEGRASS AND TALL FESCUE MIXTURES IN THE SECOND AND THIRD YEAR AFTER SOWING

The annual yield of tall fescue was higher than that of Italian ryegrass in the third year after sowing, but the total yield of herbage from grass plus clover swards was similar.
In both the second and third year after sowing, the yield of herbage in the spring grazing was higher when fescue was used as the sown grass. The method of establishment of both tall fescue and Italian ryegrass affected the total and seasonal yield in the second and third year after sowing, but the magnitude of these effects was not nearly as marked as it was earlier in the life of the leys. In the second year after sowing, swards of both Italian ryegrass and tall fescue had a higher yield of total herbage and of white clover, and a lower ingress of unsown species, when established without a cover crop and grazed frequently in the year of sowing.
The inclusion of red clover did not increase total yield of DM in the second and third year after sowing, and it slightly decreased the yield of the tall fescue mixture in the third year following sowing when N was applied. S170 tall fescue was readily grazed by sheep in spring and autumn.
The apparent recovery of applied N varied with the mixture sown, and the management given during establishment.     

EFFECTS OF METHOD OF ESTABLISHMENT ON TALL FESCUE AND ITALIAN RYEGRASS MIXTURES IN THE FOLLOWING YEAR

Tall fescue and Italian ryegrass mixtures react differently to management in the year of sowing. The decrease in yield of tall fescue and the increase in growth of red clover, caused by the use of a cover crop, was still evident in the following year. In both the spring grazing and total yield of the first harvest year, the Italian ryegrass/ white clover mixture was better than the tall fescue/white clover mixture when a cover crop was used. The reverse trend was recorded when the grass plus clover was sown without the cereal. In general, the addition of red clover to the tall fescue/ white clover, or Italian ryegrass/white clover mixture, increased the yield, but the magnitude of the increase was modified by management during establishment. The increase in total herbage yield and the reduction of white clover growth due to nitrogen application were both related to management in the year of sowing. Consideration should thus be given to method of establishment in the assessment of herbage seeds mixtures.     

Effect of dairy slurry application rate and forage type on production,soil nutrient status and nitrogen‐use efficiency

Seven forage types (diploid and tetraploid perennial ryegrass, Italian ryegrass and hybrid ryegrass, a low‐input mixture of perennial ryegrass, cocksfoot, timothy and meadow fescue, a mixture of perennial ryegrass and white clover, and monoculture of red clover) were sown in late July 2004. Each received one of four rates of dairy cattle slurry in three annual applications by trailing shoe, which supplied average nitrogen (N) inputs of 0·0, 114·9, 204·8 and 301·2 kg N ha^?1 annum^?1. Treatments were cut either three or four times annually over four years. Average dry‐matter yield (DM) response to slurry N was 15·6 kg DM kg^?1 N. Lowest recovery of slurry N was in the second application each year (after first cut). The data suggest that slurry applied to Italian ryegrass, and also to swards containing legumes on soils with high phosphorus content, will produce a lower DM response to slurry N and result in a lower slurry N recovery than on swards of perennial ryegrass or cocksfoot‐dominant low‐input mixtures. Apparent recovery of slurry N was low at the second cut, especially when first‐cut yields had been high. To maximize slurry N recovery, application to regrowths with potentially slow rates of growth or high legume content should be avoided.     

Effect of fertilizer nitrogen on six-year-old red clover/perennial grass swards

Three diploid red clover cultivars—Sabtoron, Violetta and Essex—and three tetraploid, Hungaropoly, Teroba and Red Head, were sown separately in pure culture and with each of three companion grasses: timothy (Aberystwyth S48), tall fescue (Aberystwyth S170) and perennial ryegrass (Aberystwyth S24).
The effects of fertilizer N on yield and on clover/grass ratio over a 2-year period (seventh and eighth harvest years) subsequent to 6 harvest years during which no N fertilizer was applied were investigated. The data for productivity and persistence have already been published (McBratney, 1981; 1984).
Application of fertilizer N increased DM yields in the eighth year. In this year, the highest yield, 11·9t ha^-1, averaged over the six clover cultivars, was given in association with tall fescue. Tall fescue contributed 90% of this yield. Clover content continued to decrease in all swards but the decrease was greatest in the swards receiving fertilizer N. The yield of clover DM averaged over the six cultivars under N treatment declined from 5·6t ha^-1 in the seventh year to only 0·4t ha^-1 in the eighth year.
The results from this trial demonstrate the potential of red clover sown either pure or in mixture with a suitable perennial grass, to maintain high output of quality herbage over a 6-year period without the aid of fertilizer N. They further demonstrate that following decline in red clover content, both herbage yield and quality may be restored by the application of N fertilizer, particularly where the clover was seeded with a highly productive companion grass.     

A COMPARISON OF THE REACTION OF DIFFERENT GRASS SPECIES TO FERTILIZER NITROGEN AND TO GROWTH IN ASSOCIATION WITH WHITE CLOVER

Seven species or varieties of grass, and a mixture of 3 of them, were sown in pure swards and treated with 4 levels of nitrogenous fertilizer (0,17·5, 35 and 70lb N per acre per cut). Each species, and the mixture, was also sown with white clover. The experiment was cut 4 or 5 times per year. The effect of fertilizer on the yield of each grass was compared with the effect of clover on the yield of the grass/clover swards. Mean annual yields showed an approximately linear response to N; there was a small but significant fall in response to the highest level of N. Response among the species ranged from 20 to 30 Ib of dry matter per Ib N applied for the intermediate level of fertilizer and from 14 to 23 Ib for the final increment of fertilizer.
S37 cocksfoot, S48 timothy, S24 ryegrass, and a mixture of these grasses, were high yielding and responded well to fertilizer N; Irish ryegrass and Agrostis tenuis were less productive and gave poorer responses to N. S215 meadow fescue and S23 ryegrass were intermediate in yield and response.
There were no significant differences between the annual yields of the 8 grass/clover mixtures; the yields of the grass and clover components of each mixture were inversely related. The effect of clover on the yield of the grass/clover mixtures was estimated to be equivalent to the effect of an annual application of 205 lb N per acre to Agrostis tenuis and 120 lb N to S48 timothy. The fluctuations in annual yields were greater with grass/ clover mixtures than with grass swards receiving N.
The yields of grasses when sown with clover were in similar order to their yields when sown pure; but whereas the latter tended to fall from year to year, the yields of the grass components of mixtures (except Irish ryegrass) did not.     

Performance and quality of tall fescue (Festuca arundinacea Schreb.) and perennial ryegrass (Lolium perenne L.) and mixtures of both species grown with or without white clover (Trifolium repens L.) under cutting management

There is increasing interest in tall fescue (Festuca arundinacea Schreb.) in Western Europe and elsewhere, mainly because of its better drought resistance and yield potential compared with perennial ryegrass (Lolium perenne L.). Important drawbacks of tall fescue, compared with perennial ryegrass, are its lower digestibility and voluntary intake. Mixtures of both species might combine the advantages of each, and species interactions may eventually lead to transgressive overyielding. We compared the agronomic performance of tall fescue, perennial ryegrass and tall fescue–perennial ryegrass mixtures, as pure‐grass swards or in association with white clover (Trifolium repens L.). Tall fescue–perennial ryegrass mixtures differed in the proportion and ploidy of the perennial ryegrass component. Yield, feed quality and botanical composition were measured in the 3 years after the sowing year. We found significant effects of ploidy of the ryegrass variety and of the proportion of ryegrass in the initial seed mixture on the botanical composition of the swards. Nevertheless, all swards were dominated by tall fescue at the end of the experiment. No overyielding of the mixtures compared with that of single‐species swards was found, but feed quality was intermediate between that of the single‐species swards. Mixed swards had better drought resistance than L. perenne and higher feeding quality than F. arundinacea swards.     

Establishment, persistence and nitrogen fixation of white clover and marsh trefoil on blanket peat

White clover (cv. S184 or Blanca) or marsh trefoil (cv. Grasslands 4703 or Grasslands Maku) were sown alone and in combination with a companion grass (tall fescue, red fescue, perennial ryegrass or ryegrass cleanings) on a peat area of hill land originally dominated by Juncus articulatus and Molinia caerulea.
Lime and ground rock phosphate were applied in 1973, the area rotavated to a shallow depth and seeds sown in May 1974. No fertilizers were applied except at sowing time and two harvests were taken per annum for 4 years.
Differences in yield and legume content between companion grass treatments were small. After the first full harvest year Blanca swards usually had the lowest DM yields and those containing Maku the highest. The contribution of marsh trefoil to total herbage yield increased from 4·2% in 1975 to 22% in 1978, whereas white clover decreased from 4·2 to 1·2%. Legume yields in 1978 were less than half those in 1977. Overall, N yields were low, Maku swards fixing most N estimated to be highest in 1977 at 35 kg ha^-1.
It is concluded that marsh trefoil grows well on upland peat but more information on its response to grazing is necessary before conclusions can be drawn about its value in hill land improvement.     

Effects of white clover cultivar and companion grass on winter survival of seedlings in autumn-sown swards

The aim was to study the effects of white clover cultivar and combinations with perennial ryegrass cultivars on seedling establishment in autumn‐sown swards and on winter survival of seedlings. Large‐leaved white clover cv. Alice and small‐leaved white clover cv. Gwenda, and an erect and a prostrate perennial ryegrass cultivar were sown in autumn in pure stands and as four binary grass‐clover mixtures. Mixtures of white clover cv. Huia and Aberherald with perennial ryegrass were also sown. Companion grasses had no significant impact on the establishment of white clover. The number of seedlings of white clover cv. Alice in mixtures (335 m^?2) was higher than cv. Gwenda (183 m^?2) and pure swards had similar white clover population densities as mixed swards. White clover cv. Huia tended to have more seedlings than Aberherald (355 and 205 m^?2 respectively). No stolons were produced prior to a severe winter, because of the late sowing date. Winter survival of clover seedlings was 0·56 in mixtures and 0·69 in pure stands, irrespective of white clover or companion grass cultivar. Stolon development of white clover in autumn is often considered essential for overwintering survival and spring growth. In this study, there was considerable survival of the non‐stoloniferous tap‐rooted seedlings of all four clover cultivars despite a severe winter.     

A comparison of perennial ryegrass‐ and tall fescue‐based swards with or without a cropping component for dairy production: Animal production,herbage characteristics and financial performance from a 3‐year farmlet trial

Perennial ryegrass/white clover swards have some limitations in temperate grazed dairying systems. This study tested the hypothesis that farmlets based on alternative species would be equally or more profitable than those perennial ryegrass‐based, and would produce more herbage in summer‐dry conditions. Six farmlets were established; three with perennial ryegrass‐ and three with tall fescue‐based swards. For each grass species, one farmlet was solely based on grass‐clover swards while the other two had either chicory‐red clover or lucerne crops planted on 20%–25% of the area. Animal‐ and herbage performance‐related variables were measured for 3 years, and calculated financial performance was evaluated. Using tall fescue improved total annual herbage yield compared with perennial ryegrass, but animal production and operating profit were lower. This was likely due to the reduced yield and nutritive value of tall fescue during spring and an associated decline in daily milksolids production. The deficit in spring milksolids production was never recovered, despite greater herbage production from tall fescue during summer/autumn. Incorporating chicory‐red clover or lucerne crops reduced both annual herbage and milksolids production. This reduced farm income, while increasing operating expenses as the farmlets required crop renewal and more purchased supplementary feed to maintain feed supply. Under the conditions of this study (i.e., partial irrigation, high nitrogen supply), changing the forage base from perennial ryegrass to tall fescue did not improve animal production or profitability, nor did incorporation of crops on 20%–25% of the farmlet area.     

Yield and stability of yield of single- and multi-clover grass-clover swards in two contrasting temperate environments

Diversity of clovers in grass-clover swards may contribute to greater herbage yields and stability of yield. This possible effect was evaluated in an experiment carried out over three harvest years at two contrasting sites, differing in precipitation and soil composition, using mixed swards containing either one, two or three clover species sown together with timothy ( Phleum pratense L.) and meadow fescue ( Festuca pratensis L.). The clover species were red clover ( Trifolium pratense L.), white clover ( Trifolium repens L.) and alsike clover ( Trifolium hybridum L.) sown in various proportions in a total of ten treatments. All swards were fertilized with nitrogen with amounts that increased from year to year, and three harvests were taken in three consecutive years. There was a significant interaction between site and species mixture on total dry matter (DM) yields (range 27–32 tonnes ha⁻¹) and DM yields of clovers (range 5–15 tonnes ha⁻¹); red clover as a single species or in a mixture was superior at the dry site while multi-clover species mixtures were superior at the wet site. Alsike clover was the least productive species of clover. Stability of yield of clovers was generally higher by including white and red clover in the seed mixture but total DM yield was not.     

Herbage productivity of a range of grass species in association with white clover

Red fescue ( Festuca rubra L.) cvs Reptans, Dawson and Ruby, perennial ryegrass ( Lolium perenne L.) cvs Perma and Sprinter, crested dogstail ( Cynosurus cristatus L.) cv. Southlands, smooth meadow-grass ( Poa pratensis L.) cvs Parade and Arena, Yorkshire fog ( Holcus lanatus L.) commercial types I and II, creeping bent ( Agrostis stolonifera L.) cv. Penncross and rough meadow-grass ( Poa trivialis L.) cvs Omega Øtofte, Dasas and Sabre were each sown with white clover ( Trifolium repens L.) cv. Donna and the productivity of the fourteen mixtures assessed under simulated grazing (six harvests per year) for 3 years. A single fertilizer-N rate (50 kg ha⁻¹) in early spring was applied to all treatments each year.
Over the 3 years, mean annual dry matter (DM) production of the swards ranged from 5·25 t ha⁻¹ (creeping bent) to 8·71 t ha⁻¹ (red fescue cv. Reptans), with the species ranking order of: red fescue > perennial ryegrass > crested dogstail > smooth meadow-grass > Yorkshire fog > creeping bent. Rough meadow-grass only persisted over the first harvest year. Mean annual organic matter digestibility (OMD) values of the swards ranged from 0·712 (creeping bent) to 0·782 (ryegrass cv. Sprinter). Mixtures with the lowest proportion of grass (crested dogstail and smooth meadow-grass) generally gave the highest proportions of white clover. The creeping bent mixture had exceptionally low production of both components.
Red fescue, crested dogstail and smooth meadow-grass showed compatibility with clover and, therefore, have potential for use in grass/clover swards. Yorkshire fog had lower compatability with clover. Some breeding effort to improve selected characteristics of promising species, e.g. improvement of low OMD in red fescue, seems warranted.     

How does the vertical and horizontal structure of a perennial ryegrass and white clover sward influence grazing?

Mixtures of perennial ryegrass ( Lolium perenne L.) and white clover ( Trifolium repens L.) sown in alternate rows or in a thoroughly mixed matrix were grazed by sheep, either continuously or during short grazing tests, and were used to investigate the influence of the vertical and horizontal components of the sward structure on defoliation by sheep.
In an experiment under continuous grazing, the defoliation intensity was greater for white clover compared with perennial ryegrass leaves (0·80 and 0·58 respectively). In spring, perennial ryegrass leaves were more defoliated than white clover leaves, whereas the reverse was observed in summer. The ratio of the proportion of white clover to perennial ryegrass leaves grazed was negatively correlated with the difference between the surface height of the perennial ryegrass and white clover rows in spring. In both spring and summer, white clover leaves of the same extended leaf length had a higher proportion of them grazed than perennial ryegrass leaves.
In another experiment, during short grazing tests with perennial ryegrass–white clover swards that were grazed at the same sward surface height and at the same white clover content as in the previous experiment, there were no significant differences in the proportion of white clover and perennial ryegrass leaves grazed between strips of the two species and thoroughly mixed structures. The proportion of white clover leaves grazed was higher than that of perennial ryegrass leaves.
These results show that the differential defoliation by sheep of perennial ryegrass and white clover leaves varies according to their vertical distribution in the mixed canopy, but is little affected by their horizontal distribution. Even small differences in sward surface height between mixed perennial ryegrass and white clover can affect diet selection by sheep to a rather large extent.     

Productivity of red clover grown alone and with companion grasses; further studies

Three diploid—Sabtoron, Violetta and Essex—and three tetraploid—Hungaropoly, Teroba and Red Head—red clover cultivars were sown alone and with each of three companion grasses—timothy (S48), tall fescue (S170) and perennial ryegrass (S24). The productivity and persistence of the red clover cultivars were compared. Results obtained over the first four harvest years were presented in an earlier paper (McBratney, 1981) and this paper presents results from the fifth and sixth (final) harvest years.
The low yield obtained in the fifth and sixth year from Essex under each seeding treatment indicated a deterioration in its persistency or vigour or both. In the fifth year the remaining two diploids, Sabtoron and Violetta, were as highly productive and persistent as the three tetraploids but in the sixth year the decrease in productivity was greater in the diploids than in the tetraploids. In the fifth year Teroba, the highest yielder, produced 9 8 t total DM ha⁻¹ over all seeding treatments of which 73% was clover DM; in the sixth year Teroba was again highest yielding, producing 9.3 t ha^−l with 67% clover content. Tall fescue was the most consistent companion grass in the mixed swards, in the sixth year contributing 53% grass DM to the total 8 8 t DM harvested ha⁻¹.
The trial demonstrates the potential of a number of red clover cultivars grown either pure or with a suitable perennial grass to achieve and maintain a high output of herbage of good quality over a period of 6 years without any application of fertilizer N. The practical agricultural implications of using red clover are discussed.     

Productivity of red clover grown alone and with companion grasses over a four-year period

Six red clover cultivars, three diploid—Essex, Sabtoron and Violetta—and three tetraploid— Teroba, Red Head and Hungaropoly—were sown alone and with each of three companion grasses—timothy (S48), tall fescue (S170) and perennial ryegrass (S24). The productivity and persistency of the red clover cultivars were compared over 4 years. Dry matter (DM) yield, DM digestibility and the crude protein (CP) concentration were assessed and botanical analyses conducted on herbage samples from each treatment at each of three harvests per annum. Annual fertilizer application consisted of 165 kg P and 312 kg K ha^-1. Comparing clover cultivars alone Essex was significantly less productive and less persistent than the other five cultivars. Yield and persistency of the five other cultivars did not differ markedly within years with the exception that the diploids were significantly less productive than the tetraploids in the fourth year. Over all 4 years mean annual total DM and clover DM yields of the five cultivars were between 12·2 and 13·2 t ha^-1 and between 9·2 (79·2% of total DM yield) and 10·2 (83·2%) t ha^-1 respectively, and differences were not significant. Up to the end of the third year there was little or no advantage gained by the inclusion of a companion grass, annual total DM yields being between 11·2 and 14·2 t ha^-1 for clover alone and between 10·2 and 14·2 t ha^-1 for clover-grass mixtures. In the fourth year there was an overall tendency for the yield of the clover alone to be lower, between 7·2 and 12·2 t ha^-1, than that of the clover-grass mixture, between 8·2 and 13·2 t ha^-1, and this was more pronounced with the diploid than with the tetraploid clover cultivars. Sown with companion grasses, Essex and Hungaropoly were lower in yield and in contribution than the other cultivars over the 4 years. The influence of the companion grass on total dry matter yield showed that the contribution of timothy was low relative to that made by tall fescue and perennial ryegrass. Perennial ryegrass made the most varied contribution from year to year. Tall fescue was the most consistent contributor with all clover cultivars and at the end of 4 years both yield and clover-grass balance had not changed materially. No pronounced differences in DM digestibility were evident between treatments. Crude protein concentration of the pure clover was similar to that of the clover-timothy treatments and both would appear to be superior to either the clover-perennial ryegrass or clover-tall fescue mixtures. It is considered that red clover dominant swards are suitable for use under a cutting regime and can provide high yields of DM at a low cost for up to 4 years. Such swards are self-sufficient in N and in addition soil N accumulation can be exploited in the production of succeeding crops.     

A COMPARISON OF THE EFFECT OF ITALIAN RYEGRASS AND TALL FESCUE MIXTURES ON THE FOLLOWING CROPS OF KALE AND BARLEY

The effects of preeropping with Italian ryegrass and tall fescue mixtures on marrow-stem kale (1st test crop) and barley (2nd test crop) were measured. Application of N to the grass/clover swards reduced the yield of DM, P, K, Na, Mg, crude protein and also the percentage crude protein in the kale test-crop. Following the tall fescue swards, the yield of DM, P, Na, Ca, Mg and E, crude protein and percentage crude protein in the kale were lower than after the Italian ryegrass swards. These effects were particularly evident in the kale stem. Interactions occurred which involved the method used to establish the herbage mixtures, the application of N to the swards and of fertilizers to the kale. These effects were complex, but the method of sward establishment could clearly affect the following kale crop grown several years later. The influence of grass swards on the second test-crop (spring barley) was much smaller than on kale. Precropping with Italian ryegrass or using a cover crop during grassland establishment reduced 1000-grain weight of barley. The crude protein percentage in barley grain was reduced by the compound fertilizer applied to the previous kale crop, particularly when no cover crop had been used during establishment of the herbage mixtures. The converse of these effects was recorded in the crude-protein percentage of the straw of the barley test-crop.     

The effect of time of sowing and sowing method on production of white clover in mixed swards

Two field trials were carried out in successive years in which (1) perennial ryegrass and white clover seeds were drilled together, or (2) clover was broadcast and grass drilled, or (3) both clover and grass were broadcast. The last two treatments were followed by harrowing or not of the seed bed. Sowing took place in early May or August. Four harvests were taken in each full harvest year. In establishment years, sowing in May resulted in a total dry matter (DM) production at least ten times higher than that resulting from sowing in August. No sowing method treatment effects on total DM or clover yield were significant in the establishment year (measured only in spring-sown plots). In the first harvest year the August-sown treatments produced 15% less DM than those sown in May and clover yield was, on average, 40% lower than the earlier sown treatments; clover proportion followed a similar pattern to yield. Drilling of both grass and clover without harrowing produced swards with a significantly lower proportion of clover in the first harvest year in the first trial than in treatments in which clover was broadcast but not harrowed. In the second trial at harvest 1, clover proportion just failed to be significantly lower in the treatment in which grass and clover were drilled than in the treatment when clover was broadcast and the seed bed harrowed. In the second harvest year (first trial only), annual clover yield and proportion were not affected but drilled grass and clover had lower DM yield than when grass was drilled and clover broadcast without harrowing and when both were broadcast, without harrowing. At one harvest, the yield of clover in treatments sown in May was actually lower than that in the later-sown plots. In a subsidiary controlled environment experiment to investigate the effect of stage of clover development on cold hardiness (a factor in autumn-sown swards), plants which were about to initiate stolons (58 days old) had an LD₅₀ of ?5·1° compared with ?9·3°C for plants 60 days older. It is concluded that autumn sowing delays the time at which optimum clover production is achieved (late in the first full harvest year) and method of sowing does not compensate for this. However, sowing in August under Northern Ireland conditions does not seem to jeopardize the chances of a successful establishment of white clover, and plants should be sufficiently winter hardy to withstand relatively hard freezing conditions.     

EFFECTS OF WHITE CLOVER ON THE BOTANICAL COMPOSITION OF SWARDS OF RYEGRASS, TIMOTHY AND MEADOW FESCUE AND MIXTURES OF THESE THREE

Results are presented to show the effect of white clover (Trifolium repens) and of applied N on the hotanical composition of swards sown with perennial ryegrass (Lolium perernne), timothy (Phleum pratense), meadow fescue (Festuca pratensis) and mixtures of two and three of these species. White clover had a more favourahle effect on the growth of perennial ryegrass than on either timothy or meadow fescue both in terms of yield and in terms of tiller numbers. Tiller numbers of rough-stalked meadow grass (Poa trivialis) were increased and tiller numbers of Agrostis species were reduced by application of N and by inclusion of clover. Possible reasons for these effects are considered.     

The spatial pattern of vegetation in cut and grazed grass/white clover pastures

The spatial patterns of white clover and sward surface height (SSH) that developed In established perennial ryegrass (Lolium perenne)/white clover (Trifolium repens) pastures undercutting lent every 4 weeks to 5 cm) and gracing (continuously grazed with sheep to 5 cm) were measured. While clover cover was recorded in 1000 contiguous 5 × 5-cm quadrats down 50-m permanent transects from early spring to late autumn. Measurements of SSH were made at 10-cm intervals down the same transect. Spatial pattern was analysed using two-term local quadrat variance and patch-gap analysis. At least two scales of spatial pattern existed for white clover when defoliation treatments began. White clover was not distributed at random but found in patches (mean size = 1.1 m) where it was finely intermixed with grass. Patches, separated by gaps (regions of no clover) (mean size = 2.3 m), were in turn aggregated into ‘patches of patches’, separated by larger gaps (mean size = 4.1 m). Under grazing the pattern of patches and gaps did not alter. Under cutting, patch size increased and gap size decreased, explaining in part the greater mass and cover of white clover that arose in cut than grazed swards during the experiment. No new patches of white clover due to seedling establishment or clonal growth were observed in either cut or grazed swards. The intensity of pattern increased in both cut and grazed swards, but the increase was greater m cut swards. The initial single scale of spatial pattern of SSH of tall patches (mean size = 1.2 m) separated by short patches (mean size = 2.7 m) did not change under grazing. SSH became uniform under cutting. It is suggested that the response of plants to selective (spatially heterogeneous) grazing is a crucial factor in the development and maintenance of spatial pattern in grasslands. The importance of spatial pattern to our understanding and interpretation of plant-plant and plant-animal interactions and to the composition of temperate grasslands is considered.     

The effect of companion perennial ryegrass cultivars on red clover productivity when timing of the first cut is varied

Replicated plots of Hungaropoly red clover were sown on a sterilized area in May 1975 alone (seed rate 11 kg ha^-1) or with one of six cultivars of perennial ryegrass (seed rate 3·5 kg ha^-1) viz. Cropper and S24 (early heading), Barlenna and Hora (medium heading) and Melle and Perma (late heading). In 1976 and 1977 primary growth was cut at one of four dates ranging from mid-May to mid-June and thereafter plots were harvested twice each year.
Varying the time of first cut did not have a significant effect on total dry matter (DM) yield in either year despite differences in means of cutting treatments on annual red clover yields of the order of 6–9%.
In some companion grass treatments total DM yield in 1976 was increased and total red clover yield and percentage red clover contribution were reduced relative to swards sown only with red clover. In 1977 a similar but non-significant trend was found. Swards containing early ryegrasses had higher total herbage DM yields but lower red clover yields and contents than all other swards at the first harvest in both years.
Delay in date of taking the first harvest in 1976 reduced DM digestibility in the first cut and increased it in the second in both years.
It is suggested that by cutting early and increasing the number of harvests from three to four per year, differences in the content of red clover between the first and second cut might be reduced, and it is concluded that more benefit is derived from red clover when medium or late heading ryegrasses are used as companion grasses.     

The value of timothy (Phleum pratense L.) in ryegrass — timothy mixtures managed to simulate intensive grazing

The performance of timothy in mixtures with perennial ryegrass was assessed under a simulated intensive grazing management over two harvest years in 1974–75. Three seed rates of S23 perennial ryegrass were factorially combined with three rates of Scots timothy and compared with pure stands of each grass. All were sown with Huia white clover. When cut six times at monthly intervals and with an annual N input of 350 kg ha^?1, there were no significant differences in total DM production in either year. The 2-year mean DM yield for the nine mixtures and six pure swards was 9·77 t ha^?1 (range 9·34–10·16). Compared with the pure ryegrass swards, in both years the ryegrass-timothy mixtures produced earlier spring growth but were significantly lower yielding at the second cut. Over the first five cuts the proptortion of timothy in the three mixtures with 22·4 kg ha^?1 ryegrass seed averaged 26% in the first year and 37% in the second. Corresponding calculated mean DM yields of timothy were 2·75 and 3·00 t ha^?1. It is concluded that an early timothy variety is capable of competing with a late-heading perennial ryegrass in frequently cut swards managed to simulate intensive grazing. The strong development of timothy in the dry summer of 1975 suggests that in mixtures of late perennial ryegrass varieties, an early variety of timothy should be beneficial for its spring growth in grazed swards.