Effects of forest harvesting on runoff and sediment characteristics in the Hyrcanian forests,northern Iran

Loss of canopy cover by forest harvesting generally increases the average surface runoff volume and sediment. Selective cutting (single and group selection method) is the most usual forest harvesting method in the Hyrcanian forests in the north of Iran. The purpose of this study is to find the effect of selective logging technique on the hydrological behavior of runoff and sediment in the Kheyrud forests located in northern Iran over 1 year. Four treatments were implemented: natural forest without harvesting (C), forest with selective harvesting (H) and area without canopy cover (WC) and skid trail (S). Three types of data were measured in each plot including soil chemical and physical properties, runoff and sediment load after each rainfall. The results indicate significant differences (P ≤ 0.05) in runoff generation and sediment production with respect to the treatments cover. The runoff in all treatments showed relatively similar response to rainfall, while the highest runoff and sediment were observed in skid trails, and the area without canopy cover (1.13 and 0.62 mm, and 1.2 and 0.51 g m^?2), averagely. In contrast, the natural forest without harvesting and the forest with selective harvesting treatments exhibited the lowest amounts of runoff (0.2 and 0.44 mm) and sediment (0.1 and 0.17 g m^?2), averagely. Implementation of low logging technique was useful to control the effects of logging on the runoff and sediment yield.     

Effects of forest vegetation on runoff and sediment transport of watershed in Loess area, west China

This paper aims to study the effects of vegetation on runoff and sediment transport at the watershed scale, and to provide a theoretical basis for afforestation in the Loess area, in the nested Caijiachuan watershed, Jixian County, Shanxi Province of west China. Forest watersheds and farmland watersheds with similar terrain features were selected through cluster analysis to study their runoff and sediment transport characteristics. Results showed that compared with farmland watersheds, runoff generation time in forest watersheds was delayed remarkably, and peak flow was reduced greatly, which indicates that vegetation played an important role in holding and absorbing rainfall. Besides, with the increase of forest coverage, the runoff amount, runoff depth and runoff coefficient decreased during the rainy seasons. The runoff depth and runoff coefficient of farmland watersheds in the rainy season were 5–20-fold as much as that of forest watersheds, and runoff and sediment yield of watersheds with low forest coverage were 2.7–2.9-fold and 3–6-fold as great as those with high coverage during rainstorms, and low forest coverage had larger variation in sediment hydrograph. For the complexity and scale dependence of the influence of forest vegetation on runoff, forest hydrological functions based on regional scale or watershed scale were worthy of further studies. __________ Translated from Science of Soil and Water Conservation, 2006, 4(3): 48–53 [译自: 中国水土保持科学]     

机械运输对伊朗北部山林陡峭集材道土壤的干扰

A study was conducted to investigate the effects of skid trail slope and traffic levels on soil disturbances at two soil depths (0–10 and 10–20 cm). The treatments were set at four traffic levels (2, 7, 12 and 20), two slope classes (<20% and >20%) and two soil depths (0–10 and 10–20 cm). Results show that skidder traffic, longitudinal slope and soil depth have significant effect on soil bulk density in skid trail. Comparison of average soil bulk density in different traffic levels shows that there are significant differences in average bulk density between different traffic levels and control (p<0.05). The average bulk densities in different slopes and soil depths are significantly increased with increase in traffic levels, maximized at 12 passes (p<0.05), but there are no significant differences between 12 and 20 passes. The interaction effects between traffic and soil depth are significant (F _0.05,3=0.109, p<0.001). For all traffic treatments, there are significant differences in soil moisture content between the two slope classes and the two depths (p<0.001). However, the interaction effects between traffic levels and slope classes are not significant (p >0.05), although skidder traffic and slope affected soil moisture content.     

Effect of forest vegetation on runoff and sediment production in sloping lands of Loess area

According to fixed-position data for 1985–2003 from nine runoff plots of Caijiachuan watershed which lies in Jixian County of Shanxi Province in Loess area, this paper studied the relationship between vegetation and runoff and sediment production in sloping lands in detail, which helps to provide scientific basis for vegetation re-construction and studies on environmental transformation of water and sediment in watersheds of Loess area. Although, many study results testify that forest vegetation has an important function in soil and water conservation and cutting runoff, the effect of vegetation on runoff and sediment transmission is complicated, and this needs to be studied in depth. The results of the paper showed the following. Firstly, the natural secondary forest performs better function of soil and water conservation than artificial Robinia pseudoacacia forest, and runoff and sediment produced in the former in individual rainfall were 65%–82% and 23%–92% of those produced in the latter. At the same time, better correlative relationship between runoff and sediment production and rainfall and rainfall intensity were testified by multiple regression, but the correlation decreased gradually with the increase of canopy density of forest. Secondly, the difference of runoff and sediment production in several land use types was very distinct, and the amount of runoff and sediment produced from Ostryopsis davidiana forest and natural secondary forest were the least, and runoff and sediment produced from in artificial Robinia pseudoacacia forest and Pinus tabulaeformis forest were 5-fold as much as those from O. davidiana forest. Besides, runoff and sediment produced in mixed planting of apple trees and crops were 16.14-fold and 2.96-fold than those of O. davidiana forest, respectively, but the amount decreased obviously after high-standard soil preparation in the case of the former. Thirdly, based on gray cognate analyses of factors affecting runoff and sediment production in sloping land, the factors of stand canopy density and herb and litter biomass were the most significant ones, whose gray incidence degree exceeded 0.6. Therefore, mixed forest with multi-layer stand structure and shrub forest should be developed in vegetation re-construction of Loess area, which will help to increase coverage and litter thickness in order to cut down the runoff and sediment dramatically in sloping land. __________ Translated from Chinese Journal of Applied Ecology, 2005, 16(9): 1,613–1,617 [译自: 应用生态学报, 2005, 16(9): 1,613–1,617]     

Fire intensity effects on seed germination of native and invasive Eastern deciduous forest understory plants

Low-intensity fires were important for maintaining the structure of Eastern deciduous forests (EDFs) for thousands of years before European settlement of North America, though fire suppression became a standard management practice in the 1930s. More recently, prescribed fires have been reintroduced to EDF habitats to aid in the restoration of native plant diversity, but invasions of non-native species such as Microstegium vimineum (Japanese stiltgrass) may increase prescribed fire intensity and suppress colonization of native species. As fire becomes a more common management tool in these habitats, it is vital to predict fire temperature effects on the native and non-native species present in the system. In this study, we found that prescribed fires in areas invaded by Microstegium can be 250-300 °C hotter than fires in nearby native-dominated areas. We then compared the effects of fire on germination rates of six native and three non-native EDF understory species representing the range of functional groups common in this habitat. We manipulated both fire intensity (temperature and length of exposure) and type of fire effect (direct flame and indirect furnace heat) to generate germination curves and make predictions about potential prescribed fire effects on populations of these species. There were very different responses among species to both direct (flame) and indirect (furnace) heating. Germination of three native species, Lycopus americana (American water horehound), Verbesina alternifolia (wingstem), and Vernonia gigantea (tall ironweed), showed signs of being stimulated by heating at low temperatures, while germination of all non-native species (M. vimineum, Elaeagnus umbellata, and Schedonorus phoenix) were inhibited at these lower intensities. High fire intensity (temperatures above 300 °C) effectively killed most species, though one native species, Senna hebacarpa (American senna) and one non-native species, E. umbellata (autumn olive), were capable of tolerating 500 °C temperatures. We conclude that high-intensity prescribed fires in habitats invaded by Microstegium may reduce seed germination of some non-native species, but may also inhibit the regeneration of native understory species.     

Species differences in timing of leaf fall and foliage chemistry modify nutrient resorption efficiency in deciduous temperate forest stands

Extensive variation in fractional resorption of mineral elements from plant leaves is still not fully understood. In multi-species forest stands, species leaf fall phenology and leaf constitution may significantly modify the timing of nutrient return to the soil and overall plant nutrient loss. We studied leaf fall and nutrient loss kinetics, and leaf composition in three natural, temperate, deciduous broadleaf forest stands to determine the role of timing of leaf abscission and nutrient immobilization in cell walls on nutrient resorption efficiency of senescing leaves. Nitrogen (N), phosphorus and potassium contents decreased continuously in attached leaves after peak physiological activity during mid-season. Changes in nutrient contents of attached leaves were paralleled by decreases in nutrient contents in freshly fallen leaf litter. In different species and for different nutrients, resorption of nutrients from senescing leaves proceeded with different kinetics. The maximum nutrient resorption efficiency (the fraction of specific nutrient resorbed from the leaves at the end of leaf fall) did not depend on the mid-seasonal nutrient concentration. Species with earlier leaf fall resorbed leaf nutrients at a faster rate, partly compensating for the earlier leaf fall. Nevertheless, the litter-mass weighted mean nutrient contents in leaf litter were still larger in species with earlier leaf fall, demonstrating an inherent trade-off between early leaf fall and efficient nutrient resorption. This trade-off was most important for N. Losses of the non-mobile nutrients calcium and magnesium were unaffected by the timing of leaf fall. There was large variation in the maximum N resorption efficiency among species. Correlations among leaf chemical variables suggested that the maximum N resorption efficiency decreased with the increasing fraction of cell walls in the leaves, possibly due to a greater fraction of N occluded in cell wall matrix. We conclude that species leaf fall phenology and leaf chemistry modify the timing and quantities of plant nutrient losses, and that more diverse forest stands supporting a spectrum of species with different phenologies and leaf types produce litter with more variable chemical characteristics than monotypic stands.     

红壤区生态公益林不同类型林分降雨及产流与产沙量研究

通过对湘中丘陵红壤区不同类型林分的定位观测,对林地不同年份因降雨量差异而产生的径流量、泥沙含量的差异进行对比分析.结果表明:降雨量较大的2010年林地的径流量、泥沙含量、径流系数均大于相对干旱的2009年;径流量、泥沙含量的大小与大雨量降雨的次数和降雨量密切相关.     

Characteristics of runoff and sediment generation of forest vegetation on a hill slope by use of artificial rainfall apparatus

We studied the impact of forest vegetation on soil erosion,surface runoff, and sediment generation by using field simulated rainfall apparatus. We measured runoff and sediment generation of five 4.5 × 2.1m runoff plots (a bare soil as a control; two Pinus tabulaeformis forestplots and two Platycladus orientalis forest with row spacing of 1 m × 1m and 1.5 m × 1.5 m, respectively) in Beijing Jiu Feng National ForestPark under three rainfall intensities (0.42, 0.83, 1.26 mm per minute).Forest vegetation significantly reduced soil erosion and sediment yield.Mean total runoff volume in the four tree stand plots was 93% of that inthe control plot, demonstrating the limited effectiveness of forest vegetation in runoff control. With increasing rainfall intensity, runoff reductionin forest plots declined from 28.32% to 2.1%. Similar trends in runoff coefficient and the relationship between runoff volume and rainfall duration was observed. Mean total sediment yield and mean sediment yield reduction rate under different treatments was 55.05% and 43.17% of those in the bare soil control plot, respectively. Rainfall intensity playedan important role in runoff and sediment generation processes, and had agreater impact on runoff than on soil erosion and sediment generation.When considering several factors in runoff and sediment transport processes, the P. tabulae form plot with row spacing at 1 × 1 m had a greater effect on soil and water conservation than did other forested plots.     

Dispersal of Quercus mongolica acorns in a broadleaved deciduous forest

Effects of two sets of factors on the disappearance of acorns of Q. mongolica var. grosserrata were investigated. Acorns were distributed on (a) the floor of a forest with its canopy intact, (b) the floor of a forest clearing, (c) the vegetated sloping bank of a forest road, and (d) a bare forest road. They disappeared fastest from the forest with intact canopy and slowest from the bare road, an effect thought to be directly related to amounts of cover.When the acorns were covered with frames having netting with different-sized apertures, a mesh of 3.0×4.0 cm did not appreciably alter the rate of acorn disappearance compared with the uncovered controls: a mesh of 0.5×0.5 cm totally inhibited the disappearance of acorns.Five species of small mammal were live-trapped. Of these, Clethrionomys rufocanus, Apodemus argentinus and A. speciosus are thought to be implicated in the disappearance of acorns. The rate of acorn disappearance seems to be directly related to total numbers of small mammals.Acorns disappeared more rapidly when distributed on the ground than when buried.     

Assessing the effects of initial soil characteristics,machine mass and traffic intensity on forest soil compaction

An extensive field trial was set up in eight forest stands to examine the influence of soil texture (two stands on sand, four on loam to silt loam, two on clay), machine mass (light, heavy) and traffic intensity (one and five skidding cycles) (i.e. pass back and forth on the skid trail) on soil compaction after mechanized harvesting. Dry bulk density (BD), penetration resistance (PR), micro-topography and soil carbon dioxide (CO₂) concentration were applied as response variables for soil compaction. Significant effects on BD were nearly absent (<7% increase) and occurred occasionally for PR (60–70% increase, up to 150% on clay soils). Especially for loam to silt loam and clay soils, this was in contrast with the expectation. The negligible compaction degrees for loam to silt loam are attributed to high initial compaction levels that prevented further compaction, as was found by general linear modelling (GLM) for both BD and PR. For clay soils the small compaction degrees can be explained by the high water contents that result in plastic deformation instead of strong compaction degrees, as was confirmed by the micro-topographical measurements. GLM also revealed a significant impact of machine mass (BD) and soil water content (BD, PR) on the compaction degree. Soil texture, traffic intensity and position in relation to the wheel tracks generally turned out to have an insignificant influence. With regard to clear interactions the influence of traffic intensity depends on the position in relation to the wheel tracks and the machine that was used (PR).     

Seasonal variations in radiation and energy balance in a deciduous broad-leaved forest

This study was conducted to document the effects of canopy change on radiation and energy balances in a deciduous broad-leaved forest. Global solar radiation was highly variable with a maximum close to 28.7 MJ m⁻² day⁻¹ and a minimum of about 0.7 MJ m⁻² day⁻¹. The high daily variation of net radiation was the result of the frequency and duration of rainfall. The mean annual albedo in this study site was 0.126. Seasonal variation of albedo was characterized by steadily decreasing values with leaf-fall, but at the time when new leaves sprouted the trend did turn upwards. The mean annual radiation efficiency at this study site was 0.564, and the seasonal variation of the radiation efficiency was dependent upon the net long-wave radiation rather than albedo. This study also illustrated the seasonal variation of the proportion of the actual evaporation to the equilibrium evaporation as a means for estimating the latent heat flux from forest using the equilibrium evaporation model. Seasonal variation of the proportion of the actual evaporation to the equilibrium evaporation reflected the physiological and productive activities of trees. Latent heat flux dominated the energy balance and, in particular, reached about 96% of net radiation in July.     

老挝过度采伐热带混合落叶林区的人工补植(英文)

人工补植通常用于增加低商业价值树种盛行的次生林中期望树种的密度。本研究评估了均格种植法(vis-à-vis gap)和行植法(line planting)2种人工补植方法,以及老挝过度采伐的热带混合落叶林中的青皮木(Vatica cinerea) 和龙脑香木(Dipterocarpus alatus)和3种豆科植物(缅茄 Afzeliax ylocarpa,大果紫檀Pterocarpus macrocarpus,酸枝木Dalbergia cochinchinensis)的表现。人工补植实验安捧在了一个随机完整的大块场地,做了7个重复。栽植7年后测定了存活率、高度和直径生长,并进行方差分析。用2种人工补植方法栽植的幼苗的存活率没有发生变化(p0.05)。然而,就直径和高度生长而言,均格种植法要比行植法效果更好。这与林冠在格间和行间块速封闭有关,这种情况在行间更加明显。在存活率,高度和直径生长方面,存在着明显的种间变异。耐阴的龙脑香科树种存活率和生长表现优于喜光树种(p0.0001)。个体的大小分布不规则,说明林下光照条件不均衡。考虑到行植法很难维持恒定线条宽度和均匀光线条件,及其每年用于清理的成本和僵硬的几何模式,我们推荐采用均格种植法。龙脑香木(Diperocarpus alatus)和青皮木(V.cinerea)可用于混合阔叶林人工补植的树种。缅茄、酸枝木和大果紫檀可作为人工林的混合树种,在较宽林隙地、直线区域和开放地种植或在火耕林幼林冠下种植。     

Effects of geometric design of forest road and roadside vegetation on traffic noise reduction

Geometric design of forest roads and design of their landscapes can reduce noise pollution and its harmful effects on human health. We investigated the effects of technical and biological parameters such as geometric road design and various roadside tree stands on reducing noise pollution according to the tree density and distance from roadway in Darabkola Forests, Sari, Iran. We recorded the noise generated by a car(Land Rover) relative to changes in longitudinal slope, horizontal curve radius and type of road pavement. We also measured noise levels according to roadside tree density and stand type(coniferous and hardwood) in 40 rectangular plots of three widths(25, 100 and 300 m) and 50 m length that were randomly demarcated along forest roads. The changes in noise level were recorded using a decibel meter with an accuracy of ±1.5 d B and resolution of 0.1 d B. Noise levels were higher alongside unpaved roads than alongside paved roads. There was an inverse relationship between the measured noise level and horizontal curve radius. The rate of noise level on horizontal curve with a radius less than30, 30–45 m and more than 45 m were 64.8, 70.8 and75.9 d B, respectively. The noise level increased with the increasing longitudinal slope of the road. There was a significant difference between the noise level on slopes less than 3 %(67 d B) and 3–8 %(71.2 d B) in comparison with slopes greater than 8 %(77.8 d B),(p 0.05). Pinus brutia L. reduced the noise level more(about 6 d B) in stands of1/3 density of mixed hardwoods within 25 m from middle of the road. Careful design of geometric properties of forest roads as well as planting coniferous trees with hardwoods is a suitable solution for reducing noise pollution.     

Soil changes after traffic with a tracked and a wheeled forest machine: a case study on a silt loam in Sweden

Alterations to some soil physical parameters were evaluatedafter passes by a tracked forest machine and a wheeled one ofequal mass (about 20 000 kg). Measurements were made after arange of machine passes, with a maximum of eight. The measuredsoil physical parameters were dry bulk density, penetrationresistance, intrinsic air permeability, saturated hydraulicconductivity, porosity and pore-size distribution. Rut depthwas also measured. Although the wheeled machine caused deeper ruts than the trackedone, alterations caused by the two machines to the measuredsoil parameters were similar, except in the uppermost 5–10cm. The wheeled machine caused a decrease in bulk density at5 cm depth, whereas the tracked machine caused an increase,despite its lower ground pressure.     

间伐强度对兴安落叶松林林下植被多样性及生物量的影响

研究了间伐强度对兴安落叶松林林下植物多样性和生物量的影响,为确定兴安落叶松林分的最佳采伐强度提供理论依据。以兴安落叶松林为研究对象,设置4种间伐强度(10%、20%、30%、40%)和对照的试验固定样地,对间伐5 a后各样地林下灌草多样性和生物量进行调查分析。结果表明:1)抚育间伐可以明显提高林下植物种类、Menhinick丰富度指数、Simpson指数、Shannon-Wiener指数和Pielou均匀度指数,各层均以40%间伐强度最大,而生态优势度指数则呈递减趋势(P <0.05);2)适当加大间伐强度对林分林下植被层和枯落物层生物量具有促进作用,均以间伐强度30%为优(P <0.05);3)综上所述,在30%～40%的间伐强度下,兴安落叶松林林下植被多样性和生物量最高,林分质量最佳。     

Wildfire and salvage harvesting effects on runoff generation and sediment exports from radiata pine and eucalypt forest catchments, south-eastern Australia

This study examined the effect of wildfire and salvage harvesting on runoff generation and sediment exports from three small forest catchments in south-eastern Australia. In 2006, wildfire burnt a radiata pine catchment and two adjacent natural eucalypt forest catchments which formed part of a long-term hydrological research project. Subsequently, only the pine plantation catchment was salvage harvested. The combined effect of fire and salvage harvesting in the pine catchment caused a substantial increase in runoff compared to the burnt eucalypt forest catchments and pre-fire conditions, particularly in response to high intensity, short duration summer storms. Post-fire maximum suspended sediment concentrations from fixed-interval sampling greatly exceeded pre-fire values for both eucalypt and pine catchments, while sediment (suspended and bedload) exported from the pine catchment exceeded each of the eucalypt catchments by a minimum of 180 and 33 times. However, the export increase was probably closer to 320 and 71 times based on a survey of eroded channels in the pine catchment combined with measured post-survey exports. Notably, seven summer storm events accounted for approximately 80% of the pine catchment sediment yield. Hillslope process measurements indicated that the highest runoff velocities occurred in log drag-lines formed by cable harvesting, while soil water repellency was more extensive in the harvested pine catchment than in the adjacent eucalypt catchment. The latter effect probably resulted from higher burn severity in the pines combined with reduced soil moisture due to less shading after harvesting. Runoff modelling indicated that the log drag-lines acted as an extension to the drainage network and increased peak flows at the harvested catchment outlet by 48% for a high intensity summer storm event, while substantial reductions in modelled runoff were achieved through increasing the hillslope surface roughness coefficient. It is recommended that post-fire salvage operations should avoid the formation of log drag-lines when using cable harvest techniques and maximise surface cover to limit increases to runoff, erosion and catchment sediment exports.     

Prescribed burning of Pinus oocarpa in Honduras I. Effects on surface runoff and sediment loss

The effects of low-intensity prescribed burning on surface runoff and sediment loss were studied in a stand of oocarpa pine (Pinus oocarpa Schiede) in central Honduras. The effects of burning in two seasons were compared on small paired plots on 10°, 25° and 40° slopes. Over a period of almost 2 years, mean percent runoff was 1.73% on the control plots and 5.03% on the burned plots. Mean sediment loss during the same period was 80 kg ha^?1 on the control plots and 1732 kg ha^?1 on the burned plots. The relationship between slope and surface runoff and sediment loss was negative; the effect of slope being masked by variation of other site factors. Recovery from burning was rapid and is attributed to the rapid rate of recovery of the ground vegetation of grasses and forbs which are adapted to frequent burning. Surface runoff and sediment loss on control plots were very low and the increases caused by burning are considered an acceptable alternative to the exposure of the overstory and site to the high risk of wildfires in accumulated fuels. Greater runoff and sediment loss were recorded on the plots burned in February than on those burned in June. As the period November to mid-February offers the greatest number of days with weather conditions suitable for low-intensity prescribed burning, it is concluded that burning for hazard reduction should be conducted as early as possible in this period, in November and December, to permit greater recovery of the ground vegetation before the heavy rains of May and early June.     

龙王山区落叶阔叶林林分结构特征研究

为全面掌握龙王山自然保护区落叶阔叶林的林分结构特征,设置了3个100m×100m样地,对其中的6个30m×30m样方,进行群落和林分调查。经统计分析,结果表明龙王山落叶阔叶林中树种种类多,优势树种为小叶白辛树和锐齿槲栎,林分树种组成具有一定的复杂性和多样性。林分株数密度平均为1511株/hm2,林分样地间差异不明显。胸径分布呈典型的倒"J"型;树高主要集中在4-9m的树高级,分布则呈偏左的正态分布。林分结构总体上具有多树种复层异龄的基本特征,具有良好的生物多样性保护和植物种质资源保存作用。     

Differences in leaf phenology between juvenile and adult trees in a temperate deciduous forest

In a deciduous forest, differences in leaf phenology between juvenile and adult trees could result in juvenile trees avoiding canopy shade for part of the growing season. By expanding leaves earlier or initiating senescence later than canopy trees, juvenile trees would have some period in high light and therefore greater potential carbon gain. We observed leaf phenology of 376 individuals of 13 canopy tree species weekly over 3 years in a deciduous forest in east central Illinois, USA. Our objectives were: (1) to quantify for each species the extent of differences in leaf phenology between juvenile and conspecific adult trees; and (2) to determine the extent of phenological differences between juvenile Aesculus glabra Willd. and Acer saccharum Marsh. trees in understory and gap microhabitats. All species displayed phenological differences between life stages. For 10 species, bud break was significantly earlier, by an average of 8 days, for subcanopy individuals than for canopy individuals. In 11 species, completion of leaf expansion was earlier, by an average of 6 days, for subcanopy individuals than for canopy individuals. In contrast, there were no significant differences between life stages for start of senescence in 10 species and completion of leaf drop in nine species. For eight species, leaf longevity was significantly greater for subcanopy individuals than for canopy individuals by an average of 7 days (range = 4-10 days). Leaf phenology of subcanopy individuals of both Aesculus glabra and Acer saccharum responded to gap conditions. Leaf longevity was 11 days less in the understory than in gaps for Aesculus glabra, but 14 days more in the understory than in gaps for Acer saccharum. Therefore, leaf phenology differed broadly both between life stages and within the juvenile life stage in this community. A vertical gradient in temperature sums is the proposed mechanism explaining the patterns. Temperature sums accumulated more rapidly in the sheltered understory than in an open elevated area, similar to the canopy. Early leaf expansion by juvenile trees may result in a period of disproportionately higher carbon gain, similar to gains made during summer months from use of sun flecks.     

Cohabitation of tree holes by ants and breeding birds in a temperate deciduous forest

A previously unknown association of ants with birds breeding in tree holes is described. Ants Lasius brunneus Latreille (Hymenoptera: Formicidae) were found in c. 15% of nests of Parus major L. and Poecile palustris L. (Paridae) breeding in tree holes in the primeval deciduous forest located in the Bia?owie?a National Park (Poland). The ants preferably used holes located higher above the ground. As such holes are warmer than the unused holes or other nest sites, it is suggested that the ants cohabit holes utilized by the tits to gain thermal benefits.