Quantification and Understanding of Above and Belowground Biomass in Medium Saline Zone of the Sundarbans,Bangladesh: The Relationships with Forest Attributes

ABSTRACT

The relationship between stand attributes and biomass accumulation pattern in a mangrove forest has been intensively studied in this study. We assessed above (AGM) and belowground mass (BGM) and examined the relationship between forest attributes and aboveground mass in the Sundarbans, Bangladesh. The study was conducted with 18 plots having total area of 1.08 ha. The mean AGM and BGM of the study sites were 234.08 and 132.85 Mg ha^?1 respectively. H. fomes contributed the highest amount (82.9% of total AGM and 80.53% of total BGM) of above (193.56 Mg ha^?1) and belowground mass (107.09 Mg ha^?1) at the study site. Our study revealed structural attributes (tree diameter, height, and basal area) positively correlated with AGM. In contrast, species richness and species diversity negatively correlated with AGM. Our study indicated that lack of positive relationship between species diversity and AGM which may be attributable to high AGM of the dominant species (H. fomes) and may have a considerable consequence in AGM of the study area. Thus, maintaining large trees (DBH and height) rather than species diversity in the Sundarbans mangrove forest might be an effective approach for increasing aboveground mass.     

异速模型评估森林植被生物量有机碳储量

在孟加拉的吉大港南部森林地区,利用异速模型评估森林植被的有机碳的储量.异速模型被分别应用测试树木(被划分两个胸高直径级)、灌木和草本植物.采用基部面积估算胸高直径级为从> 5 cm 到 ≤ 15 cm 和> 15 cm树木的生物量有机碳储量模型最好,分别有很高的决定系数(胸高直径级> 5 cm 到 ≤ 15 cm 的r2 为0.73697,胸高直径级> 15 cm 的r2为0.87703),且回归系数(P = 0.000)显著.其它模型(包括采用树高,胸高直径,树高和胸高直径,以及综合树高、胸高直径和木材密度)的线性和对数关系都表现出很低的决定系数.分别建立了20种优势树种的异速模型,采用树木基部面积的模型都得到很高的决定系数值.单独采用灌木和草本植物总生物量的异速模型有较高的决定系数(灌木的r2 为0.87948,草本植物的r2 为0.87325),且回归(系数)性显著(P = 0.000).生物量有机碳的评估是复杂的和耗时的研究,本研究所建立的异速模型可以应用于孟加拉和其它热带(地区)国家的森林植被的有机碳储量的测算.     

Influence of canopy structure on the survival and growth of underplanted seedlings

This study explores the indirect relationship between forest structural measures and initial seedling survival and growth along a structural gradient between 64% to 92% canopy closure. The gradient was created by applying various levels of midstory removal to fifty 0.05 ha areas located within a mixed-hardwood riparian forest corridor. Twelve yellow-poplar (Liriodendron tulipifera L.) and cherrybark oak (Quercus pagoda Raf.) containerized seedling pairs were underplanted within each area. Canopy closure was estimated using hemispherical photography; height-to-canopy and basal area were recorded at each seedling pair. Survival, basal diameter, and height were monitored through two growing seasons. Species-specific mortality and height growth models were developed for one and two growing seasons following underplanting. The interaction of height-to-canopy and basal area along with canopy closure were found to be the most strongly related to mortality. Height to the forest canopy and initial seedling size explained the most variance in height increment. Although the height increment models possess limited predictive power (R² range from 0.22 to 0.36), both mortality and growth analyses emphasize the importance of quantifying vertical canopy structure, along with the more commonly considered horizontal measures of forest structure (basal area and stem density), when evaluating seedling development beneath a forest canopy.     

Forest variable estimation using photogrammetric matching of digital aerial images in combination with a high-resolution DEM

Abstract

The rapid development in aerial digital cameras in combination with the increased availability of high-resolution Digital Elevation Models (DEMs) provides a renaissance for photogrammetry in forest management planning. Tree height, stem volume, and basal area were estimated for forest stands using canopy height, density, and texture metrics derived from photogrammetric matching of digital aerial images and a high-resolution DEM. The study was conducted at a coniferous hemi-boreal site in southern Sweden. Three different data-sets of digital aerial images were used to test the effects of flight altitude and stereo overlap on an area-based estimation of forest variables. Metrics were calculated for 344 field plots (10 m radius) from point cloud data and used in regression analysis. Stand level accuracy was evaluated using leave-one-out cross validation of 24 stands. For these stands the tree height ranged from 4.8 to 26.9 m (17.8 m mean), stem volume 13.3 to 455 m³ ha^?1 (250 m³ ha^?1 mean), and basal area from 4.1 to 42.9 m² ha^?1 (27.1 m² ha^?1 mean) with mean stand size of 2.8 ha. The results showed small differences in estimation accuracy of forest variables between the data-sets. The data-set of digital aerial images corresponding to the standard acquisition of the Swedish National Land Survey (Lantmäteriet), showed Root Mean Square Errors (in percent of the surveyed stand mean) of 8.8% for tree height, 13.1% for stem volume and 14.9% for basal area. The results imply that photogrammetric matching of digital aerial images has significant potential for operational use in forestry.     

Practical large-scale forest stand inventory using a small-footprint airborne scanning laser

Mean tree height, dominant height, mean diameter, stem number, basal area and timber volume of 116 georeferenced field sample plots were estimated from various canopy height and canopy density metrics derived by means of a small-footprint laser scanner over young and mature forest stands using regression analysis. The sample plots were distributed systematically throughout a 6500 ha study area, and the size of each plot was 232.9 m². Regressions for coniferous forest explained 60–97% of the variability in ground reference values of the six studied characteristics. A proposed practical two-phase procedure for prediction of corresponding characteristics of entire forest stands was tested. Fifty-seven test plots within the study area with a size of approximately 3740 m² each were divided into 232.9 m² regular grid cells. The six examined characteristics were predicted for each grid cell from the corresponding laser data using the estimated regression equations. Average values for each test plot were computed and compared with ground-based estimates measured over the entire plot. The bias and standard deviations of the differences between predicted and ground reference values (in parentheses) of mean height, dominant height, mean diameter, stem number, basal area and volume were ?0.58 to ?0.85 m (0.64–1.01 m), ?0.60 to ?0.99 m (0.67–0.84 m), 0.15–0.74 cm (1.33–2.42 cm), 34–108 ha^?1 (97–466 ha^?1), 0.43–2.51 m² ha^?1 (1.83–3.94 m² ha^?1) and 5.9–16.1 m³ ha^?1 (15.1–35.1 m³ ha^?1), respectively.     

Comparing an individual tree growth model for <Emphasis Type="Italic">Pinus halepensis</Emphasis> Mill. in the Spanish region of Murcia with yield tables gained from the same area

To support forest management decisions on converting stands from even-aged to uneven-aged management, tree models are needed that can simulate both options. Therefore, an individual tree model for Pinus halepensis Mill. is compared with the respective yield table. The individual tree model is built from data of two Spanish forest inventories in this province and evaluated with published yield tables, which have been built from permanent observational plots. The individual tree model consists of a basal area increment model and a height increment model, both based on two measurements of about 4,000 trees from single species stands of Aleppo pine. The R2 values of 0.362 and 0.107 for the basal area increment model and the height increment model, respectively, are within the range of other published models of this type. Comparing the model with the yield tables for different site indices, we find that our model matches the yield table well. Only in higher ages where the database for the yield table was poor, our model indicates higher basal area growth rates than the yield table.     

Response of Three Young Douglas-Fir Plantations to Forest Fertilization with Low Rates of Municipal Biosolids

Abstract

The growth of three young (5-6 years since planting) Douglas-fir plantations fertilized with a single low-rate application of municipal biosolids, ranging from 17-19 dry Mg ha^-1, was studied. Stand measurements showed increases in DBH, height, average and total basal area, average and total volume and average and total dry-weight differences in the biosolids-fertilized vs. control. For instance, average per tree DBH was 14.7% greater than the controls, height by 2.7%, per stem basal area by 27%, per stem average volume by 33%, per stem average dry weight by 38%, basal area per hectare by 28%, volume per hectare by 32% and dry weight per hectare by 36% compared to untreated controls. Comparing differences in volume increases from initial growth periods with the latest growth period indicates that volume increases due to the biosolids treatment are continuing and apparently increasing over time. However, none of these observed differences were statistically significant (0.05 level) due to high variation within and between stands.     

Determination of number of stems in coniferous forest stands by means of aerial photo‐interpretation

Two models for determination of the number of stems per hectare in forest stands (N) from attributes derived by aerial photo‐interpretation were developed. The models relied on the assumption that N could be determined by dividing the total stand volume per hectare with the volume of the “average tree”; defined by stand mean height and the diameter corresponding to mean basal area of a stand. Input variables of the models were stand mean height, crown closure and site quality. Additionally, model II required input of average stand volume per hectare and average mean diameter derived from stratified field sample plot inventories. Material for 143 coniferous stands was used for the testing of the models. The stands were recorded by intensive field measurements. Aerial photographs at the approximate scale of 1:15 000 were used for photo‐interpretation. The N value was underestimated in model I by 5.4–47.0%. The standard deviation for the differences was 15.2–26.2% for mature stands and 41.4–44.2% for young thinning phase stands. For model II, the mean difference between the predicted and observed N value was in the range ‐16.1% to 12.2%.     

Modelling irregular and multimodal tree diameter distributions by finite mixture models: an approach to stand structure characterisation

Tree structural diversity is assessed by modelling stand diameter at breast height (DBH) distribution. The aim of this study was to verify the suitability of a mixture of two- and three-component Weibull and gamma models for describing irregular and multimodal DBH distributions. Investigations were carried out in natural Abies alba Mill. and Fagus sylvatica L. stands, representing the growing-up stage, in the Świętokrzyski National Park (Central Poland) and in the Pieniny National Park (Southern Poland). Sample plots (0.25–0.5 ha in area) were selected from a database used for forest dynamics analysis. The parameters of the mixture model were estimated using maximum likelihood estimation (MLE) and a combination of an expectation–maximisation (EM) algorithm with a Newton-type (NT) method. The multistart method (MM) of choosing initial values was used. The DBH distributions in mixed Abies–Fagus forests of complex structure were accurately approximated by two- and three-component mixture models. The root mean square error (RMSE) values varied from 3.4515 to 7.6682 for the Weibull mixture and from 3.6457 to 7.1419 for the gamma mixture. The results indicated that the mixture Weibull and gamma models were, in general, equally suitable for modelling irregular and multimodal DBH distributions. Finite mixture models may be used to characterise tree structural diversity, especially in uneven-aged stands of complex structure.     

Prediction of tree height,basal area and stem volume in forest stands using airborne laser scanning

The aim of this study was to develop prediction models using laser scanning for estimation of forest variables at plot level, validate the estimations at stand level (area 0.64 ha) and test the effect of different laser measurement densities on the estimation errors. The predictions were validated using 29 forest stands (80×80 m²), each containing 16 field plots with a 10 m radius. For the best tested case, mean tree height, basal area and stem volume were predicted with a root mean square error of 0.59 m (3% of average value), 2.7 m² ha^?1 (10% of average value) and 31 m³ ha^?1 (11% of average value), respectively, at stand level. There were small differences in terms of prediction errors for different measuring densities. The results indicate that mean tree height, basal area and stem volume can be estimated in small stands with low laser measurement densities producing accuracies similar to traditional field inventories.     

Tree and stand structure of the non-native Pinus contorta in relation to native Pinus sylvestris and Picea abies in young managed forests in boreal Sweden

Managed forest stands are typically younger and structurally less diverse than natural forests. Introduction of non-native tree species might increase the structural changes to managed forest stands, but detailed analyses of tree- and stand-structures of native and non-native managed forests are often lacking. Improved knowledge of non-native forest structure could help clarify their multiple values (e.g. habitat for native biodiversity, bioenergy opportunities). We studied the structural differences between the introduced, non-native Pinus contorta and the native Pinus sylvestris and Picea abies over young forest stand ages (13–34 years old) in managed forests in northern Sweden. We found that P. contorta stands had greater mean basal areas, tree heights, diameters at breast height, and surface area of living branches than the two native species in young stands. The surface area of dead attached branches was also greater in P. contorta than P. abies. Although this indicates greater habitat availability for branch-living organisms, it also contributes to the overall more shaded conditions in stands of P. contorta. Only one older 87 years old P. contorta stand was available, and future studies will tell how structural differences between P. contorta and native tree species develop over the full forestry cycle.     

Soil organic matter in European forest floors in relation to stand characteristics and environmental factors

Published data on the quantities of organic matter in the forest floor (SOM_fl) of European forest stands were collated. Studies of SOM_fl were included if the total sampled surface was at least 0.15 m², the surface of a single sample was at least 100 cm², live material was sorted out, and ash or carbon content was determined. Data from 59 forest stands were compared with regard to stand characteristics and environment. Using a single variable, tree genus was most important for the amount of SOM_fl (R ² _adj = 0.34). The amounts in spruce stands (41 × 10³ kg ha^?1) were significantly higher than those in larch, Douglas fir, oak and birch stands (4–11 × 10³ kg ha^?1). The best significant multiple model was genus combined with stand age and basal area (R ² _adj. = 0.72). The importance of litter quantity and quality for differentiating the tree genera is discussed.     

Structure of a Central-European mountain spruce old-growth forest with respect to historical development

This study examines the structural characteristics of the tree layer, dead wood, canopy openings, and regeneration patterns of a spruce old-growth forest in the Bohemian Forest, Czech Republic. An old-growth stand with minor human influence and a stand that was presumably logged about 200 years ago were analyzed and compared, as some forest managers considered the presumable human impact as a reason for salvage logging. Even though the stands differed in tree density, height and DBH structure, it was not possible to conclude whether it was due to management history or the environmental differences. The volume of dead wood also differed between the stands. There was about 142 and 83 m³ ha⁻¹ of dead wood in the old-growth stand and presumably logged stand, respectively. The amount of dead wood found in the old-growth stand was comparable with values reported from spruce old-growth stands across Central Europe. In both stands, many canopy trees were arranged in linear patterns, which was a result of spruce regeneration on nurse logs. This suggests that the origin and development of the stands were characterized by natural processes and during the past 200 years typical old-growth structural characteristics have already evolved.     

Volume equations for poplars growing on farmland in Sweden

Abstract

Volume equations for poplars growing on farmland in Sweden. The aim of the study was to construct a simple volume equation for poplar and to evaluate the equation together with seven published equations. Data from 72 poplars were collected from 37 stands in central and southern Sweden (lat. 55–60°N). The stands’ mean age and density was 21 years and 940 stems ha^?1, respectively and the trees’ mean diameter at breast height (DBH) was 25 cm. One simple volume equation with three parameters to be estimated for poplar was constructed with DBH and total height (H) as independent variables. This equation was analyzed and compared to seven published simple equations. The best performance, in terms of low values of Root Mean Square Error (RMSE) and Absolute Bias, was achieved by the equations developed by Eriksson and by Scott. However, due to these equations high level of multicollinearity an alternative recommendation was made where the constructed equation is recommended.     

Crown radius of pedunculate oak (Quercus robur L.) depending on stem size,stand density and site productivity

Crown size is a good indicator of the growth potential of trees and is often used in forest management for outlining thinning guidelines or constructing forest growth models. The aim of this study was to analyse mean crown radius as a function of stem size, stand density and site productivity in even-aged stands of pedunculate oak (Quercus robur L.). Data included measurements of 620 trees from 53 plots in nine thinning experiments and one operational stand in Sweden, Denmark and Great Britain, representing a wide spectrum of thinning practices ranging from the strictly unthinned control to extremely heavy thinning with essentially solitary trees. Three sets of models were constructed based on different predictor variables, including indicators of individual stem size (diameter at breast height, DBH), stand density/thinning grade (quadratic mean diameter and stand basal area) and site productivity (stand top height). Preliminary results indicated a significant effect of DBH and (nominal) thinning grade on crown radius. The response pattern of the final models indicated an increasing crown radius with increasing DBH, with increasing thinning grade (decreasing stand density) and with decreasing site productivity. The models are valid for predicting the crown radius of pedunculate oak in even-aged forest stands.     

Altitudinal variability of stand structure and regeneration in the subalpine spruce forests of the Pol’ana biosphere reserve,Central Slovakia

The structure of natural subalpine spruce forest in the Zadná Pol’ana massif of the Western Carpathians was analysed. We focused on the variability of different aspects of stand structure, tree decay and regeneration processes in altitudinal gradient. We used systematic sampling, covering an area of 2 km², to detect even subtle changes in stand structure within one forest type over a range of less than 200 m in elevation. Mean stand density was 290 trees (>7 cm DBH) per hectare, average basal area was 41 m² ha⁻¹, and the volume accumulation in living trees amounted to 500 m³/ha⁻¹. Stand volume decreased by more than 50% between 1,260 and 1,434 m a.s.l. This means for an increase of altitude of 100 m that stand volume decreased by nearly 200 m³. Neither stand density nor basal area was related to elevation. Maximum tree height was strongly correlated to elevation, and it decreased on average by 6 m for each 100 m increment of altitude. No significant changes in the maximum spruce diameter were recorded in relation to the elevation gradient. Spatial distribution of trees was biased toward regularity at lower altitudes. Tree clustering increased with increasing altitude. The stock of coarse woody debris (CWD) decreased slightly along the altitudinal gradient, but changes were not significant. Density of spruce saplings and their number growing on CWD significantly increased across the elevation gradient. Despite the fact that the analysed forest tract was relatively large, highly variable in respect to environmental factors, and that stand volume, spatial structure, and tree height displayed strong variability along the elevation gradient, the diameter structure of stands and regeneration measures were uniform. Our results suggest that the recruitment of new trees in the Zadná Pol’ana subalpine spruce forest is not temporally continuous even at a scale of several square kilometres.     

Application of the 3PG forest growth model to a <Emphasis Type="Italic">Eucalyptus globulus</Emphasis> plantation in Northwest Spain

This paper presents an assessment of the ability of the 3PG process-based model to simulate height (h) and diameter at breast height (DBH) growth in three plots located in a Eucalyptus globulus plantation on the Atlantic coast of Galicia (Northwest Spain). The obtained results show how this model estimates adequately height (h) and DBH growth, and determination coefficients (R ²) between observed data and data predicted by the model are always higher than 0.97, root mean square error (RMSE) of 0.45–1.05 cm in DBH and 0.46–0.85 in h, and a model efficiency (EF) close to 1 (0.98–0.99). In order to improve the results predicted by the model, we propose modifying the nS and fullCanAge parameter values given by default so as to adjust growth rates in the plantation to the weather conditions in the site. The ability of the model to discriminate growth rates in the three plots must be noted, even when plots are located in the same plantation and they only show differences in aspects related to site factors. Given the obtained results in the three plots used in this study, its simplicity and the small number of parameters needed as input data, the 3PG model stands out as a very useful tool for forest plantation management.     

Pioneer tree responses to variation of soil attributes in a tropical semi-deciduous forest in Brazil

The occurrence of pioneer tree species inside tropical forests is usually associated with canopy openness due to disturbances. The distribution of these species under different environmental conditions, aside from light presence, can be influenced by other variables such as soil attributes, water availability, and non-arborous species presence. This work evaluates pioneer tree distribution in the Pindorama Biological Reserve, Brazil, with respect to altitude, soil attributes, and non-arborous species in 65,400-m² plots in two toposequences of semi-deciduous forest. We evaluated the physical and chemical soil attributes altitude, basal area, height, and number of individuals of tree species with diameter at breast height (DBH) ≥5 cm in a randomly chosen quadrant in each plot. Pioneer trees were characterized by the following higher occurrence species: Acacia polyphylla, Aloysia virgata, Casearia sylvestris, and Croton floribundus. Cluster analysis suggested five similar groups among sampling plots. For each group, the mean of altitude, physical and chemical soil attributes, and degree of non-arborous species infestation was calculated. Principal components analysis correlated variables with pioneer tree data. C. floribundus occurred at low altitudes at lower or higher fertility, C. sylvestris occurred in lower fertility plots, and A. polyphylla and A. virgata occurred in higher altitude plots.     

Stem volume models for Cryptomeria japonica of Jeju Island,Korea

The accurate estimation of the stem volume of trees is an important task for forest managers. Two common variables used in stem volume models are the diameter at breast height (DBH) and the total height of a tree. These two variables can be used together, or only the DBH can be used. The present study was conducted to develop a simple volume model for Cryptomeria japonica found in Jeju Island, South Korea. The performance of six models using only the DBH and using both the DBH and total height as variables were evaluated. Fit statistics including coefficient of determination (R²), standard error of the estimate (SEE), mean bias (ē), absolute mean difference (AMD), Akaike information criterion (AIC), and Akaike weight (AIC_w) were used to evaluate the models. Lack-of-fit statistics were also used for further evaluation. The results showed that the models using the DBH and total height as variables performed better than the models using only the DBH as a variable.     

Structure and composition of streamside management zones following reproduction cutting in shortleaf pine stands

Streamside management zones (SMZs) in the Ouachita Mountains of Arkansas and Oklahoma are frequently established along headwater ephemeral and intermittent streams to protect water quality, provide wildlife habitat, and increase landscape diversity. To better understand the function of these riparian forest corridors, we characterized the tree density and composition, forest floor mass, and downed woody debris volume within SMZs located in undisturbed, mature, upper mid-slope shortleaf pine stands and then compared these attributes to those in upland portions of these stands. In addition to evaluate the impact of upland forest harvesting on these riparian corridors, we compared the amounts and distribution of forest floor, downed woody debris (DWD), snags, and windthrows in SMZs within shortleaf pine stands that had been clearcut, had a shelterwood harvest, and had no recent management activity (uncut stands). Total tree and hardwood basal area was significantly higher (4.4 and 4.2 m² ha⁻¹) while forest floor mass was significantly lower (0.5 kg m⁻²) in the SMZs than in the upland portion of the undisturbed stands. Five years following the reproduction cuttings tree basal area, DWD volume, and forest floor mass within SMZs did not significantly differ among stands that had or had not been harvested. Snag density was significantly lower within SMZs that occurred in clearcut stands compared to those in the uncut or shelterwood stands. Harvesting activities that retain few or no residual trees appear to increase the degradation of snags. This study provided evidence that clearcutting may also increase the risk of windthrow in SMZs as well. There was little difference in the distribution of forest floor within SMZs regardless of whether the stand was harvested or the type of harvesting that occurred in the stand. However, DWD amounts were higher near the SMZs edge than in the interior of the SMZs with the greatest differences in distributions in stands that were clearcut.