不同沙丘部位和不同结皮类型对土壤种子库的影响

采用萌发法对古尔班通古特沙漠沙垄不同部位生物结皮类型的种子库进行研究,发现不同沙丘部位和结皮类型种子分布特征有一定的规律性:藻类结皮的种子数量、丰富度和多样性均显著高于地衣和苔藓结皮。生物结皮从藻类到地衣和苔藓的演替,有降低种子数量和种子多样性的现象。在不同沙丘部位,沙丘顶部流沙的种子数量和多样性较低,与丘顶流沙相比沙丘底部的藻类结皮显示出较高的丰富度和多样性,但沙丘底部的地衣和苔藓结皮与丘顶流沙相比未表现出明显的差异。背风坡中部的藻类结皮种子多样性较高,显著高于迎风坡中部的藻类结皮。     

Spatial heterogeneity of plant species on the windward slope of active sand dunes in a semi-arid region of China

Species richness and abundance are two important species diversity variables that have attracted particular attention because of their significance in determining present and future species composition conditions.This paper aims to explain the qualitative and quantitative relationships between species diversity pattern and grain size(i.e.size of the sampling unit),and species diversity pattern and sampling area,and to analyze species diversity variability on active sand dunes in the Horqin Sandy Land,northeastern Inner Mongolia,China.A 50 m×50 m sampling plot was selected on the windward slope,where the dominant species was annual herb Agriophyllum squarrosum.Species composition and abundance at five grain sizes were recorded,and the species-area curves were produced for thirteen grain sizes.The range of values for species abundance tended to increase with increasing grain size in the study area,whereas,generally,species richness did not follow this rule because of poor species richness on the windward slope of active sand dunes.However,the homogeneity of species richness increased significantly.With the increase in sampling area,species abundance increased linearly,but richness increased logarithmically.Furthermore,variograms showed that species diversity on the windward slope of active sand dunes was weakly anisotropic and the distribution pattern was random,according to the Moran Coefficient.The results also showed that species richness was low,with a random distribution pattern.This conflicts with the results of previous studies that showed spatial aggregation in lower richness in a sampling area within a community and inferred that the physical processes play a more important role in species diversity than distribution pattern on active sand dunes.Further research into different diversity patterns and mechanisms between active sand dunes and interdune lowlands should be conducted to better understand biodiversity conservation in sand dune fields.     

The influence of the soil management systems on aboveground and seed bank weed communities in olive orchards

The biodiversity of farmed landscapes is, in the context of agricultural intensification, a key aspect with regard to improving the sustainability of agroecosystems. Olive groves are undergoing rapid changes because of the spread of intensive farming systems, which may have negative environmental impacts. This paper reports a survey on the aboveground flora and seed banks in five olive groves located in Andalusia (Southern Spain). In this study, the following three management systems have been compared: no‐tillage, with the mowing of spontaneous weedy vegetation; no‐tillage, with the mowing of planted cover crops (Poaceae); and conventional tillage practices. Results showed that coverage and an abundance of vegetation are favored by spontaneous weedy vegetation with mowing management, while the richness of aboveground species was affected by landscape diversity and the presence of edges, which increases the richness and diversity of aboveground flora species in olive groves. Seed bank composition showed a low relationship with aboveground flora in the three cover crop management systems. The multivariate analysis performed pointed to those seed species that have a major influence on the aboveground flora communities of each of the three agricultural systems. The seed bank was clearly impoverished in terms of both abundance and species richness after the long‐term conventional tillage practices. We conclude that the intensive long‐term conventional tillage dramatically reduces weed communities in olive orchards and the subsequent ecosystem services provided by them.     

黄土丘陵沟壑区植被自然恢复过程中土壤种子库特征

采用野外调查取样和室内实验相结合的方法,研究了黄土丘陵沟壑区植被自然恢复过程中土壤种子库的物种组成、密度、物种多样性以及相似性特征.结果表明:阴、阳坡10个样地中,共萌发了55种2203株幼苗,隶属于23科48属;总密度达到5 505±625.537粒/m2,且0～5 cm土层远远大于5～10 cm土层的土壤种子库密度,约是其4.6倍;丰富度、多样性、均匀度指数均未与恢复年限达到显著相关水平,但生态优势度随恢复年限的增加呈上升趋势,且阳坡上升的趋势比阴坡明显;组成成分相似性系数变化范围为0.0952～0.7143,恢复年限相距越大,其相似性越小,说明植被恢复演替的过程也是土壤种子库空间异质性加大的过程.     

新疆野生樱桃李天然林土壤种子库特征

采取野外调查、网筛分选和温室萌发相结合的方法,对新疆霍城县大西沟山区野生樱桃李天然林的土壤种子库物种组成、密度和生活史型、垂直空间分布、物种多样性及其与地上植被间关系进行了研究。结果表明：① 试验鉴定出的幼苗共有48种,隶属于19科,且一年生植物多达20种（41.7%）,地下芽植物为15种（31.3%）;② 种子库中种子的数量自上而下呈先增加后减少的特征,有超过90%的种子分布在距地表5 cm以上的表层土壤及枯枝落叶层中,近90%的樱桃李种子分布在距地表2 cm以上的表层土壤及枯落层中;③ 土壤种子库中草本层物种的Margalef指数、Shannon-Wiener指数均为最高,其次为乔木层和灌木层,种子库物种多样性主要受草本层丰富度的影响;④ 野生樱桃李天然林土壤种子库与地上植被的相似性系数为0.315,表明地上植被对土壤种子库的贡献较大。但是,由于过度放牧,使得种子库的物种多样性在垂直空间上的分布规律受到干扰。     

短花针茅荒漠草原土壤种子库对不同放牧强度的响应

在内蒙古短花针茅荒漠草原,采用野外随机定点取样与室内萌发相结合的方法,对不同放牧强度下围封草地土壤种子库的密度、垂直结构、多样性、丰富度、均匀度、以及种子库与地上植被的相似性进行了研究。结果表明,在各试验区土壤种子库主要分布在0～20cm土层,占种子库总密度的87%～96%;随着放牧强度的增加,土壤种子库总密度减少,同时地上植被与土壤种子库的相似物种数减少,种子库组成的相异性增加;轻度放牧对土壤种子库中一二年生草本所占比例影响较小,而中度和重度放牧使土壤种子库中一二年生草本所占比例增加;土壤种子库的物种丰富度、多样性、均匀度均随放牧强度的增加而减小。     

Effects of vegetation cover on recruitment of Ulmus pumila L. in Horqin Sandy Land,northeastern China

Understanding the effects of vegetation cover on seedling survival is helpful for promoting vegetation restoration in environmentally fragile zones. This study was conducted in the desertified, moving sand dunes of Horqin Sandy Land, Inner Mongolia, northeastern China. We hyphothesized that(1) seed density(i.e., number/m2) increases as vegetation cover increases, and(2) there will be more surviving seedlings in locations with higher vegetation covers. Total vegetation cover and initial densities of seeds, germinated seeds and surviving seedlings of Ulmus pumilia were evaluated under various vegetation covers in trying to clarify the effects of vegetation cover on the early stages of the plant life history. In agreement with the first hypothesis, initial seed densities were greater(P<0.05) under higher vegetation covers. The relationship between vegetation cover and initial seed density was represented by a quadratic regression, where a threshold occurred with a vegetation cover of 36%(P<0.05). The higher total vegetation covers, however, did not result in increased densities of germinated seeds(P>0.05), which on average represented 16.7% of initial seed densities. Even more, three months after the study initiation, total vegetation covers were similar(P>0.05) at all positions in the dunes, and they determined a similar number(P>0.05) of surviving seedlings at those positions(i.e. the second hypothesis had to be rejected). The mean number of seed- lings that survived at all positions was only 4.5% of germinated seeds. The number of surviving elm seedlings(0 to 1.7 seedlings/m2) under various vegetations covers(12.2% to 20.8%) at all dune positions by late summer would most likely not contribute to vegetation restoration in the study area.     

Effect of spatial scale and topography on spatial heterogeneity of soil seed banks under grazing disturbance in a sandy grassland of Horqin Sand Land, Northern China

Soil seed banks play an important role in the distribution and composition of plant communities in semiarid grassland ecosystems.However,information on how spatial scale influences the spatial heterogeneity of soil seed banks in a grassland under grazing disturbance is still lacking.Based on field sampling and greenhouse germination,we measured the species composition and seed density of soil seed banks at different spatial scales (30 m×30 m,30 m×60 m and 30 m×90 m)along a topographical gradient in a sandy grassland in Horqin Sand Land, Northern China.By applying geostatistical methods,we examined how spatial scale and topography affected the spatial distribution of soil seed banks in the study area.Our results showed that the total number of species in soil seed banks,as well as the number of dominant annuals,increased with the increase of spatial scales.Seed density in soil seed banks decreased with the increase of spatial scales due to an increase in the slopes and relative heights of the sampling points.Geostatistical analysis showed that the relative structural variance(C/(C0+C))of seed density and species richness were over 65%for all spatial scales,indicating that these variables had an obvious spatial autocorrelation and the spatial structured variance accounted for the largest proportion of the total sample variance.Spatial autocorrelation of seed density in soil seed banks increased with the increase of measured scales,while that of species richness showed a reverse trend.These results suggest that the total number of species in soil seed banks is spatial scale dependent and lower topography may accommodate more seeds.Spatial distribution of seed density in soil seed banks is also scale dependent due to topographic variation.Grassland management,therefore,needs to consider local grazing disturbance regime,spatial scale and topography.     

Effect of sand-fixing vegetation on the hydrological regulation function of sand dunes and its practical significance

Soil water content is a key controlling factor for vegetation restoration in sand dunes. The deep seepage and lateral migration of water in dunes affect the recharge process of deep soil water and groundwater in sand dune ecosystems. To determine the influence of vegetation on the hydrological regulation function of sand dunes, we examined the deep seepage and lateral migration of dune water with different vegetation coverages during the growing season in the Horqin Sandy Land, China. The results showed that the deep seepage and lateral migration of water decreased with the increase in vegetation coverage on the dunes. The accumulated deep seepage water of mobile dunes (vegetation coverage<5%) and dunes with vegetation coverage of 18.03%, 27.12%, and 50.65% accounted for 56.53%, 51.82%, 18.98%, and 0.26%, respectively, of the rainfall in the same period. The accumulated lateral migration of water in these dunes accounted for 12.39%, 6.33%, 2.23%, and 7.61% of the rainfall in the same period. The direction and position of the dune slope affected the soil water deep seepage and lateral migration process. The amounts of deep seepage and lateral migration of water on the windward slope were lower than those on the leeward slope. The amounts of deep seepage and lateral migration of water showed a decreasing trend from the bottom to the middle and to the top of the dune slope. According to the above results, during the construction of sand-control projects in sandy regions, we suggest that a certain area of mobile dunes (>13.75%) should be retained as a water resource reservoir to maintain the water balance of artificial fixed dune ecosystems. These findings provide reliable evidence for the accurate assessment of water resources within the sand dune ecosystem and guide the construction of desertification control projects.     

浑善达克沙地南缘沙化草地围封过程中土壤种子库与地上植被的耦合关系

在浑善达克沙地南缘,以围封不同年限的沙化草地(1年、4年、9年)为试验样地,并以未围封沙化草地作对照,深入研究了沙化草地围封过程中,土壤种子库与地上植被的耦合关系。结果表明,对沙化草地实施围栏封育,可显著提高土壤种子库与地上植被的物种丰富度;不同围封年限间,二者在植被群落组成上,均表现出较高的相似性,且相临阶段相似性较高,随围封年限差异的增加,相似性逐渐减小,但这一变化前者滞后于后者,表明草地在恢复演替中,前者较后者具有滞后性;4种类型样地之间,土壤种子库密度以及地上植被密度差异均极显著,且二者之间均存在显著正相关性,其关系均可用二次曲线描述。     

民勤荒漠区植被退化动态定位研究

白刺、沙蒿、麻黄是民勤荒漠区的主要植物种类,梭梭是当地主要固沙造林树种。定位监测表明:民勤5月下旬土壤含水率与灌木、草本植物和优势种密度及植被总盖度之间呈正相关关系。6~7月的降水量与灌木密度和植被总盖度之间呈正相关关系。5~7月的降水量和土壤水分是限制当地荒漠植被密度、盖度的关键因子。从群落类型看,深根性的麻黄和梭梭盖度比较稳定。从地貌看,流动沙丘和半固定沙丘上的植被较稳定,固定沙丘植被普遍严重退化。设置黏土沙障易形成沙面结皮,阻止降水下渗,不利于接收和保存有限的天然降水。     

青海湖东沙地不同植被恢复措施下土壤水分变化特征

土壤水分是沙地植被格局和过程改变的主要驱动力,也是干旱半干旱区植物生长最大的限制因子.沙漠地区的土壤水分表现出明显的空间异质性.以高寒半干旱沙地青海湖东沙地为研究区,以栽植人工固沙植物(小叶杨、樟子松、沙棘和乌柳)的沙丘和自然固定沙丘(沙蒿为优势种)作为研究样地,探讨不同固沙植物影响下的沙丘不同地貌部位土壤水分变化特征...     

古尔班通古特沙漠不同类型生物结皮对草本植物多样性影响

选择古尔班通古特沙漠的北部(一号点)、中部(二号点)、南部(三号点)3个不同样点的裸沙和藻结皮、地衣结皮与苔藓结皮3种生物结皮类型,对比研究了草本植物多样性的差异性及其主要环境影响因素.结果表明:(1)不同生物结皮类型的土壤理化性质有明显差异,土壤有机质、全氮、全磷、全钾含量以及黏粒、粉粒和细沙的含量随生物结皮演替显著...     

青海湖流域沙柳河草甸群落结构与数量特征

沿青海湖流域沙柳河河岸,选取多个断面进行样方调查,分析沿沙柳河距河流入湖处不同距离、垂直河岸方向上草甸群落结构、地上生物量以及物种多样性的变化特征,并讨论影响群落结构变化的生境因子。结果表明：① 在所调查的样地中,有草本植物52种,隶属39属,13科。平均总盖度为70%,平均地上生物量为131 g•m^-2。② 随着离湖距离的增加,Shannon-Wiener指数、物种丰富度、物种均匀度指数、地上生物量呈现先增加后略微减小的趋势,盖度的变化趋势是先增加后不变。土壤含水量和土壤含盐量、海拔等可能是影响植物群落空间分布差异的主要原因。③ Shannon-Wiener指数、物种丰富度与地上生物量呈现不明显的线性关系。     

Grain size distribution at four developmental stages of crescent dunes in the hinterland of the Taklimakan Desert,China

Although scientists have performed many studies on crescent (barchan) dunes in the Taklimakan Desert,few studies reported the changes in grain size at different development stages of crescent dunes.In order to evaluate the changing trends of surface sediment grain size with dune development,we investigated the grain size characteristics at four developmental stages (oval sand pile,shield dune,incipient crescent dune and mature crescent dune) of crescent dunes by measuring the morphology of sand dune and observing the near-surface wind regime.The dunes have developed in a wide inter-dune corridor between high compound longitudinal ridges in China's Taklimakan Desert.The surface sediments at four developmental stages of the crescent dunes were primarily composed of fine sands,followed by very fine and medium sands.Mean grain sizes ranged from 2.8 to 3.1 φ,with a unimodal distribution.The sands were moderately well-sorted,their distribution varied from platykurtic to very platykurtic,and symmetrical or skewed towards the fine particles.From oval sand piles through shield and incipient crescent dunes to mature crescent dunes,incipient grain size gradually increased,particles became finer,sorting became better,kurtosis and skewness increased.Grain sizes on the surface layer became coarser upwards from the toe of the windward slope and then became finer towards the bottom of the leeward slope.We found that the coarsest particles at different positions at the four developmental stages were different.The coarsest particles were distributed at the top of the oval sand piles and shield dunes,versus at the middle of the windward slope of the incipient and mature crescent dunes.Correlations between the mean grain size and other grain size parameters showed that as mean grain size became finer,sorting became better and kurtosis became wider,but skewness changed only slightly.In addition,grain size variation in the surface sediments correlated with the movement speed of the dunes in the study area.In the open ground among tall-complex longitudinal ridges in the hinterland of the Taklimakan Desert where aeolian environment is characterized by comparatively strong wind and unsaturated sand flow,faster dune movement corresponded to coarser grain size.     

Sand-fixing characteristics of Carex brunnescens and its application with straw checkerboard technique in restoration of degraded alpine meadows

Carex brunnescens(Pers.), a typical clonal species, is considered to be the only herb found to date that can develop on sand dunes in Maqu alpine region of northwestern China. However, the characteristics that C. brunnescens resists to harsh alpine environment have not been documented. In this study, we conducted a field investigation to determine the morphological, reproductive, and sand-fixing characteristics of C. brunnescens. Concomitantly, we transplanted the belowground rhizomes of C. brunnescens to sand dunes and compared the abilities to restore degraded alpine meadows among sand dunes that had no further treatment(SD+N), sand dunes that had straw checkerboard technique but no transplanted rhizomes of C. brunnescens(SD+SCT), and sand dunes that had both SCT and transplanted rhizomes of C. brunnescens(SD+SCT+P). We found that belowground vertical rhizomes and horizontal rhizomes(including branching rhizomes and main rhizomes) of C. brunnescens were highly developed and that population reproduction was dominated by horizontal rhizomes. C. brunnescens exhibited a significant sand-fixation effect under following conditions: population density was 145–156 ramets/m~2, vegetation cover was 31.2%–39.3%, total length of belowground rhizomes was 11,223 cm/m~2, total length of belowground first-order roots was 9161–10,524 cm/m~2, fresh weight of aboveground part was 198.5–212.6 g/m~2, and fresh weight of belowground part was 578.8–612.4 g/m~2. It should be particularly noted that SD+SCT+P treatment(sand dunes that had both straw checkerboard technique and transplanted rhizomes of C. brunnescens) was the best and SD+N(sand dunes that had no further treatment) was the worst in terms of following biotic indicators: total number of reproductive ramets, total number of belowground rhizomes, and fresh weight of aboveground and belowground parts of C. brunnescens, contents of soil organic carbon, available nitrogen, microbial biomass carbon, and microbial biomass nitrogen. It implies that applying SCT in sand dunes and transplanting belowground rhizomes to sand dunes with SCT could improve both soil fertility and growth of C. brunnescens. These results suggest that the SCT-promoted high reproductive abilities of belowground rhizomes of C. brunnescens can successfully facilitate the establishment of ramets and can thus be an effective strategy to restore degraded vegetation in Maqu alpine region of northwestern China.     

甘肃河西绿洲边缘积沙带所表征的气候环境信息

绿洲边缘积沙带的研究是一个新的科学问题。甘肃河西地区经过半个多世纪的防沙治沙,在绿洲边缘形成了一条积沙带。为了研究积沙带的形成和发育与气候环境因子之间的关系,对河西走廊东端至走廊中西部绿洲边缘的积沙带进行了调查。结果表明：① 年降水量是决定积沙带高度和宽度的关键因子;② 降水量是决定植被盖度的关键因子,降水对天然植被盖度的影响大于对包括人工植被在内的植被总盖度的影响,随着降水增加,背风坡植被盖度增加幅度大于迎风坡;③ 积沙带不同样点之间植被盖度的差异主要是由植株个体大小引起的,亦即降水对植株个体大小的影响大于对植株密度的影响。     

浑善达克沙地榆种子扩散及成苗影响因素分析

以浑善达克沙地榆为研究对象,以种子扩散规律及成苗率为主要研究内容,采用样线法调查了平沙地、沙丘迎风坡、丘间草地、稀疏灌木地、灌木地5种不同地表类型下沙地榆种子数量.分析种子分布数量与母树距离、地表状况、植被种类、盖度和高度及成苗率与土壤含水率的关系.结果证明:影响种子分布数量的主要因子是母树的距离,其他因子对沙地榆种子...     

Effects of temperature and light on seed germination of ephemeral plants in the Gurbantunggut Desert,China: implications for vegetation restoration

Seed germination is a key transitional stage in plant life cycle and is strongly regulated by temperature and light. Therefore, research on the effects of temperature and light on seed germination is extremely meaningful for vegetation restoration, especially in desert ecosystems. Seeds of 28 ephemeral plants collected from the Gurbantunggut Desert of China were incubated at different temperatures (5°C/1°C, 15°C/5°C, 20°C/5°C, 25°C/10°C and 30°C/15°C) in 12-h light/12-h darkness or continuous darkness regimes, and the responses of seed germination to temperature and light and the germination speed were studied in 2016. Results showed that seed germination percentage of the 28 ephemeral plants significantly differed to temperature and light. We classified the studied plants as the following groups based on their responses to temperature: 1 low temperature responsed plants, 12 moderate temperature responsed plants, 7 high temperature responsed plants, 4 non-responsed plants and 5 plants of no germination. It should be noted that Corispermum lehmannianum Bunge is sensitive to both moderate and high temperatures. There were 4 groups of plant in response to light, i.e., 7 light responsed plants, 10 dark responsed plants, 6 light non-responsed plants and 5 plants of no germination. Based on seed germination speed of the 28 ephemeral plants, we divided them into 4 patterns of germination, i.e., very rapid, moderately rapid, moderate and slow. Combining variations of temperature, precipitation and sand dune types in the study area, we suggested that very rapid and moderately rapid germinated plants could be used to moving sand dunes in early spring during vegetation restoration, moderate germinated plants could be used to semi-fixed sand dunes in late autumn, and slow germinated plants could be used to sand plain in summer. Thus, seedling establishment and vegetation restoration would be improved by considering seed germination characteristics of these ephemeral plants in the Gurbantunggut Desert, China.     

甘肃马鬃山地区种子植物多样性研究

对甘肃肃北马鬃山地区的种子植物多样性进行了调查,结果表明:马鬃山地区的种子植物有25科65属112种。其中裸子植物1科1属3种,被子植物24科65属112种;马鬃山地区的植被类型大致分为5个植被型组;马鬃山地区分布着国内罕见、甘肃省面积最大而且保存最完整的天然梭梭林;最后论述了马鬃山地区天然梭梭林的保护价值并且提出了保护建议。