The digestion by cattle of grass silages made with no additive or with the application of formic acid or formic acid and formaldehyde

The aim of the experiment was to assess the effect of applying, at commercially recommended rates, formic acid or formic acid/formaldehyde at ensilage upon the subsequent digestion of the silages by cattle. Three wilted grass silages were made from perennial ryegrass ensiled at a DM concentration of 206 g kg⁻¹ after a poor wilting period of 49 h without additive application and with application of 2.5 litres t⁻¹ of formic acid or of 4.5 litres t⁻¹ of a mixture of (gkg⁻¹) 500 formic acid, 200 sulphuric acid and 200 formatin giving an application rate of 15 g formaldehyde kg⁻¹ herbage crude protein (N × 6.25).
The silages were fed to cattle equipped with rumen cannulae and duodenal re-entrant cannulae. Results from analyses of silage composition and from the digestion of organic matter and N showed no major differences between silages. The efficiency of rumen microbial N synthesis, the rumen degradability of silage N (determined in vivo or in sacco ) and in vivo digestion of individual amino acids were also unaffected by additive treatment. These results indicate that poor wilting conditions before ensilage restricted the effectiveness of additive treatment; the ineffectiveness of formaldehyde in reducing the rumen degradability of silage N may also have been related to the low rate of formaldehyde application.     

The effect of supplements of fishmeal and lactic acid on voluntary intake of silage by calves

In order to assess the relative effects of protein content and acid on intake, grass silage was given ad libitum to twelve entire male calves, initially 4 months of age and 117 kg liveweight (LW), either alone or supplemented with fishmeal at 50 g dry matter (DM) per kg silage DM. These two diets were offered alone or with additions of lactic acid at 50 g per kg silage DM in a partial changeover design in which lactic acid levels only were reversed between two 28-d measurement periods. In the absence of lactic acid, fishmeal significantly increased absolute intake but not intake relative to LW. There was a significant interaction between treatments in that the addition of lactic acid depressed DM intake in the absence of fishmeal (23·9 vs 21·1 g kg^-1 LW) but not in its presence (23·2 vs 23·1 g kg^-1 LW). Addition of fishmeal increased the digestibility of DM from a mean of 0·708 to 0·744 (P < 0·05), of gross energy from 0·646 to 0·694 (P < 0·01) and of N from 0·422 to 0·592 (P < 0·001) but had no significant effect on the digestibility of cellulose (0·766 vs 0·788 for treatments without and with fishmeal respectively). Lactic acid addition had no effect on the digestibility of plant components and, further, interaction effects between treatments were not significant. Supplementation with fishmeal reduced the total time spent eating and ruminating (Ro value) from a mean of 284 to 240 min kg^-1 DM (P < 0·01) and increased urine pH from 8·34 to 8·50. Lactic acid influenced neither Ro value nor urine pH and the interaction effect did not reach significance for these two parameters. The results indicate that the addition of fishmeal can overcome a depressant effect of lactic acid on intake and it is suggested that the responses to treatment were mediated via changes in the ratio of energy to protein supply rather than as a result of treatment effects on digestive efficiency or acid-base balance.     

The effect of silage chop length on the voluntary intake and rumination behaviour of sheep

Two experiments were conducted with sheep to investigate the effect of silage chop length on voluntary intake and eating and ruminating behaviour. The silages, made from Italian rye-grass, either received no additive or formic acid at a rate of 4 g kg^?1 and were chopped to lengths of 5.3 cm (long), 1.8 cm (short) before ensiling or 1.8 cm (long/short) before feeding. The addition of the additive and short chopping improved the fermentation characteristics of the silage. Voluntary intake of silage dry matter and organic matter was higher with the short than with the long silage. Differences in silage digestibilities and cellulolytic activity did not explain the increase in silage intake. Sheep on the long silage had a lower rate of intake and a shorter ruminating time than on short silage, and thus the effective breakdown of the silage in the rumen was delayed, i.e. an increased latency time. Pseudo-rumination was significantly higher on the long than on the short silage.     

Protein supplementation of formic acid- and enzyme-treated silages. 2. Nitrogen and amino acid digestion

Two silages were made from primary growth perennial ryegrass and ensiled after the application of either formic acid or an enzyme mixture of cellulase and hemicellulase. Silage analysis showed both silages to be well preserved with low pH of 3·70 and 3·62 for the formic and enzyme treatments respectively. Formic acid-treated silage had a higher total amino acid concentration than enzyme-treated silage. The silages were offered to growing steers either as the sole diet or supplemented with rapeseed meal at 60 g or 120 g fresh weight kg⁻¹ silage DM offered, in a 6 × 6 Latin square arrangement.
Non-ammonia nitrogen and microbial nitrogen flows at the duodenum (g d⁻¹) were significantly ( P < 0·05) increased by supplementation of enzyme-treated silage compared with formic acid-treated silage (enzyme, 83·6, 58·7; enzyme + 60 g, 101·7, 75·3; enzyme + 120 g, 112·5, 80·7; formic, 91·9, 63·7; formic + 60g, 88·3, 67·9; formic + 120 g, 95·5, 67·1) respectively. Efficiencies of microbial protein synthesis were increased for supplemented enzyme-treated silage diets and values were reduced for supplemented formic acid-treated silage diets compared with the silage only diets (enzyme, 27·9; enzyme + 60 37·7; enzyme + 120 g, 38·6; formic, 33·7; formic + 60g, 31·2; formic + 120 g, 28·8). Total amino acid flow at the duodenum increased with supplementation of both silages; however, microbial amino acid flow increased significantly ( P < 0·05) with supplementation of enzyme-treated silage compared with formic acid-treated silage diets. Significantly greater amounts of cystine, methionine, alanine, valine and aspartic acid entered the small intestines of animals receiving supplemented enzyme silages compared with supplemented formic acid silages.     

Effect of acetic acid,caproic acid and tryptamine on voluntary intake of grass silage by growing cattle

The objective of this study was to identify and quantify fermentation end‐products, detected with chromatographic techniques, that were negatively related to intake of grass silage by cattle. Further, the aim was to verify the intake‐depressing effect of these compounds in a feeding trial. A set of twenty‐four silages that had been used in a previous study to model variations in intake owing to fermentation quality was reanalysed with liquid and gas chromatography. Known and unknown chromatogram peaks were subjected to a regression analysis to determine which were negatively related to intake. Compounds were identified and quantified using a liquid chromatography–mass spectrometry system; acetic acid (AcA), caproic acid and tryptamine were chosen for verification. Growing steers were offered wilted silage with these three compounds added, separately or as a mixture, in proportions similar to the maximum values detected in the silages of the previous study. Dietary addition of AcA, either separately or mixed with the other two compounds, reduced silage dry matter (DM) intake. However, the reduction in silage DM intake equalled the amount provided by the added substances, so that no differences in total DM intake were observed for any of the dietary treatments.     

The chemical composition and nutritive value of grass silages prepared with no additive or with the application of additives containing either Lactobacillus plantarum or formic acid

A mixture of perennial and hybrid ryegrasses(234 g DM kg^-1) was forage harvested and ensiled after a 24-h wilt in good ensiling conditions in 2-t capacity silos with no additive application (control) or with the application of either Lactobacillus plantarum , 4 × 10⁶ (g fresh weight of grass)^-1, or of 31 formic acid t^-1. Sufficient 2-kg capacity laboratory silos were also filled with grass to monitor the changes in chemical composition of the ensiled grass with time. In laboratory silos, inoculation with L. plantarum resulted in a more rapid fall in silage pH ( p < 0.001) and a more rapid production of lactic acid ( P < 0.001) than in the control silage. At the end of the storage period (laboratory silos, 80 d; 2-t silos, 200-300 d), the inoculated silos had lower pH ( p ammonia-N (g kg N¹) and acetic acid contents ( p < 0.01) and higher water soluble carbohydrate (WSC), lactic acid (P<00.1) and ethanol ( p < 0.05) contents than the control silage. The formic acid-treated silage had significantly lower contents of ammonia-N (g kg N^-1, p < 0.05), acetic and tactic acids (p<0.01) and higher contents of WSC and ethanol ( p < 0.01) than the control silage. When fed to wether sheep, the digestibilities of DM, organic matter and gross energy were not altered by additive treatment. The digestibility of modified acid-detergent fibre was lower for both the inoculated ( P < 0.01) and formic acid-treated silages ( p < 0.05). However, N retention was improved ( p < 0.05) by both additive treatments. Silage intake was improved ( p < 0.01) by additive treatment from 53.4 (control) to 58.0 (inoculated) and 60.4 (formic acid) g DM (kg live weight^0.75)^-1d^-1.     

The effects of different protein supplements on the utilization of untreated, formic acid-treated or enzyme-treated silages by growing steers

Three grass silages were made from perennial ryegrass ensiled without additive application (U) or with the application of formic acid (F) or an enzyme mixture of hemicellulases and cellulases (E).
Analysis of silages showed that both untreated and enzyme-treated silages had higher lactic acid concentrations than formic acid-treated silage. Enzyme-treated silage had lower levels of ADF and NDF but higher concentrations of residual WSC than other silages.
The silages were fed to growing steers supplemented with either rapeseed meal (RSM) (60 g kg⁻¹ silage DM) or fishmeal (FM) at a level isonitrogenous with RSM diets, so providing six diets (UR, UF, ER, EF, FR, FF). Organic matter intakes were similar, but ADF intakes were significantly ( P < 0·001) lower with enzyme-treated silage diets (UR, 1163; UF, 1160; ER, 1104; EF, 1035; FR, 1216; FF, 1213), as were intakes of NDF ( P < 0·01) (UR, 1946; UF, 1955; ER, 1877; 1772; FR, 2031; FF, 2041). Apparent whole tract digestibilities of organic matter were significantly ( P < 0·001) higher with enzyme-treated silages (UR, 0·644, UF, 0·644; ER, 0·668; EF, 0·678; FR, 0·633; FF, 0·633). Liveweight gains were generally higher with treated silage diets and RSM supported a greater response than FM (UR, 0·496; UF, 0·498; ER, 0·567; EF, 0·489; FR, 0·543; FF, 0·506) with both enzyme and formic acid-treated silages, although none of these differences were significant.     

The effect of ammoniation using urea on the intake and nutritive value of chopped barley straw

Chopped barley straw was sprayed with a 10% urea solution at the rate of 400 ml kg^-1 straw and stored in sealed containers. Two experiments were conducted with this straw.
In experiment 1 the effect of ensiling periods of 1, 15, 30, 45, 60, 75 and 90 d on the in vitro digestibility, total N and free NH3-N of straw were examined. In vitro digestibility reached a maximum 45 d post-treatment and remained constant thereafter; the overall increase in digestibility of treated over untreated straw was 0·21 units. Forty-four% of the added N was retained in the straw and free NH3-N accounted for 31% of the total N. In experiment 2 when straw was given alone the voluntary intake and crude fibre (CF) and DM digestibilities of urea-treated straw were increased by 47, 40 and 26% respectively over the untreated straw. The digestibilities of the diets were similar when either treated or untreated straw formed 20% of the diets. The CF digestibility of the two high-concentrate diets was similar to that of the untreated straw. Treatment of straw with urea increased the digestible energy value of straw by up to 20%. Feeding urea-treated straw resulted in a significant increase of ruminal NH₃-N concentration without any change of rumen pH.     

The effect of formic acid on the rate of water loss from cut grass and lucerne

The drying rates of grasses and lucerne, following treatment with 1% formic acid, were compared with those of untreated crops. Material was fully exposed to a flow of air in a ventilated laboratory oven set at 23 °C. Treatment with formic acid initially enhanced the rate of drying of grass, but this was followed by retardation. This reduction in the rate of drying was caused by the rapid desiccation of laminae, which reduced their effectiveness in providing a pathway for water loss from the stem. The rate of water loss directly from the stem surface was not increased by formic acid treatment, probably because the enveloping leaf sheaths acted as a protective layer. In contrast, formic acid increased drying rate in lucerne. whose stems have no such protective layer. The results suggest that chemical desiccants. which have a contact action are unlikely to have any application in making hay from grass.     

The effect of herbage allowance upon the herbage intake and performance of suckler cows and calves

Twenty-four Hereford × Friesian cows and their South Devon cross calves were allocated to three herbage allowances allotted daily for three periods of 8 weeks in a Latin square design. The daily allowances were 17, 34 and 51 g dry matter per kg cow plus calf live weight. Milk production was depressed by 0·2 and 1·2 kg d^-1 at the medium and low allowances. The corresponding falls in liveweight gain were for cows 0·26 and 0·25, and for calves 0·27 and 0·24 kg d^-1. Residual sward height after grazing gave a better indication of the animals' reaction to sward conditions and the management imposed than actual herbage allowance. The quantity per unit area and the composition of material present were important factors influencing intake. Calves were unable to compete with their dams to maintain herbage intake at the lower allowances and therefore are likely to benefit from additional feeding or creep grazing when residual sward height falls below 6cm for periods in excess of 1–2 weeks.     

Effects of formic acid on silage fermentation, digestibility, intake and performance of young cattle

Data from twenty-two comparisons carried out at ADAS Experimental Husbandary Farms are used to compare untreated and formic acid-treated silages. Additive treatment led to an improved fermentation in some crops, particularly those of low DM concentration (<262 g kg^-1). Where this occurred there were associated benefits in silage digestibility (+0·234 units), intake (+16%) and the growth rate of young cattle (+0·28 kg d^-1). Where the fermentation of the untreated silage was good, both digestibility and animal performance associated with treated and untreated silages were similar. It is suggested that the justification for using formic acid in a commercial situation is thus restricted to occasions where the untreated crop would be liable to develop a clostridial fermentation. These may be when crops contain less than 35 g water-soluble carbohydrate kg^-1.     

The effect of cell wall degrading enzymes or formic acid on fermentation quality and on digestion of grass silage by cattle

A first cut of timothy, treated with water (untreated), formic acid (FA), cellulase + lactic acid bacteria (CB), cellulase + hemicellulase (CH) or cellulase + hemicellulase + a lignin-modifying enzyme (CHL), was ensiled in pilot-scale silos. Silages, except CB, were fed to four male cattle, each equipped with a rumen and duodenal cannula, in a digestibility trial designed as a 4 × 4 Latin square. The animals were fed a diet of 400 g of concentrate and 600 g of silage at a level of 70 g DM kg^?1 live weight (LW^0·75). All enzyme-treated silages were well-preserved with a more extensive fermentation than in FA silage. The quality of untreated silage was poorer as indicated by higher pH and ammonia-N content. The amount of effluent from enzyme-treated silages ranged from 116 to 127 g kg^?1; for FA and untreated silages values were 101 g kg^?1 and 80 g kg^?1, respectively. Total DM losses from enzyme-treated silages were higher than from FA silage (P < 0·05). No significant differences were noticed between silages in the apparent digestibility of organic matter (OM), neutral-detergent fibre (NDF), acid-detergent fibre (ADF) or nitrogen (N). The apparent digestibility of cellulose was higher with enzyme-treated silages than with FA silage (P < 0·05). The values for microbial N flow at the duodenum were 80·0, 91·9, 80·7 and 70·5g N d^?1, and for the efficiency of rumen microbial N synthesis 38·6, 47·6, 36·9 and 32·5 g N kg^?1 OM apparently digested in the rumen for untreated, FA, CH and CHL silages, respectively. In the rumen the molar proportion of propionate was higher (P < 0·01) and that of butyrate lower (P < 0·01) with enzyme-treated silages when compared with FA silage. The proportion of butyrate was also lower with untreated than with other silages (P < 0·01). The rumen residence time of NDF and ADF was longer (P < 0·05) with enzyme-treated silages than with FA silage.     

The effect of formic acid on the fermentation of ryegrass ensiled at different stages of growth and dry matter levels

In the first experiment the buffering effect of Italian and perennial ryegrass to fonnic acid was determined with three varieties (Leda, S24 and Presto) cut at weekly intervals throughout the growth cycle. The buffer curves showed no significant differences between the three varieties at any one stage of growth and the results were combined to produce regression equations predicting pH values for the three ryegrasses cut at first ear emergence (e.e.), 50% e.e. and 100% e.e. after addition of formic acid. In the second experiment Italian ryegrass was harvested at three stages of growth, wilted to four different DM contents and ensiled in laboratory silos with five levels of formic acid. The interaction of stage of growth, formic acid level and DM content on the water-soluble carbohydrates, ethanol and volatile N contents of tbe resultant silages is discussed.     

The herbage intake and performance of set-stocked suckler cows and calves

Groups of eight Hereford × Friesian cows and their South Devon cross calves were set stocked over a 24-week grazing season at 3·23 (low), 3·21 (medium) or 4·24 (high) cows ha^-1 together with their calves. For the first 8 weeks only two-thirds of the total area was grazed. Increasing the stocking rate from low to medium reduced daily milk yield and cow and calf liveweight gains by 1·2, 0·24 and 0·29 kg d^-1 respectively, and from medium to high by 1·2, 0·24 and 0·23 kg d^-1. The main sward factor influencing faecal output and herbage intake was the quantity of organic matter on the pastures but the digestibility of the herbage selected also exerted a significant effect on the intake of cows. Major depressions in the herbage intake of cows occurred once the average sward height fell below 7 cm. Output of calf live weight was 628, 658 and 743 kg ha^-1 for the 3 stocking rates from low to high, and for cows 246, 179 and 30 kg ha^-1. It was concluded that decisions on pasture management should be taken in relation to the cow rather than the calf on set-stocked swards.     

Comparison of the fermentation quality and nutritive value of sulphuric and formic acid-treated silages fed to beef cattle

Five experiments were carried out in the years 1980-1983 and 1986 to study the effect of treating grass at ensiling with sulphuric acid (850 g kg^?1) and formic acid (850 g kg^?1) additives alone, and in mixtures with or without formalin on the preservation of grass, in vivo digestibility in sheep, in-silo loss, intake and performance of finishing cattle. Primary growth grass was ensiled in experiments 1 (3–4 June 1980), 2 (12-15 June 1981) and 3 (31 May-2 June 1982), primary regrowth grass in experiment 4 (1-2 August 1983) and secondary regrowth grass in experiment 5 (7-10 October 1986). During the ensiling period within each experiment, approximately 60 t of unwilted, double-chopped, additive-treated or untreated grass was packed into covered concrete-walled 60-t capacity silos. The dry matter (DM) contents of the ensiled grass in experiments 1, 2 and 5 ranged from 155-180 g kg^?1 and were lower than those recorded in experiments 3 and 4 (214 g kg^?1). With the exception of grass ensiled in experiment 2, where water-soluble carbohydrate (WSC) contents were low, at 104 g kg^?1 DM, grass in all other experiments contained relatively high WSC contents ranging from 140-154 g kg^?1 DM. In experiments 1, 3 and 4 all silages were well-preserved. However, in experiment 2 the 450 g kg^?1 sulphuric acid-treated and formic acid-treated silages displayed significantly lower pH, buffering capacity (Bc) and ammonia nitrogen contents than the untreated silage. In experiment 5, the sulphuric acid-treated and formic acid-treated silages displayed significantly lower pH, Bc, ammonia nitrogen, butyrate and volatile fatty acid (VFA) contents than the untreated silage. Each of the silages was offered daily with various levels of a supplementary concentrate for approximately 70-d periods to twelve animals of mixed breed in experiments 1, 3, 4 and 5 and to fifteen animals in experiment 2. All animals weighed between 380-470 kg at the start of the experiments. In experiments 1, 2 and 3 there were no significant differences between silages for any of the intake or animal performance parameters. In experiment 4, cattle fed the formic acid-treated silage displayed significantly higher silage DM intakes and daily liveweight gains than those fed the sulphuric acid-treated and untreated silages; in experiment 5, cattle fed the formic acid-treated silage displayed significantly higher silage DM intakes than those fed the untreated silage. It was concluded that formic acid was a more effective silage additive than sulphuric acid. Increasing the level of supplementation significantly decreased silage DM intakes in cattle in experiments 3 and 4, and significantly increased daily liveweight gains and daily carcass in cattle in experiments 1, 3 and 4.     

The effect of heat-treatment and gamma radiation on the composition of unwilted and wilted lucerne silages

Three unwilted and three wilted silages were prepared from lucerne. Unwilted and wilted material was ensiled either without treatment (UWC and WC), after gamma irradiation (UWI and WI), or following heat-treatment and inoculation with Lactobacillus plantarum (UWH and WH). Gamma radiation was from a cobalt 60 source. The UWH material was heat-treated using steam, and WH material was treated with dry heat. At ensiling, irradiated material was considered to be sterile, and heat-treated material had lost 80% of its original protease activity. Fifteen 1-litre silos were prepared for each treatment (90 in total), and three from each treatment were opened after 1, 4, 14, 70 or 200 d ensiling. In irradiated silages, there was no accumulation of organic acids, but water soluble carbohydrate (WSC) concentration was almost double (116 g kg^-1dry matter) that at ensiling. This indicated extensive liberation of WSC from more complex carbohydrates during ensiling. The extent of silage fermentation in heated silages was similar to control silages. The WH silage had a higher proportion of the organic acids as lactic acid relative to the other treatments, possibly as a result of inoculation with L. plantarum. Proteolysis was unaffected by irradiation. However, heat-treatment markedly reduced the extent of proteolysis, particularly in the first four days post-ensiling. Deamination appeared to be related to the type of microbial fermentation rather than to the extent of proteolysis.     

The effect of an acid salt-type additive on the fermentation of grass silages made in bunker silos on commercial dairy farms in Wales

The fermentation characteristics and chemical composition of 57 first-cut and 30 second-cut samples of grass silages, made in bunker silos on commercial dairy farms in Wales in 1990, and treated with a nominal 61 t⁻¹ of an acid salt-type additive at ensilage, is described. Typical chemical composition of grass cut for ensilage was 156 g kg⁻¹ dry matter (DM) and 28 g kg⁻¹ water soluble carbohydrate (WSC), with 181 g (kg DM) ⁻¹ crude protein (CP) and 232 g (kg DM) ⁻¹ modified acid detergent fibre (MADF). The effect of additive use was to produce silages with DM 230 g kg⁻¹ pH 3·93, ammonia N 70 g kg⁻¹ total N, with residual WSC 35 g (kg DM) ⁻¹, lactic acid 83 g (kg DM) ⁻¹, total acids 118 g (kg DM) ⁻¹ and butyric acid 0·7 g (kg DM) ⁻¹. No significant differences were found between first- and second-cut silages. Silage fermentation was restricted (i.e. lactic acid less than 60 g kg DM⁻¹) in only 20% of the samples.
It is suggested that on commercial farms the application rate achieved may be insufficient to produce a restricted fermentation.     

Effect of stage of maturity and chop length on the chemical composition and utilization of formic acid-treated ryegrass and formic acid silage by sheep

Two areas of an early-heading perennial ryegrass cv. Cropper were harvested by either a precision-chop or a flail harvester at around 50% ear emergence (15 May 1978) and 14d later (29 May). Formic acid (85%) was applied at the rate of 2·2 litres t^-1. Mature crossbred wethers were used in a 2 × 2 factorial design to determine the effect of stage of maturity and method of harvesting (chop length) on the in vivo digestibilities of formic acid-treated grass in experiment 1 and formic acid silage in experiment 2. Apparent digestibility coefficients were determined at a fixed level of feeding for both grass and silage and at ad libitum access to feed for silage only. There were no significant differences in the concentrations of crude protein (CP), ether extract (EE), acid-detergent fibre (ADF) or acid-detergent lignin (ADL) in grass or silages of differing chop lengths but the later cut forages had significantly higher ADF and ADL concentrations and lower CP concentrations than the early-cut forages. The ADF and ADL concentrations were also higher in the silages than in the corresponding grasses. In general, the fermentation characteristics of precision-chopped silage were better than for the corresponding flail-cut silage but date of harvest was a more important determinant of quality and the late, flail-cut silage had the highest butyrate and ammonia N concentrations and the highest pH (411) of any treatment. There was a significantly higher intake of precision-chopped as compared with flail-cut silage with both the early-and the late-cut silage but there were no significant differences attributable to stage of maturity (i.e. date of harvest) or significant interaction between chop length and maturity. The slightly increased intake of early harvested, precision-chopped silage as compared with late precision-chopped silage was not significant. Dry matter digestibility (DMD) of the grass decreased at a rate of 0004 units d^-1 post 50% ear emergence. The results of experiment 2 indicated a decrease of 0·207 units d^-1 in silage fed at a similar level. The late-cut silage (DMD 0·292, mean of both harvesting treatments) thus had a significantly lower digestibility than the corresponding grass (mean DMD 0·247). Chop length had a variable influence on the DMD of both grass and silage fed at a fixed level but treatment differences were non-significant. However, a trend towards higher digestibility of flail-cut as compared with precision-chopped silage was apparent and this became statistically significant when the animals were allowed ad libitum access to feed. This may be a response to the generally lower intake of flail-cut silage.     

The effect of daily herbage allowance and sward characteristics upon the ingestive behaviour and herbage intake of calves under strip-grazing management

Measurements of herbage intake and ingestive behaviour were made on British Friesian calves 4–9 months old, which were strip-grazed on swards of S24 perennial ryegrass at three (Experiment A) or four (Experiment B) levels of daily herbage allowance in three (A) or four (B) 12-d periods. Herbage intake declined by approx. 18% as daily herbage allowance was reduced from 90 to 30 g DM per kg LW in both experiments, but the relationship between intake and allowance was not consistent between periods. Bite size and rate of biting were greater in calves on low than on high allowances after entry to a new strip of herbage but the positions were reversed towards the end of grazing. Reduced herbage intakes at low allowance resulted from reductions of approx. equal magnitude in bite size, rate of biting and grazing time, though not all of these effects were significant. It is suggested that these responses result from the increasing difficulty of prehending and ingesting herbage as swards are grazed down. The calves at low herbage allowances did not increase grazing time, probably because their behaviour was conditioned by anticipation of a new allowance of herbage. Bite size and daily herbage intake were greater on a spring sward than on an autumn sward, but within experiments herbage intake was not consistently related to any of the sward characteristics measured.