The effect of daily herbage allowance and sward characteristics upon the ingestive behaviour and herbage intake of calves under strip-grazing management

Measurements of herbage intake and ingestive behaviour were made on British Friesian calves 4–9 months old, which were strip-grazed on swards of S24 perennial ryegrass at three (Experiment A) or four (Experiment B) levels of daily herbage allowance in three (A) or four (B) 12-d periods. Herbage intake declined by approx. 18% as daily herbage allowance was reduced from 90 to 30 g DM per kg LW in both experiments, but the relationship between intake and allowance was not consistent between periods. Bite size and rate of biting were greater in calves on low than on high allowances after entry to a new strip of herbage but the positions were reversed towards the end of grazing. Reduced herbage intakes at low allowance resulted from reductions of approx. equal magnitude in bite size, rate of biting and grazing time, though not all of these effects were significant. It is suggested that these responses result from the increasing difficulty of prehending and ingesting herbage as swards are grazed down. The calves at low herbage allowances did not increase grazing time, probably because their behaviour was conditioned by anticipation of a new allowance of herbage. Bite size and daily herbage intake were greater on a spring sward than on an autumn sward, but within experiments herbage intake was not consistently related to any of the sward characteristics measured.     

The effect of herbage allowance upon the herbage intake and performance of suckler cows and calves

Twenty-four Hereford × Friesian cows and their South Devon cross calves were allocated to three herbage allowances allotted daily for three periods of 8 weeks in a Latin square design. The daily allowances were 17, 34 and 51 g dry matter per kg cow plus calf live weight. Milk production was depressed by 0·2 and 1·2 kg d^-1 at the medium and low allowances. The corresponding falls in liveweight gain were for cows 0·26 and 0·25, and for calves 0·27 and 0·24 kg d^-1. Residual sward height after grazing gave a better indication of the animals' reaction to sward conditions and the management imposed than actual herbage allowance. The quantity per unit area and the composition of material present were important factors influencing intake. Calves were unable to compete with their dams to maintain herbage intake at the lower allowances and therefore are likely to benefit from additional feeding or creep grazing when residual sward height falls below 6cm for periods in excess of 1–2 weeks.     

The effect of herbage allowance on intake and performance of ewes suckling twin lambs

The effect of herbage allowance on the herbage intake and performance of ewes and their twin lambs at pasture was investigated. Daily herbage allowances of 40,80.120 and 160 g organic matter (OM) kg^-1 ewe live weight, based on herbage mass measured to ground level, were offered during the first 12 weeks of lactation.
The sheep were grazed rotationally around four paddocks of a perennial ryegrass pasture for 7-day periods and herbage mass, extended tiller length and digestibility of the herbage consumed by the animals were estimated. Herbage intake by the ewes was estimated during weeks two to twelve and live weights were recorded weekly: during the last two weeks of the experiment grazing behaviour of one ewe on each treatment was recorded continuously.
Herbage intakes by the ewes (164, 1–81, 2–42 and 268 ± 0153 kg d^-1) and live weight gains of the lambs (202, 245, 274 and 300 ± 7–3 g d^-1) increased with increasing herbage allowances. Herbage intakes by the ewes and growth rates of their lambs increased up to a herbage allowance which was over five times the amount of herbage eaten by the ewes.
As the animals reduced herbage mass and sward height, biting rates by the ewes during grazing increased by 4 (± 0·08) bites min^-1 cm^-1 and masticating rates decreased. Mastications reached a maximum of 90 (± 3·5) min^-1 at a sward surface height of 9 cm.     

Herbage intake and milk production by grazing dairy cows 1. The effects of variation in herbage mass and daily herbage allowance in a short-term trial

Twenty-four British Friesian cows were allocated between six grazing treatments (two levels of herbage mass x three levels of daily herbage allowance) in a balanced change-over design with four periods of 12 d each at monthly intervals. Herbage OM mass ranged from 3790 to 5770 kg ha^-1 measured to ground level and daily herbage DM allowances were 30, 60 and 90 g per kg animal live weight. Herbage OM intake was lower at high than at low herbage mass (24·6 vs 26·3 g per kg LW), and was 26·9, 26·6 and 22·9 g per kg LW respectively at daily herbage DM allowances of 90, 60 and 30 g per kg LW. Milk yield was not affected by herbage mass but was depressed at the low herbage allowance.     

Herbage intake and milk production by grazing dairy cows 2. The effects of level of winter feeding and daily herbage allowance

Two experiments are described in which two levels of winter feeding and three levels of herbage allowance during the grazing season were imposed upon March/April calving British Friesian dairy cows. The winter treatments resulted in differences in live weight and milk yield at turnout of 35 and 53 kg and 3·4 and 3·2 kg d^-1 for the two trials. Subsequently, when grazed at generous herbage allowances, the cows were able to compensate for much of this difference but when herbage was restricted the milk yield differences were accentuated. Groups of cows from each winter treatment were offered 25, 50 or 75 (Experiment 1) and 30, 50 or 70 (Experiment 2) g herbage DM per kg LW daily during the grazing season. Daily herbage intakes on the three allowances in each trial were 14·1, 13·3, 10·7 and 12·5, 12·1, 11·5 kg OM and milk yields were 16·0, 15·3, 12·5 and 15·2, 14·3, 11·8 kg SCM respectively. Both intake and milk production were depressed once the cows were forced to consume more than 50% of herbage on offer or to graze the sward down to a mean height of less than 8–10 cm. Grazing behaviour observations indicated that under rotational managements the cows did not compensate for restrictions in available herbage by grazing longer. Highest levels of milk production per unit area were observed in both trials when production per cow was depressed by 20–25%.     

Effects of ruminal fill on short-term herbage intake rate and grazing dynamics of beef heifers

The impact of manipulating ruminal fill (RF) on intake rate of herbage and grazing dynamics was measured with three rumen‐cannulated beef heifers grazing Bermudagrass pastures individually. The treatments compared were removal of proportions of rumen contents of 0 (treatment RF0), 0·33 (treatment RF33), 0·66 (treatment RF66) and 1·00 (treatment RF100). Treatments were randomly applied in a 3 × 4 Youden‐square design. The rumens were emptied before and after planned grazing sessions (30 min) to set up the treatments, and to estimate intake rate and bite mass, respectively. Measurements were made of bite rate, bites per feeding station, feeding stations per minute, intake per feeding station, time per feeding station, eating and searching step rates and times. Apparent bite area and area grazed per feeding station were calculated. Ruminal fill affected short‐term intake rate and changed grazing dynamics. As RF increased, step rates, searching times, bite mass, apparent bite area, bites per feeding station, area grazed per feeding station, time per feeding station and intake per feeding station decreased (P < 0·01) while step times, eating step rates and bite depth increased (P < 0·01). The results of the present study indicate that RF is an important factor governing the intake characteristics and behaviour of grazing beef heifers.     

Effect of herbage allowance on the efficiency of defoliation by the grazing animal

An experiment examining the relationship of daily herbage disappearance (DHD) and daily herbage allowance (DHA) is described. Cows with calves were grazed at three stocking rates on Kleingrass and Coastal Bermudagrass pastures. DHD and DHA were monitored at 14-d intervals. A significant positive linear relationship was noted between DHD and DHA ( P 0·25). Equations derived for Coastal Bermudagrass and Kleingrass were not significantly different and resulted in a combined equation Ŷ= 0·275X + 0·209 with an r² vaiue of 0·27. As DHA exceeded 6–9 kg DM per 100 kg live weight, efficiency of defoliation by the cows and calves declined.     

Defoliation pattern, foraging behaviour and diet selection by lactating dairy cows in response to sward height and herbage allowance of a ryegrass-dominated pasture

The effects of sward surface height (SSH) and daily herbage allowance (HA) on the defoliation pattern and grazing mechanics of early lactation dairy cows grazing on irrigated perennial ryegrass–white clover pasture were studied. The hypothesis tested was that SSH and HA affect intake and diet selection through their effects on the pattern of defoliation which is influenced by the resistance to prehension bites. Factorial combinations of two initial SSH (14 and 28 cm) and two daily HA (35 and 70 kg DM cow^?1 d^?1) were examined in a replicated experiment. The peak longitudinal tensile force required to break the sward portion encompassed in a 100 cm² area [bite fracture force (BFF₁₀₀)] was measured as an index of the resistance to prehension. The volume of herbage defoliated and herbage intake increased with SSH (P < 0·05) and HA (P < 0·01). Corresponding to an increase in HA from 35 to 70 kg DM cow^?1 d^?1, there was a proportional increase in the total defoliation area (TDA) and intake by 0·24 and 0·55 in the short sward compared with 0·16 and 0·32 in the tall sward respectively. The results of this experiment suggest that a consistent spatial pattern of reduction of the canopy exists during defoliation by cows and that the volume of sward canopy defoliated is the major variable affecting herbage intake. The BFF increased down the sward profile at a rate that was higher (P < 0·05) for the taller sward than for the shorter sward. It is proposed that a relatively lower resistance to prehension in the short sward compared with the tall sward explains the greater proportionate increase in TDA and intake corresponding to an increase in HA. The rate at which BFF₁₀₀ increases down the sward profile is suggested as a sward physical variable that can influence the defoliation process. The estimated time and energy costs of prehension bites are discussed in the context that larger bites are handled more efficiently than smaller bites.     

Effect of herbage mass and allowance on sward characteristics,milk production,intake and rumen volatile fatty acid concentration

The objective of this study was to examine the effect of herbage mass and daily herbage allowance (DHA) on sward characteristics and animal performance, dry‐matter intake, rumen pH and volatile fatty acid production of unsupplemented spring‐calving dairy cows throughout the main grazing season. Sixty‐eight Holstein‐Friesian dairy cows were randomly assigned across four treatments (n = 17) in a 2 × 2 factorial design. Two swards were created with different levels of pre‐grazing herbage mass [allocated above 4 cm (>4 cm); 1700 kg DM ha^?1 (medium; M) or 2200 kg DM ha^?1 (high; H)] and two levels of DHA (>4 cm; 16 or 20 kg DM per cow d^?1). An additional eight lactating ruminally cannulated Holstein–Friesian dairy cows were randomly assigned to each treatment in a replicated 4 × 4 Latin square design. Sward and animal measurements were collected across four periods each of 1 week duration in April and May (PI) and July and August (PII). Maintaining the medium‐mass sward across the season improved the nutritive value of the sward in the latter part of the grazing season compared with high‐mass swards, thus resulting in increased animal intakes and milk production throughout PII. The higher organic matter digestibility of the medium‐compared with high‐masses during PII indicates that grazing severity and herbage mass in the spring to mid‐summer period will determine sward quality parameters in the late summer period.     

Effect of maize grain and herbage allowance on estimated metabolizable energy intake and animal performance in beef cattle finishing systems

This study investigated the effects of levels of supplementation with maize grain and herbage allowance (HA) on grass herbage and maize intake, animal performance and grazing behaviour in two replicated grazing experiments with Angus beef cattle in Argentina. In Experiment 1, the response to increasing HA (2·5, 5·0 and 7·5 kg DM herbage 100 kg^?1 live weight (LW) d^?1 with and without 0·5 kg DM maize grain 100 kg^?1 LW d^?1) was investigated. In Experiment 2, the responses to level of maize grain offered (0, 0·5 and 1·0 kg DM maize grain 100 kg^?1 LW d^?1) at an HA of 2·5 kg DM herbage 100 kg^?1 LW d^?1 and an HA of 5·0 kg 100 kg^?1 LW d^?1 without maize grain were assessed. In Experiment 2, soyabean meal was added to control the crude protein concentration in the diet. Two methods were used for intake estimations: pre‐ and post‐feeding herbage mass difference, and the use of the n‐alkane and ¹³C technique. The latter predicted most accurately the metabolizable energy requirements calculated from live weights and liveweight gain of beef cattle attained in each treatment in both experiments. Increasing HA significantly increased herbage intake and liveweight gain (P < 0·01), and general quadratic relationships between these variables could be fitted across experiments despite differences in animal and pasture characteristics. Increasing the amount of maize grain offered significantly reduced herbage intake and grazing time, but increased liveweight gain and digestibility of the diet. Substitution rate increased with increasing HA in Experiment 1 but was not affected by level of maize supplementation in Experiment 2. These relationships will aid the development of grazing management models for Argentinean conditions.     

Effect of structure of a tropical pasture on ingestive behaviour, digestibility of diet and daily intake by grazing cattle

The effect of the structure of a tropical pasture, based on Dichanthium spp., on the ingestive behaviour, in vivo digestibility of the diet and herbage intake by eight Creole tethered heifers was studied. Two levels of nitrogen fertilizer (0 and 50 kg ha^?1) were applied to plots after each grazing cycle and there were 28 d between each of the three grazing cycles. Four heifers grazed individual subplots daily on each plot for 14 d in each of the successive grazing cycles. Simultaneous measurements of bite depth, bite mass, biting rate, short‐term intake rate and daily grazing time were made in two 4‐d periods at the end of each 14‐d period. The in vivo organic matter digestibility (OMD) and daily herbage organic matter intake (OMI, expressed on a kg LW^0·75 basis) were also measured at the same times. Relationships among pasture characteristics and ingestive behaviour were similar to those reported in other short‐term studies: pasture height was highly correlated with bite depth, bite mass and biting rate (r = 0·91, r =0·79 and r = ?0·68, respectively, P < 0·001). Pasture variables had lower correlations with grazing time and short‐term intake rate than with bite depth, bite mass and biting rate. Pasture structure was more highly correlated with OMD than OMI: leaf mass and length and also the extended tiller length were highly correlated with OMD (r = 0·77, r = 0·76 and r = 0·72, respectively, P < 0·001) whereas the crude protein concentration of the herbage was correlated with OMI and digestible OMI (r = 0·50 and r = 0·69, respectively, P < 0·001). Ingestive behaviour variables, as well as OMD, were correlated more with pasture characteristics than was OMI.     

The intake of maize silage by self-fed heifers allowed restricted access

The intake of maize silage by self-fed heifers was measured where access to the silage was limited either by reducing the width of the feed face or by restricting the time allowed for feeding. In the first experiment twenty heifers were allowed 0.4 or 0.2 m feed face per head for 7 h daily. In the second experiment fourteen heifers were allowed 0.5 m feed face per head for 5 or 3 h daily. Intake was measured either by estimating faecal output by the chromic oxide technique and then using the in vitro estimate of silage digestibility to calculate individual intakes, or by using the results of eight individually fed heifers to provide a direct relationship between the chromic oxide concentration of the faeces and total intake, and using this to predict the intake of the self-fed heifers from the concentration of chromic oxide in their faeces. The latter method was found to be the most satisfactory because it took into account the effect of level of feeding on the digestibility of maize silage. Restriction of access to the silage by limiting feed face width (Experiment 1) and time (Experiment 2) significantly reduced intakes by 4 and 7% respectively. In both experiments feeding time was significantly reduced by restriction, but this was partially compensated for by a significant increase in rate of eating.     

The effect of adding surface water to herbage on its digestion by ruminants

In two experiments the effects of adding surface water to herbage on its digestion by ruminants were examined. In experiment 1, with sheep, adding water to silage reduced DM intake and the diet DM digestibility. Offering a high DM alternative forage, hay, increased DM digestibility with the wetted silage, but tended to decrease it with the dry silage. More hay was eaten with the wetted than dry silage. In experiment 2, with steers, adding water to fresh herbage reduced DM intake but had no effect on rumen pH or ammonia levels. Adding water to the herbage reduced the rate of DM disappearance from nylon bags placed in the rumen and the proportion of rumen liquor particles of low specific gravity. The proportion of very short and very long particles in the rumen liquor was also reduced by adding water to the herbage.     

The effect of herbage surface water and the provision of supplementary forage on the intake and feeding behaviour of cattle

High rainfall and the corresponding low herbage DM content has been shown to adversely affect cattle intakes and production. This could arise from physical restrictions on intake or digestion rate, or other behavioural limitations such as reduced palatability. A knowledge of the reason for the intake reduction would assist in determining the optimum supplement to feed. Eight steers were offered four diets in a changeover design. The diets were: A, cut herbage; B, cut herbage with added surface water; C, as B but with a low dry matter (DM) forage supplement (silage); and D, as B but with a high DM forage supplement (hay). The addition of surface water to herbage reduced DM intake and feeding time but had no effect on biting rate or rate of intake. Neither hay nor silage was eaten in sufficient quantities to restore DM intake, even though the silage was of similar nutrient composition to the herbage.     

The effects of level of Molinia utilization on diet selection and herbage intake by cattle grazing Molinia grassland

Groups of mature, non-lactating, cows grazed two Molinia -dominant grassland communities in central and southern Scotland during six consecutive summers. Two treatments, designed to use either 33% or 66% of the estimated annual Molinia leaf production by grazing to different leaf lengths, were imposed at each site. Grazing was restricted to the period of Molinia growth each season. During the first 4 years, diet composition, diet digestibility and herbage organic matter intake were determined during either one or two measurement periods each year. There were differences between sites in the floristic content of the sward and these differences were reflected in the diet selected by the cattle. Cows grazing the taller (33% utilization) plots had higher percentages of Molinia , grass stem, sheath and inflorescences and lower percentages of broad-leaved grasses, sedges, rushes and dead herbage in the diet than those grazing the shorter (66% utilization) plots. Differences between the floristic composition of the sward and the diet were explicable by (a) the height at which cattle grazed in relation to the distribution of components within the sward or (b) the selective grazing of small areas dominated by a particular species. The organic matter digestibility of diets differed between sites but there was no significant difference in digestibility or organic matter intake between the treatments. On average less than 50 d grazing was provided by the experimental sites each year. During this period the liveweight gains of cows grazing the two treatments did not differ significantly. The implications of these results for the management of Molinia -dominant communities are discussed.     

The effect of drainage on herbage growth and soil development

The effects of drainage on the yield of perennial ryegrass ( Lolium perenne ) swards grown for silage, and on soil development under grassland, were studied between 1982 and 1985.
Over the 4 years of the experiment, drainage resulted in a mean increase in herbage production of 1-5 t dry matter (DM) ha⁻¹, 16% above the yield recorded on the undrained site. Drainage also led to increased herbage nitrogen contents at every cut except one, so that total nitrogen recovery was increased by an average of 24-8 kg N ha⁻¹ (15%) when drainage was installed. These total differences were mainly due to differences that were present at the first cut taken in June each year; drainage led to an average 24% increase in yield, and an average 27% increase in nitrogen recovery at the first cut.
Detailed sward measurements taken in 1985 endeavoured to explain these differences; drainage increased depth to free water over winter, enabling increased root efficiency and nitrogen availability, greater tillering and more rapid leaf extension in early spring. There was some suggestion that drainage also led to an improved topsoil structure in terms of improved aggregation and aeration. This is important for long-term soil development as well as short-term herbage yield.     

The effects of nitrogen and phosphorus fertilizer application on herbage yield of natural pastures

This study was carried out in a natural pasture in Van province of Turkey between the years of 2004 and 2005 to determine the effects of nitrogen and phosphorus fertilization on herbage yield. The study was performed in randomized block with three replications. 0, 4, 8, 16 kg da(-1) nitrogen and 0, 6, 12 kg da(-1) phosphorus applications were examined. The effects of different nitrogen and phosphorus fertilizer applications on plant height, green herbage and crude protein yield were significant for both years. Depending on the increasing nitrogen and phosphorus applications, significant increases were in green herbage, hay and crude protein yields. According to results, differences between 8 and 16 kg da(-1) doses of nitrogen and 6 and 12 kg da(-1) doses of phosphorus applications were insignificant. In the first year, in terms of nitrogen applications, the highest green herbage, hay and crude protein yield (1423.1, 263.3 and 29.2 kg da(-1)) were obtained from 8 kg N da(-1), in the second year, the highest values (1426.1, 602.7 and 67.8 kg da(-1)) were obtained from 16 kg N da(-1) application. As for the phosphorus applications, in the first year, the highest green herbage yield was 1142.2 kg da(-1) at the 6 kg P2O5/da application, hay and crude protein yields (218.2, 23.1 kg da(-1)) were recorded from 12 kg P2O5/da application. In the second year, the highest green herbage, hay and crude protein yields were (1335.8, 549.6 and 66.1 kg da(-1)) determined at the 12 kg P2O5/da application.     

Influence of defoliation regime on herbage production and characteristics of intake by dairy cows as affected by grazing intensity

Three contrasting defoliation regimes for dairy cows—four cuts annually, severe rotational grazing and lax rotational grazing integrated with cutting—were compared in terms of herbage production, selection and intake per cow. Lax and severe grazing were compared simultaneously and grazing intensity was characterized by means of changes in herbage mass and sward height during grazing.
Herbage growth and yield were similar under cutting, severe grazing and lax grazing/cutting (120 t organic matter (OM) ha⁻¹). Herbage quality and leaf production were highest with severe grazing, which also had a less marked seasonal pattern of growth. The herbage intake per cow was 111 kg OM d⁻¹ with severe grazing and proportionately (0-20) higher at lax grazing/ cutting. 050 of the herbage yield was harvested by cutting at lax grazing/cutting. Selectivity was described with high certainly by the nutrient content prior to defoliation and by the degree of defoliation. Models to quantify this are presented.
Grazing intensity could be described by the size and degree of utilization of the fouled area, and herbage intake was quantified by means of herbage height before and after grazing. Within the range of 80-240 mm sward height prior to grazing, height measurements led to realistic and more accurate estimates than those obtained by measuring organic matter disappearance.
Herbage utilization was depressed significantly by increasing maturity of herbage due to lower nutritive value, but in particular due to reduced green leaf content and increased reproductive development. If sward height did not exceed 250 mm at any time, good utilization by grazing could be obtained.     

Concentrate supplementation of grazing dairy cows. 2. Effect of concentrate composition on herbage intake and milk production

The effect of feeding either traditional concentrates containing starch or high quality fibrous concentrates on the performance of grazing dairy cows was examined in a trial in which cows were given concentrates with either 350 g starch and sugars (kg dry matter (DM))^-1 (high-starch) or 100 g starch and sugars (kg DM)^-1 (high-fibre). The swards used consisted predominantly of perennial ryegrass and were usually aftermaths following cutting. Each area was grazed for 3 or 4 d at each grazing and a two-machine sward-cutting technique was used for estimating herbage intake.
The effect of concentrate composition on the herbage intake of grazing cows at a high daily herbage allowance of 28 kg OM above 4 cm cutting height was investigated in 1983 and 1984. With 54 kg OM d^-1 of high-starch concentrates the mean herbage intake was 11·5 kg OM d^-1 per cow while cows fed 5.3 kg d^-1 of high-fibre concentrates consumed on average 12–6 kg OM d^-1. The mean substitution rate of herbage by concentrates was reduced from 0·45 kg herbage OM (kg concentrate OM)^-1 with the high-starch concentrate to 0·21 with the high-fibre concentrates.
The effect of the treatments on milk production was studied in 1984. The cows consumed 5·5 kg OM d^-1 as concentrates and grazed at a lower herbage allowance of 19 kg OM above 4 cm cutting height. With high-fibre concentrates milk production and 4% fat-corrected milk production were 13 and 1·8 kg d^-1, respectively, higher than with the high-starch treatment. The daily live weight gain with the high-starch concentrates was 0·17 kg per cow more than with the high-fibre concentrates.     

The effect of some pre-treatments on proteolysis during the ensiling of herbage

Two experiments were carried out to examine the nitrogenous changes occurring in herbage during harvesting and ensiling. In the first, ryegrass-clover was wilted rapidly in the laboratory (6 h) and in the field under good (48 h) and poor (48 h and 144 h) weather conditions. Protein breakdown and ammonia formation were negligible in herbage wilted rapidly and it was only when the crop was exposed to a prolonged wilt in humid conditions that appreciable proteolysis occurred. In the second experiment ryegrass-clover was ensiled in laboratory silos after treatment with varying levels of formic and sulphuric acids. An additional treatment included herbage inoculated with a mixture of lactic acid bacteria and glucose. The silos were opened after 4 and 50 d and samples were analysed for protein-N, ammonia-N, water-soluble carbohydrates and organic acids. There was a high negative correlation between level of addition of either acid and the degree of proteolysis and deamination. Even at the highest levels of formic acid (7·7 g per kg) and sulphuric acid (4·0 g per kg) additions, however, about 45% of the original herbage protein was degraded after 50 d in the silo. The inoculum treatment was also effective in reducing proteolysis, the effect after 50 d being comparable with formic acid applied at a rate of 4·1 g per kg.