针叶树材纵向气体渗透的三维流阻网络

气体在木材中的渗透是通过木材内的孔隙通道进行 ,而木材内的孔隙结构非常复杂 ,在纵向、径向、弦向的连接特性均不相同。以往关于针叶树材气体渗透的模型或理论 ,只考虑针叶树材内气体渗透通道在 1个或 2个方向上的连接。本文根据针叶树材管胞流阻和管胞在纵向、径向、弦向的连接特性 ,建立了一个描述针叶树材纵向气体渗透性的三维流阻网络 ,并运用重正化变换求解出流阻网络的流阻 ,计算了针叶树材气体纵向渗透系数。所得马尾松木材气体纵向渗透系数计算值与其测量结果进行比较 ,两者十分吻合 ,表明本文关于针叶树材气体纵向渗透三维流阻网络理论的科学性     

针叶树材径向和弦向气体渗透的三维流阻网络

将针叶树材气体渗透的三维流阻网络理论进行了扩充 ,提出了气体径向和弦向渗透次级流阻网络的二维等效流阻网络及二维等效流阻网络流阻基元的确定和等效流阻的求解方法 ,将三维流阻网络从关于针叶树材气体纵向渗透性研究 ,扩展到针叶树材气体径向和弦向渗透性研究。提出了针叶树材径向和弦向气体渗透三维流阻网络理论 ,计算了马尾松试件径向和弦向渗透系数 ,并与测量值进行比较 ,结果表明两值相符合。将本文的针叶树材径向和弦向气体渗透三维流阻网络理论与此前关于针叶树材纵向气体渗透的三维流阻网络工作相结合 ,可以使用三维流阻网络计算针叶树材纵向、径向、弦向的气体渗透系数 ,其在木材干燥、改性、防腐有应用价值     

针叶树木材流体纵向渗透性与其构造关系的理论分析

本文首次提出搭接面及侧面胞壁上纹孔在流体纵向渗透时的不同作用,并运用流体力学的原理,对针叶树木材流体纵向渗透性与其构造的关系进行了系统分析。结果表明：针叶树木材流体渗透系数等于早、晚材对其贡献之和,早、晚材对其的贡献,分别等于单位面积上流体流动路径数、管胞平均长度与纹孔膜膜缘厚度比值及纹孔膜微孔的比渗透性三者之积。经验证,研究结果与实测值较好的相符。     

针叶树木材细胞力学及纵向弹性模量的计算--试件纵向弹性模量的预测

依据针叶树木材管胞和射线细胞的结构模型。使用计算机抽样模拟解剖结构参数。以及使用针叶树木材纵向弹性模量计算公式和方法,计算人工林杉木,马尾松幼龄材和成熟材试件纵向弹性模量,计算结果与常温条件下气干试件测定结果十分符合。在试件晚材率和管胞解剖结构参数改变的条件下。计算预测了人工林杉木,马尾松幼龄材和成熟材纵向弹性模量的变化。结果表明：试件纵向弹性模量随晚材率,管胞长度,管胞壁厚度的增加而增加,而试件纵向弹性模量随管胞直径增加而减小。本文提出的纵向弹性模量计算的预测方法,对于运用现代生物技术控制和改变针叶树木材的材质,材性有实际意义。     

木材流体渗透性研究的发展与趋势

介绍了木材流体渗透性研究领域中的主要理论和常用的木材结构模型;简述了我国木材流体渗透性研究的发展和现状,以及自60年代以来所取得的主要成果。木材渗透性研究的理论主要包括线性粘滞液体流的Hagen-Poiseuile方程,非线性粘滞液态流的Erk或Bar方程,气体滑流效应的线性气体流Adzumi方程;常用于木材渗透性研究的木材结构模型有:均匀并联毛细管模型、Sebatian针叶材模型、Pety串联流导模型、Comstock针叶材模型,以及Bramhal提出由Bolton改进的渗透截面随长度衰减模型;此外还有关于短毛细管的Couete修正和Clausing修正。     

针叶树木材细胞力学及纵向弹性模量计算——纵向弹性模量的理论模型

本文是关于针叶树木材弹性特性研究的理论分析。设定针叶树木材细胞主要由管胞和射线组成 ,根据管胞和射线细胞的解剖构造特征 ,建立了两端劈尖、矩形截面、中空的管胞模型与长方体状、中空的射线细胞模型。沿用MP层 (胞间层 初生壁 )内与 3S层 (次生壁的S1、S2、S3层 )内力学性质相同 ,而MP与 3S层间力学性质相异的假定 ,利用管胞和射线细胞模型 ,导出了管胞和射线细胞纵向弹性模量的计算公式。同时 ,根据管胞和射线细胞在针叶树木材中的排列规律 ,结合线形弹性体串并联的特性 ,进一步给出了计算针叶树木材试件宏观纵向弹性模量的方法。本项研究从木材细胞的结构和弹性特性出发 ,分析研究针叶树木材的宏观弹性行为 ,一方面可以获得关于针叶树木材弹性特性的认识 ,另一方面可以用于针叶树木材纵向弹性模量的计算和预测。     

木材可压缩流体渗透中滑流的研究

在气体平均压力倒数为1.4-48atm~(-1)范围内测定了长白鱼鳞云杉和红松木材可压缩流休渗透中滑流效应及影响滑流的因素,论述了木材中流体流动形态和机理。研究表明,两种木材可压缩流体渗透中均产生滑流,但长白鱼鳞云杉中滑流高于红松。前者滑流渗透性与真渗透性百分比、比名义渗透性与真渗透性比、滑流因素分别为69.39％、1.69、0.356atm;后者分别为37.64％、1.376、0.197atm。滑流的大小与木材有效毛细管半径、真渗透性和气体平均压力有高度紧密相关。木材有效毛细管半径愈小,真渗透性愈低,或气体平均压力愈低,则滑流效应愈大。     

中国裸子植物木材管胞瘤状层的电镜观察

本文对中国裸子植物11科、41属、99种木材管胞瘤状层进行了系统观察。瘤状层的有无及形态特征是区分裸子植物材的重要标志。此外并对5种针叶树材在一个年轮内,瘤状层从早材到晚材大小、分布密度的变化做了观察;12种针叶树材的切片,用0.5％次氯酸钠溶液处理后,瘤状物减小、以至消失;在另6个树种的管胞内,观察到瘤状层被白腐菌或软腐菌溶解。     

龙竹竹材纵向气体渗透性测定

参照木材流体渗透特性相关理论和研究方法,对大型丛生竹种———龙竹竹材的纵向气体渗透性进行了测定和分析,结果表明预处理方法和含水率对竹材纵向渗透性有显著影响。预处理可提高竹材气体渗透性,水煮预处理试件的渗透性优于汽蒸预处理试件的渗透性;随竹材试件含水率降低,竹材纵向气体渗透性增加。这些结论可为竹材防霉处理、染色处理、防火处理等改性处理提供有益的参考。     

黄花落叶松木材超微结构及其对渗透性的影响

本文详细地介绍了黄花落叶松木材超微结构特征及其对木材渗透性的影响,心材和边材管胞具缘纹孔的超微结构及其在不同部位和不同处理方法中的变化;晚材管胞具缘纹孔超微结构的变化;细胞壁各层次中的微纤丝及其周围的基质和结壳物质等的渗入或沉积性状。同时还针对黄花落叶松木材渗透性差,及一些晚材具缘纹孔少闭塞或不闭塞的原因,从木材的超微结构特点进行了较详细的探讨。     

Comparative study on penetration characteristics of modern wood coatings

Summary The penetration characteristics of five modern wood coatings (three waterborne, one high solid and one solvent borne) into pine sapwood, spruce and dark red meranti have been systematically compared. The degree of coating penetration is mainly determined by the ability of the coating to flow into wood capillaries. Binder type, pigmentation, solid matter content and drying speed appeared to influence this ability. In softwoods the following different coating penetration routes are observed: the flow into open ends of longitudinal early-and latewood tracheids, the flow into ray cells and the transport from rays through the cross-field into longitudinal tracheids adjacent to rays. The possibility for the coating to follow the latter route is strongly influenced by the existing type of cross field pitting and to a lesser degree by the pigmentation of the paint. Clear differences between pine and spruce have been found with respect to the flow into ray parenchym and ray tracheids. The flow into open ends of longitudinal tracheids is strongly influenced by the grain angle of tracheids. Penetration into dark red meranti is mainly limited to vessels and rays. Tylose membranes can prevent the complete filling of vessels. The impact on penetration of the removal of extractives and of sanding of the surface has also been studied but appears to be of only minor importance.The authors want to thank Akzo Nobel Coatings, DSM Resins and Sigma Coatings for their technical support. This research was financed by the the Dutch Innovative Research Program on Coatings under contract number IVE 93-812. Authors also thank the University of Hamburg (BFH) for the use of their electron microscopy facilities.     

Impact of soil pressure and compaction on tracheids in Norway spruce seedlings

We tested the effect of soil compaction on Norway spruce seedlings in terms of the size and theoretical volume flow rate of the tracheids. The results show that soil pressure limits growth in the diameter of the lumens of tracheids in all parts of seedlings studied. The tracheids of the roots with primary xylem had larger lumens than those of the roots and shoots with secondary xylem in both unloaded and loaded seedlings. This corresponds to the higher cumulative theoretical volume flow rate of the tracheids from roots with primary xylem than those from roots and shoots with secondary xylem. Although the volume flow rate of tracheids, according to the Hagen-Poiseuille law, was directly proportional to the quadratic power of the capillary diameter (tracheid lumen), the cumulative curve of the theoretical hydraulic volume flow rate was higher or relatively comparable in loaded seedlings. An explanation for these findings is that there were higher gradients of water potential values in roots and leaves in loaded seedlings because the lengths of the conductive pathways were 27% shorter than in unloaded seedlings. We hypothesise that trees have adapted to different stresses by shortening their conductive pathways to maintain a transpiration rate similar to that of non-stressed trees. These results concerning the impact of soil compaction on tracheid diameter and volume flow rate improve our understanding of the growth and functioning of different conifer organs and the mechanisms underlying the efficiency of water transport through the root xylem to the shoot.     

The potential of SilviScan’s X-ray diffractometry method for the rapid assessment of spiral grain in softwood, evaluated by goniometric measurements

The orientation of the tracheid cells with respect to the stem axis (grain angle, spiral grain) exerts a strong influence on numerous material properties. Therefore, several methods have been developed to assess this wood anatomical feature. The current study compares SilviScan spiral grain measurements, which uses X-ray diffractometry, with direct goniometer readings. The correlation between these two methods yielded an r² of 0.874 and a root mean square error of prediction (RMSEP) of 1.21 clearly demonstrating that X-ray diffractometry can provide accurate and rapid information on the grain orientation in Norway spruce (Picea abies (L.) Karst.) or similar structured softwoods.     

Variation of measured cross-sectional cell dimensions and calculated water vapor permeability across a single growth ring of spruce wood

A statistical study of the cell dimensions in a growth ring of spruce along the radial and tangential directions is performed. The data are used to study the variation of the cell vapor permeability in the growth ring. Studying cell rows within one growth ring, the frequency distributions of the cell wall thickness in the radial direction and of the lumen dimension in the tangential direction are found to be both unimodal. In contrast, the frequency distributions of these dimensions in the other directions are bimodal, where the different modes can be attributed to earlywood and latewood. Analysis of the bimodal distributions results in the determination of threshold values of cell wall thickness and the lumen dimension for earlywood and latewood tracheids. The cell dimensions are used to predict cell porosity and water vapor permeability distribution within a growth ring. The bimodal frequency distributions of the tangential cell wall thickness and the radial lumen dimension provide an explanation for the observed bimodal frequency distribution of the cell water vapor permeability both in radial and in tangential directions. Contrary to measured macroscopic vapor permeability results, the tracheid geometry results in lower cell vapor permeability in radial than in tangential direction. This confirms that rays play an important role in the vapor permeability of wood, as they can be considered as pathways for vapor transport in radial direction. The dataset analyzed in this paper leads to a set of parameters characterizing the earlywood and latewood cell dimensions. Such characterization can be used, for example, for producing synthetic data for computational modeling studies.     

Hydraulic and mechanical properties of young Norway spruce clones related to growth and wood structure

Stem segments of eight five-year-old Norway spruce (Picea abies (L.) Karst.) clones differing in growth characteristics were tested for maximum specific hydraulic conductivity (k(s100)), vulnerability to cavitation and behavior under mechanical stress. The vulnerability of the clones to cavitation was assessed by measuring the applied air pressure required to cause 12 and 50% loss of conductivity (Psi(12), Psi(50)) and the percent loss of conductivity at 4 MPa applied air pressure (PLC(4MPa)). The bending strength and stiffness and the axial compression strength and stiffness of the same stem segments were measured to characterize wood mechanical properties. Growth ring width, wood density, latewood percentage, lumen diameter, cell wall thickness, tracheid length and pit dimensions of earlywood cells, spiral grain and microfibril angles were examined to identify structure-function relationships. High k(s100) was strongly and positively related to spiral grain angle, which corresponded positively to tracheid length and pit dimensions. Spiral grain may reduce flow resistance of the bordered pits of the first earlywood tracheids, which are characterized by rounded tips and an equal distribution of pits along the entire length. Wood density was unrelated to hydraulic vulnerability parameters. Traits associated with higher hydraulic vulnerability were long tracheids, high latewood percentage and thick earlywood cell walls. The positive relationship between earlywood cell wall thickness and vulnerability to cavitation suggest that air seeding through the margo of bordered pits may occur in earlywood. There was a positive phenotypic and genotypic relationship between k(s100) and PLC(4MPa), and both parameters were positively related to tree growth rate. Variability in mechanical properties depended mostly on wood density, but also on the amount of compression wood. Accordingly, hydraulic conductivity and mechanical strength or stiffness showed no tradeoff.     

Simulation of effects of wood microstructure on water transport

A tracheid-level model was used to quantify the effects of differences in wood microstructure between coastal and interior Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco var. menziesii and var. glauca) wood on larger scale properties like hydraulic conductivity. The model showed that tracheid length, the ease of flow through a bordered pit and effective tracheid diameter can all limit maximum hydraulic conductivity. Among the model parameters tested, increasing bordered pit conductivity and tracheid length resulted in the greatest increase in maximum conductivity in both the inland and coastal ecotypes. A sensitivity analysis of the uncertainty between parameters governing flow through the bordered pit and air-seeding potential showed that, although decreased pit flow resistance increased maximum hydraulic conductivity, increased cavitation led to lower conductivity over time. The benefits of increasing the number of bordered pits depended on the intensity of the meteorological driving function: in drier environmental conditions, wood with fewer pits was more conductive over time than wood with more pits. Switching the bordered pit characteristics between coastal and interior wood indicated that the conductivity time course of coastal and interior wood was primarily governed by differences in the number of bordered pits and not differences in tracheid dimensions. The rate at which tracheids refilled had little effect on the conductivity time course of either coastal or interior wood during the first two summers when the wood was highly saturated, but had a marked influence in subsequent years once the cavitation profile stabilized. Our work highlights the need for more empirical work on bordered pits to determine whether variation in their number and properties is related to changing environmental conditions. In addition, a detailed simulation model of a bordered pit is needed to understand how variation in pit properties affects the relationship between ease of flow through a bordered pit and its potential for facilitating air-seeding.     

How timing of stem girdling affects needle xylem structure in Scots pine

While needles represent a proportionally large fraction of whole-plant hydraulic resistance, no studies to date have investigated how source–sink disturbances affect needle xylem structure. In this study, we evaluated structural changes in xylem in current-year needles of Scots pine 227 and 411 days after stem girdling (hereafter referred to as DAG). Maximum and minimum tracheid lumen diameters and therefore also the size of tracheid lumen areas increased in needles 227 DAG compared to control needles. In contrast, tracheid dimensions were similar in needles 411 DAG as in the control needles, but smaller xylem area and lower number of tracheids resulted in the lower theoretical needle hydraulic conductivity of those needles. Several needle xylem parameters were intercorrelated in both control and girdled trees. These observed changes provide a new understanding of the processes that occur following a source–sink disturbance. Considering anatomical parameters such as the number of tracheids, tracheid dimension, or needle xylem area, which are rarely described in physiological studies, could be helpful, for example, in understanding to tree hydraulic systems or for modeling gas exchange. Finally, empirical equations were developed to calculate needle theoretical hydraulic conductivity and the number of tracheids in needles using an easily measurable parameter of needle xylem area.