Contribution of understory vegetation to minimizing nitrate leaching in a Japanese cedar plantation

Understory vegetation may affect nitrate (NO₃ ⁻) leaching, even in coniferous forests. Our objective was to estimate the contribution of understory vegetation to nutrient cycling, especially nitrogen, in a Japanese cedar (Cryptomeria japonica) stand. We therefore cut down and removed understory vegetation in one plot of the stand (the cutting plot) to compare nutrient budgets in the cutting plot with those in a control plot in which understory vegetation was allowed to grow. We also examined neutralization of the acid produced due to an increase in NO₃ ⁻ leaching. A monitoring study on precipitation and soil-percolated water was carried out in both plots. When the understory vegetation was cut down, NO₃ ⁻ flux at a soil depth of 10 cm increased remarkably in summer, with values significantly higher than those in the control plot. This resulted in an increase in proton load associated with N transformation ([H⁺]load). The increase in [H⁺]load enhanced mobilization of Ca²⁺, Mg²⁺, and SiO₂ ([SiO₂]mob). In addition, the correlations between [SiO₂]mob and mobilization of each base cation were distinct in the cutting plot. These results indicated that the acids produced because of N transformation were buffered not only by ion exchange but also by chemical weathering. The contribution of understory vegetation to minimizing NO₃ ⁻ leaching suggested that understory vegetation might reduce the risk of N saturation because of chronic atmospheric N inputs.     

Species richness and biomass of understory vegetation in a Eucalyptus globulus Labill. coppice as affected by slash management

The aim of this study was to assess the effect of different slash management practices on understory biodiversity and biomass in Eucalyptus globulus coppices in Central Portugal. The experiment consisted of four treatments: (a) removal of slash (R), (b) broadcast over the soil (S), (c) as in S but concentrating woody residues between tree rows (W) and (d) incorporation of slash into soil by harrowing (I). Understory vegetation was surveyed during 1–6, 9, and 10 years, the proportion of soil cover by plant species estimated, and diversity and equitability indexes determined. Above ground understory biomass was sampled in years 2–6, 9, and 10. The highest number of species in most years occurred in plots where slash was removed. Differences between treatments in the proportion of plant soil cover were never significant, whereas differences in diversity index were only occasionally significant and apparently related to the number of species. Thus, differences in the equitability index were not significant. Understory biomass did not decrease during the rotation period, and was usually highest in R and I, and lowest in S, but not significantly different. At the end of the rotation period, understory biodiversity indices and biomass were apparently independent of slash treatment.     

Elevation patterns of woody taxa richness in the evergreen Afromontane vegetation of Ethiopia

Plant species distributions show patterns along elevation gradients. Regardless of the diverse Afromontane vegetation in Ethiopia(AFE), studies of elevation patterns of woody plants are limited and they are restricted to small areas or single/few Mountains. Moreover, there is no general consensus on the patterns of woody taxa distribution by elevation. The objectives of this study were to examine the elevation patterns of woody taxa richness and their relationship with elevation in the AFE. Data were collected and compiled from the Flora of Ethiopia and Eritrea(Vols. 1–7). About 5918 plant species of 243 families were examined and their elevational distributions were recorded. The distributional ranges of woody taxa were aggregated at 19 points(sites) at 100 m intervals starting from 1500 to 3400 m a.s.l. Single-factor analysis of variance(one-way ANOVA) was used to test the hypothesis that mean species richness of woody taxa decreases from the lower to the upper limit of AFE. Simple linear correlation and regression were used to show the relationships of woody taxa richness with elevation. We documented the presence of 505 woody taxa(441 species,31 subspecies and 33 varieties) representing 267 genera and 90 families in the AFE. In terms of habit, 279 taxa are shrubs, 178 are trees and 48 are lianas. The distribution of woody taxa(trees, shrubs and lianas) showed a slight increase at the lower portion and a monotonic decline with increasing elevation. Species richness of trees, shrubs and lianas was negatively and significantly correlated with elevation(r =-0.985,-0.984,-0.981, respectively; all p 0.001). Our hypothesis was accepted because mean richness significantly decreased from the lower to the upper limit of AFE(p 0.001). Generally, monotonic patterns of decline in richness were observed for trees, shrubs and lianas. The contribution of shrubs to total richness increased with increasing elevation whereas that of trees and lianas decreased.     

侧柏人工林下层木本植物的组成及多样性

为明确徐州山地现有侧柏人工林下层木本植物的自然更新状况,依据40个20m×20m样地群落的调查数据,分析了林下自然更新的木本植物的组成及其相似性。结果表明,侧柏人工林下层自然更新的木本植物中单科种、单属种优势明显,林下木本植物区系组成复杂,出现的12种区系地理成分中,温带成分占主体,热带成分占较大比例。侧柏人工林下层出现的木本植物种类较多,但不同生境下群落的物种丰富度和多样性不高、组成树种的相似性极高,尤其是乔木树种的相似性明显高于灌木树种。林下木本植物的种类组成受立地条件和种子传播方式的显著影响,建议改善立地条件以提高林下木本植物的丰富度和多样性,同时控制外来树种的侵入以促进人工林的地带性演替。     

Effects of root zone trenching on soil nitrogen dynamics in Japanese cedar and cypress plantations

The effects of root exclusion and planted tree species on soil nitrogen (N) dynamics were examined at two plantations, one planted with Japanese cedar and the other with Japanese cypress. We set up ten 1 × 1 × 0.2-m-deep trenched sites and ten untrenched control sites at each plantation. We measured the pool size and leaching of inorganic N at each site for 2 years and the net N mineralization 1 and 2 years after trenching. Despite similar soil conditions, the cedar plantation showed higher net N mineralization than the cypress plantation. Stopped tree uptake of N was expected to cause an increased pool size and leaching of inorganic N at the trenched sites. Nevertheless, we found no significant increase in those variables at both plantations. The trenched cypress sites showed no decrease in the net N mineralization during the 2 years after trenching. However, the net nitrification at the trenched cypress sites increased remarkably at the deeper horizons in comparison with that at the control sites. Enhanced nitrification might result from improved ammonium availability through root exclusion. Net N mineralization at the trenched cedar sites decreased more than 60% compared with that at the control sites 2 years after trenching. Higher nitrification potential at the cedar plantation and enhanced nitrification potential at the trenched cypress sites never resulted in increased leaching of N, due to added fine root litter which acted as an immobilization agent for excess N, thus preventing N loss.     

Line thinning promotes stand growth and understory diversity in Japanese cedar (<Emphasis Type="Italic">Cryptomeria japonica</Emphasis> D. Don) plantations

We studied the effects of line thinning on stand structure, microclimate and understory species diversity of two Cryptomeria japonica D. Don plantations in south-central Japan. In each of two study sites we compared stand structure between the thinned stand and an adjacent unthinned stand and found that line thinning increased the growth rate of residual trees such that stand basal area may recover within 10 years after thinning. In the thinned stand, more open canopy conditions resulted in higher maximum temperatures on the forest floor during the early growing season than in the unthinned stand. The thinned stand had greater understory plant species richness and biomass than the unthinned stand. This study suggested that line thinning could potentially enhance biodiversity while simultaneously increasing tree-growth rates in overstocked Cryptomeria japonica plantations.     

Factors affecting the occurrence of woody plants in understory of sugi (Cryptomeria japonica D. Don) plantations in a warm-temperate region in Japan

We compared the occurrence of woody plants present in sugi (Cryptomeria japonica D. Don) plantations and evergreen broad-leaved natural forests in a warm-temperate region in order to characterize the understory vegetation of sugi plantations. The influence of stand attributes on species occurrence in the plantations was evaluated using logistic regression analysis. Out of 163 species, 65 plantation-preferring species and 31 natural forest-preferring species were detected. The plantation-preferring species group contained deciduous species (38 of 65 species) that are characteristic of coppice woodland, but all the natural forest-preferring species were evergreen plants. This indicates that the composition of the woody understory in sugi plantations of the region is characterized by the species of open, disturbed habitats. Stand age and the distance from the nearest natural forests were found to significantly affect the occurrence of evergreen shrubs and plants with gravity-dispersed seeds, respectively, but the light environment of the stand had less effect. According to the regression model analyses, long-term strategies such as lengthening the rotation (logging age) or creating patch mosaics of plantations in the matrix of natural forests appeared to be effective in conserving woodland species that are characteristic of evergreen, broad-leaved forests. On the other hand, increasing light penetration through thinning practices is a short-term strategy that is less likely to be effective in promoting these species.     

Species richness and guild composition in rubber plantations compared to secondary forest on Hainan Island, China

Currently, the one of great threats to tropical biodiversity is the conversion of natural ecosystems to agriculture, and this threat is particularly critical on Hainan, the largest tropical island in China and a global biodiversity hotspot. Rubber (Hevea brasiliensis) has been planted on Hainan since the 1950s, resulting in extensive replacement of native forest, and the rate of this transformation has increased with latex prices since the 1990s. Rubber plantations now cover ca. 506,680 ha on Hainan and reach ca 1,000 m in elevation. To investigate the effect of this land-use change, we compared avian communities between a rubber plantation and a native secondary semi-deciduous monsoon forest. We found that species richness was higher in the native forest than in rubber, and that community composition differed greatly between the two habitats. No strict frugivores were recorded in rubber and no granivores in semi-deciduous monsoon forest. In both richness and abundance, more nectarivores and fewer insectivores occurred in rubber than in the native forest. Some common forest species, as well as protected species, were found only in native forest, including Hainan Partridge (Arborophila ardens), Red Junglefowl (Gallus gallus), Emerald Dove (Chalcophaps indica), Black-browed Barbet (Megalaima oorti), Blue-rumped Pitta (Pitta soror), Puff-throated Bulbul (Alophoixus pallidus), and Chestnut Bulbul (Hemixos castanonotus). Although ten babbler species were found in the native forest, only two were in rubber. Among the species missing in rubber was the endemic Hainan species Grey Laughingthrush (Garrulax maesi). Its endangerment through habitat conversion is of particular conservation concern.     

Nutrition management of cedar and hemlock plantations in coastal British Columbia

Recent re-measurements of silvicultural trials in conifer plantations on nutrient-poor cedar-hemlock (CH) cutovers on northern Vancouver Island have confirmed co-limitation by nitrogen and phosphorus. Repeated fertilization increased volumes of both cedar and hemlock on CH sites (at 2,500 stems ha^?1) by about 100 m³ ha^?1 relative to unfertilized plots 22 years following initial fertilization, and increased the productivity of regenerating conifers to a level approximating that of neighbouring hemlock-amabilis fir (HA) sites. More surprising was the response to fertilization on the more-productive HA sites. After 22 years, cedar in fertilized HA plots had produced an extra 180 m³ ha^?1 compared to unfertilized HA plots, while hemlock had produced an extra 250 m³ ha^?1 in fertilized plots (at 2,500 stems ha^?1). Thus, contrary to expectations, the greatest volume responses of both hemlock and cedar to fertilization occurred on the good (HA) sites rather than on the poor (CH) sites. Ecological studies of CH and HA sites supported the hypothesis that the poor nutrient supply and productivity of CH sites is a long-term consequence of excessive moisture, and that the two site types bracket a critical ecological threshold of moisture, aeration and redox.     

相思人工林生物量分配格局与林下植被多样性分析

对位于漳州平和7年生的3种相思人工林生物量的分配格局和林下植被多样性进行研究,结果表明:马占相思林下植被Shannon-Wiener多样性指数最高(3.0936),黑木相思其次(3.0649),卷荚相思最低(2.2430);而与物种数目无关的Shannon均匀度指数则以黑木相思最高,表明其物种组成比较均匀;3种相思人工林林下植被群落相似性指数均在0.5~0.75之间,表现为中等相似;在3种相思人工林单株各器官生物量组成中,干材生物量所占的比例最大(55.30%~68.28%),其它组分依次为:根(14.72%~19.30%)>皮(10.59%~10.82%)>枝(3.96%~9.19%)>叶(2.42%~8.39%);3种相思人工林林下植被生物量大小顺序为:卷荚相思(6.69 t.hm-2)>马占相思(6.24 t.hm-2)>黑木相思(4.21 t.hm-2);3种相思人工林林分生物量的大小顺序为卷荚相思(75.21 t.hm-2)>黑木相思(49.19 t.hm-2)>马占相思(48.22 t.hm-2)。对3种相思人工林生物量分布格局的比较研究表明:卷荚相思生产力高,适宜在闽南山地营林生产实践中推广。     

木兰围场林管局华北落叶松人工林林下植物多样性研究

对河北省木兰围场林管局北沟林场的华北落叶松人工林林下植被物种组成和群落特征进行了调查和研究,结果表明,不同年龄华北落叶松人工林中,35年生华北落叶松人工林林下植物丰富度较高,有利于林下植被发育和群落结构改善;灌木层的多花胡枝子和土庄绣线菊,草本层的繁缕、糙苏、升麻和歪头菜在不同的群落演替阶段占据重要地位;灌木层Simpson指数和不均匀度指数在林龄梯度上的变化趋势基本一致,均为23年生林分稍大。     

Effects of site preparation and slash management on growth and understory vegetation of Eucalyptus globulus plantations along a rotation time span in Portugal

The removal or maintenance of harvest residues in eucalypt plantations may influence site quality and productivity. Removal of slash from the site may facilitate further management operations and provide a valuable energy resource, but effects on site productivity and sustainability for a rotation time span were not yet assessed under Mediterranean conditions. Therefore, a study was carried out to assess the effects of slash (harvest residues plus forest floor litter) management and soil preparation options on stand productivity and understory vegetation dynamics, hypothesizing that those options influence tree growth, forest floor dynamics and understory biomass and diversity. An experiment was installed in West Central Portugal, consisting on: removal of slash without soil preparation (R); broadcast of harvest residues on the soil surface without soil disturbance (S); as in S, but concentrating the woody debris between tree rows (W); incorporation of slash into the soil by harrowing (I); removal of slash followed by harrowing (RH); and as in I followed by ripping (IS). The experiment was monitored for a rotation time span (140 months). Maintenance of slash followed by deep soil disturbance led to the highest wood production, but differences between treatments were not significant (p > 0.05). Forest floor load and understory biomass were also similar between treatments. Ground vegetation played an important role on nutrient cycling in early rotation stages, such effect being irrespective on slash management options. Incorporation of slash into the soil followed by ripping is probably the best option to match production and environmental sustainability of eucalypt plantations in Mediterranean conditions.     

Line thinning fosters the abundance and diversity of understory Hymenoptera (Insecta) in Japanese cedar (<Emphasis Type="Italic">Cryptomeria japonica</Emphasis> D. Don) plantations

We investigated the effects of line thinning on the abundance of Hymenoptera in two Japanese cedar plantations in northern Hyogo Prefecture, central Japan, in order to evaluate the effectiveness of this silvicultural treatment for ecosystem management. Line thinning is a silvicultural treatment practiced in Japan, where linear stand sections are cut (25%–35% of the total number of trees) retaining one to two rows of trees. We used Malaise traps to capture Hymenoptera from the line-thinned stand (treatment plot, including thinned and retained sections) and the unthinned stand (control plot). Overall, the total number of Hymenoptera was greater in the treatment plot than in the control plot. The treatment plot hosted more functional groups and families of Hymenoptera than the control plot in both plantations. In the Kuchiotani plantation (high-elevation site), the overall abundance of Hymenoptera and of many Hymenopteran functional groups were positively correlated with both species richness and biomass of understory plants. In the Sugi plantation (low-elevation site), understory vegetation was less developed and there was no correlation with abundance of Hymenoptera. Our study indicates that line thinning is an effective silvicultural treatment for future management of overstocked Japanese cedar plantations because it can fulfill biodiversity objectives, but the effectiveness may depend on silvicultural and landscape factors.     

Diversity of naturally-regenerated native woody species in forest plantations in the Ethiopian highlands

Woody species diversity and ground layervegetation cover were studied in plantations ofPinus patula, Cupressus lusitanica,Grevillea robusta, and Juniperusprocera, and in surrounding natural forests inWondo Genet, Ethiopia. In the understory of theplantations, a total of 53 naturallyregenerated tree and shrub species belonging to31 families were recorded. Important indigenoustimber species were also represented. Treesaccounted for 72% of all naturally-regeneratedwoody plant species. On the average, 78% ofthe woody species found in plantations werealso found in the natural forest. There was nosignificant variation in understory woodyspecies richness among plantations. C.lusitanica plantations had a significantlyhigher abundance of woody species than those ofP. patula, but the difference wasnot significant in relation to J. procera,G. robusta and the adjacent natural forest.The herbaceous ground cover percentage in G. robusta and P. patula stands wasconsiderably higher than in C. lusitanicaand J. procera stands. Relative toplantations of other species, plantations ofthe native tree species J. procera hadhigher understory species richness, mediumwoody species abundance and low ground cover.Although there was a marked variation inoverstory plantation density, its effect onunderstory species richness and abundance wasnot significant. The density ofnaturally-regenerated woody species inplantations was over three times the usualplanting density in Ethiopia, indicating a highpotential of forest plantations for restoringthe natural forest ecosystems on degraded landsat a comparatively low cost.     

Simultaneous prediction of the development of loblolly pine and woody competitors in young plantations

A system for examining the effects of hardwood density and cover of herbaceous components on mean size in newly established loblolly pine plantations was developed using a site preparation study located in the piedmont province of Georgia (USA). Multispecies density models were derived to predict the sum of crown heights for eight important hardwood species using both the current or age 1 number of rootstocks and herbaceous vegetation to account for intra- and interspecific effects. The predicted sums of crown heights for the woody species were then used as predictors of mean height, diameter, and volume for loblolly pine. A seemingly unrelated regression procedure was used to compensate for correlations in error components within each system of equations that result from using predicted crown heights as regressor variables. The effects of varying sweetgum density and andropogon cover on the inter-related components of the systems based on current and age 1 competition were examined for mean height and mean diameter of the planted loblolly pines. The cumulative impacts of associated vegetation on loblolly pine growth and the relative sensitivity of height and diameter to competing vegetation were demonstrated.     

Pyrolysis reactions of Japanese cedar and Japanese beech woods in a closed ampoule reactor

The chemical structures of hemicellulose and lignin are different for two distinct types of wood, i.e., softwood and hardwood. Such differences are expected to affect pyrolysis behavior. In this article, the differences are discussed for Japanese cedar wood (a softwood) and Japanese beech wood (a hardwood) pyrolyzed in a closed ampoule reactor (N₂/600°C/40–600 s). Oven-dried samples were used to eliminate the influence of initial water. Demineralized samples (prepared by acid washing) were also used to determine the influence of the minerals contained in the wood samples. As a result, some features were disclosed for secondary char (coke) formation, char reactivity, tar formation, and subsequent decomposition.     

华北落叶松林下植被水平结构研究

采取种群空间分布格局进行研究与分析,运用扩散系数,丛生指数,扩散指数,聚块性指数,Cassie R．M．指数等聚集度指标对塞罕坝大唤起林场华北落叶松林下植被的水平空间结构进行分析研究,发现多数草本在华北落叶松林下呈聚集分布,极少种类如景天三七、柳兰、柳叶沙参和狭叶荨麻等呈随机分布。结果表明,该地区华北落叶松林生物群落稳定,林下草本生物多样性高。     

Modeling production and decay of coarse woody debris in loblolly pine plantations

Forest management can have large impacts on the production and yield of coarse woody debris (CWD) in terrestrial ecosystems, yet few modeling tools exist to inform such efforts. The goal here was to develop a set of prediction equations for use in conjunction with loblolly pine (Pinus taeda L.) modeling and inventory systems to estimate CWD yields at scales ranging from individual trees to whole plantations. Permanent field plots from a 21-year study of thinning effects on plantation growth and yield across the commercial range of the species in the southern United States were surveyed to obtain sample data on CWD volume, density, and mass. Measured CWD properties were combined with inventory records of tree mortality over the study duration to characterize CWD production, decay and yield in a series of prediction equations. The resulting equations predict CWD attributes of dead trees including dry weight (kg) and fraction of standing versus downed woody material based on the time since death (years), tree diameter at breast height (cm) and height (m) at time of death and geographic coordinates of latitude and longitude. A stand-level equation predicts total CWD yield (Mg ha⁻¹) for thinned or unthinned stands based on plantation age, stem density (trees ha⁻¹), and the average height of dominant and codominant trees (m). Piece-level equations predict dry density (kg m⁻³) or nitrogen concentration (%) of CWD pieces based on their position (standing or down), ordinal decay classes, and latitude. The tree and stand-level prediction equations are designed for use in GIS or growth and yield modeling systems. The piece-level equations are designed to be used in inventory applications that survey CWD. The equations should facilitate the accurate and facile determination of mass, carbon, and nitrogen contents of CWD in planted loblolly pine forests of the southern United States.