Using sap flow gauges to quantify water uptake by tree roots from beneath the crop rooting zone in agroforestry systems

Grevillea (Grevillea robusta A. Cunn.; Proteaceae) is used in agroforestry in many areas of the highlands of East and Central Africa, and is reported to be mainly deep rooted, with few shallow roots and correspondingly low levels of competition with associated crops for water and nutrients. To examine the extent of below-ground complementarily in water use between grevillea and cowpea (Vigna unguiculata L.; Leguminosae), experiments were carried out at the International Centre for Research in Agroforestry (ICRAF) Field Centre at Machakos, Kenya. Sap flux was measured using heat balance gauges attached to the stems of young grevillea trees (10–18 months old), both before and after excavating the crop rooting zone (upper 60 cm of soil) around the stem base, in order to establish the capacity of the grevillea to extract water from below this zone. After excavation, the trees maintained sap fluxes of up to 85% of the unexcavated values, suggesting a high degree of below-ground complementarity.     

Effects of mulching with multipurpose-tree prunings on soil and water run-off under semi-arid conditions in Kenya

The effect of adding leaf mulches ofGrevillea robusta, Cassia siamea andGliricidia sepium on the rate of soil and water run-off from a crop field were studied during two cropping seasons in an alfisol under semi-arid conditions at Machakos, Kenya. The experiment was conducted in a randomized complete block design with three replications; two rates of mulch of each species (2.24 t and 4.48 t, on dry matter basis, per ha) and a no-mulch control constituted the seven treatments. Soil and water run-off losses after each major rainfall event and the changes in ground and crop cover were measured. Rainfall erosivity and changes in soil bulk density and infiltration rate were also determined.Soil losses from the plots with mulches ofCassia siamea, Gliricidia sepium andGrevillea robusta were significantly lower than those from the control (p=0.10). Over the two seasons, the cumulative soil losses from plots mulched with cassia, gliricidia and grevillea were 11%, 57% and 81% lower than that of the control plot. Similarly, water run-off losses from cassia, gliricidia and grevillea mulch plots were 28%, 48% and 58%, respectively, lower than that of the control plot. Thus, cassia was found to be better than gliricidia and grevillea in reducing both soil and water run-off losses.Soil bulk density did not change while infiltration rate at the end of the experiment was significantly more than in the beginning; however, there were no significant differences in these soil physical properties among the treatments.     

Coffee shade with Mimosa scabrella Benth. for frost protection in southern Brazil

Damage by radiative frosts is a major limiting factor for coffee cultivation in southern Brazil (south of 20° S latitude). The use of Mimosa scabrella (bracatinga) as a shade tree, to modify the local energy balance and thus prevent damage to the coffee plants, has been evaluated from 1986 to 1994. The study was carried out near Londrina, Parana State (23°23' S, 51°11′ W). During the experimental period, several radiative frosts with intensity ranging from moderate to very severe occurred at the site. Minimum coffee leaf temperatures during these events were 2 to 4 °C higher in the shaded plots. Due to frost protection, coffee bean yields on the average of 7 harvests were higher on the shaded plots. The potential of this system for frost protection in southern Brazil is discussed.     

Spice crops agroforestry systems in the East Usambara Mountains,Tanzania: growth analysis

A scarcity of cultivation land calls for more intensive and productive land use in the East Usambara Mountains in NE Tanzania. Spice crops could generate cash in higher parts of the mountains, but the present cultivation methods are depleting the valuable forest resources. The trial was established at the end of 2000 to find out how the two popular cash crops, cardamom (Elettaria cardamomum (L.) Maton.) and black pepper (Piper nigrum L.), normally grown under the natural forest, will produce in intensive agroforestry system with two multipurpose farm trees, Grevillea robusta A.Cunn. and nitrogen fixing Gliricidia sepium Jacq. Results from 6 years showed that cardamom produced better with grevillea than in natural forest; 5.5 times more in the fourth year than the average in the area. The Land Equivalent Ratios for black pepper and cardamom showed that pepper intercropped with grevillea produced 3.9 times more than in monoculture whereas cardamom intercropped with grevillea and pepper produced 2.3 times more than in monoculture. Gliricidia improved the nitrogen and organic matter content of the soil over the levels found in natural forest. Soil acidity was, however, preventing the plants from using the available mineral nutrients more effectively.     

Allometric estimation of above-ground biomass and leaf area in managed Grevillea robusta agroforestry systems

Non-destructive methods for determining the biomass and leaf area of individual trees throughout their growing cycle are an essential tool in agroforestry research, but must be capable of providing reliable estimates despite the influence that management strategies such as pruning may have on tree form. In the present study, allometric methods involving measurements of the diameter of all branches provided reliable estimates of canopy leaf area and biomass for grevillea trees (Grevillea robusta A. Cunn.; Proteaceae) grown as poles, but proved unsuitable for routine measurements because of their time-consuming nature. An alternative, less laborious method based on measurements of trunk cross-sectional area immediately below the first branch of the canopy provided satisfactory allometric estimates of leaf area and canopy biomass. Trunk biomass was determined from measurements of tree height and diameter at breast height using established methodology based on the assumption that trunk volume may be calculated using a quadratic paraboloid model; biomass was determined as the product of trunk volume and the specific gravity of the wood. The theoretical basis, development and validation of allometric methods for estimating tree growth are discussed and their wider applicability to other agroforestry systems is assessed. This revised version was published online in June 2006 with corrections to the Cover Date.     

Mechanical parameters during static bending of <Emphasis Type="Italic">Pinus radiata</Emphasis> growing in a silvopastoral system I: elasticity and strength

 The effects of pasture fertilization frequency and two vertical positions in the stem on elasticity and strength parameters during static bending (modulus of elasticity, stress at proportional limit, modulus of rupture) of Pinus radiata wood growing in a silvopastoral system were evaluated. Twenty-seven trees were selected randomly from three silvopastoral trials established at Tanumé Experimental Center (34°9′–34°15′ S; 72°53′–72°59′ W). The results indicated that pasture fertilization frequency had no significant effect on the physical and mechanical parameters evaluated. However, the vertical position in the stem did have a significant effect on stress at the proportional limit and on the modulus of rupture due to different average values for the annual ring width and nominal density found in the specimens obtained from logs at two different heights of the stem. Received: February 20, 2002 / Accepted: June 5, 2002 Acknowledgments The authors thank the following institutions and people for their support: CONAF, VI Region; Dr. Rolando Rodríguez, CONAF VIII Region; Cesar Cabrera, Forestry Engineer, CONAF VI Region; Osvaldo Herrera, Director of Experimental Center “Tanumé”; Dr. Manuel Sánchez, Faculty of Forestry Sciences, University of Concepción; Dr. Glenn Hofmann, Faculty of Physics and Mathematics, University of Concepción; Alex Opazo, M.Sc.(C), Faculty of Forestry Sciences, University of Concepción; Staff of Experimental Center “Tanumé”. Correspondence to:M.H.R. Vidal     

Allometric models for estimating aboveground biomass of shade trees and coffee bushes grown together

Allometric models for dominant shade tree species and coffee plants (Coffea arabica) were developed for coffee agroforestry systems in Matagalpa, Nicaragua. The studied shade tree species were Cordia alliodora, Juglans olanchana, Inga tonduzzi and I. punctata. The models predict aboveground biomass based on diameter at breast height (for trees), and the stem diameter at a height of 15 cm and plant height (for coffee plants). In addition, the specific gravity of the studied species was determined.The total aboveground biomass of the shade trees varied between 3.5 and 386 kg per tree, and between 0.005 and 2.8 kg per plant for coffee. The aboveground biomass components (foliage, branch, and stem) are closely related with diameter at breast height (r > 0.75). The best-fit models for aboveground biomass of the shade trees were logarithmic, with adjusted R ² between 0.71 and 0.97. In coffee plants, a high correlation was found (r = 0.84) with the stem diameter at 15 cm height, and the best-fit model was logarithmic, as well. The mean specific gravity was 0.52 (± 0.11) for trees and 0.82 (± 0.06) for coffee plants.     

Growth and yield of sorghum or cowpea in an agrisilviculture system in semiarid India

This study was conducted near Hyderabad, India during 1991–1994 to quantify the effects of shoot pruning, fertilization, and root barriers around Leucaena leucocephala trees on intercropped sorghum(Sorghum bicolor) or cowpea (Vigna unguiculata) crop production under rainfed conditions. Crop plants grown with pruned trees attained higher dry matter and leaf area index than did those with unpruned trees. Two-year mean grain yields of sorghum with no root barriers were76% and 39% of pure crop yield (1553 kg ha–1)for pruned and unpruned trees, respectively. Corresponding values for cowpea were 49% and 26% of pure crop yield (1075 kgha–1). Sorghum or cowpea intercropped with trees responded to fertilizer application more strongly than did their respective pure crops, suggesting an increased need for fertilizer application in this agrisilviculture system over that currently used for pure crops. Impact of root barriers was small on either crop. Irrespective of root barriers, a high response to tree pruning suggested above ground competition for light dominated tree/crop interactions in this agrisilviculture system. This revised version was published online in June 2006 with corrections to the Cover Date.     

Establishment of planted Norway spruce and Scots pine seedlings: effects of light environment,fertilisation, and orientation and distance with respect to shelter trees

The effects of stand stem density (SSD), orientation and distance with respect to shelter trees, and fertilisation on planted Pinus sylvestris and Picea abies seedlings were examined at two sites, one with a southerly aspect (64°14′N, 19°46′E, 225 m a.s.l.) and one with a northerly aspect (64°09′N, 19°36′E, 274 m a.s.l.) in boreal forests in Sweden. The damage and growth were compared at these sites in stands with three SSD’s and different light regimes: uncut forest (SSD 500, ~500 stems/ha), shelterwood (SSD 150,~150 stems/ha), and clear-cut (SSD, 0 stems). Half of the seedlings were irrigated with fertiliser (10 mM N). Fertilisation and SSD affected the height growth of P. abies, since fertilised seedlings in SSD 0 grew the most (22.2 cm). Fertilised seedlings in SSD 0 also generally had the greatest biomass (twigs, leading shoot and needles), even P. sylvestris seedlings on the North slope, where fertilised and unfertilised pine seedlings in SSD 0 had 62.6 and 39.7 g biomass, respectively. P. sylvestris increased about 150% in dry mass of leading shoot of the increase of P. abies with equal height growth, when comparing SSD 150 and SSD 0. The results indicate that the general conditions of the stand have stronger effects on the growth of planted seedlings than their orientation and distance with respect to the nearest tree and that light requirements cannot be moderated by fertilisation. We postulate that a system in which forest gaps are formed under a shelterwood could provide a way to regenerate forests that exploits the beneficial features of both shelter trees and clear-cuts.     

Fruits from shade trees in coffee: how important are they?

Agroforestry systems often receive attention and support in the literature for what is perceived as the benefits from multiple products associated with the trees that create the “forest” component of the setting. A comparison of small coffee growers’ use of fruits derived from the coffee agroforestry holding in Guatemala and Peru reveals that significant differences exist between these groups—not merely in the importance of the fruits themselves, but in the ways they are used. The overall importance of fruits from the coffee system accounts for a relatively small portion of the total value coming from the coffee area (about 10%), but the consumption and sales of the various products do generate needed income or sustenance for most farmers. The fate of fruits shows significant differences between the two countries. Whether at the farm level or on a per hectare basis, Guatemalan coffee farmers are more linked into a market economy and sell significantly more fruits than Peruvian farmers. The opposite is the case when on-farm consumption (use value) of the fruits is compared. While the potential value of these products may be quite large (from 95 to95 to 270/ha), we find that little gets consumed or sold, resulting in tremendous loss of potential benefits that could flow from these sources. Both groups lose more fruits than are sold or used, with Guatemalans foregoing more than three times the dollar value per hectare than Peruvians (151/ha vs.151/ha vs. 44/ha). Data about the economic context within which these growers and the fruits from coffee are found reveal possible reasons as to why we see the differences in use and exchange values realized in the two countries.     

Transpiration of Arabica Coffee and Associated Shade Tree Species in Sub-optimal, Low-altitude Conditions of Costa Rica

Sap flows of coffee (Coffea arabica L. cv ‘Costa Rica 95’) and associated timber trees (Eucalyptus deglupta or Terminalia ivorensis) or leguminous tree (Erythrina poeppigiana) were measured simultaneously during 12 months in 4-year-old coffee agroforestry systems in sub-optimal ecological conditions of Costa Rica. In the wet period, coffee and shade tree transpiration followed the daily patterns of photosynthetic photon flux density (PPFD) and reference evapotranspiration (ETo) while their transpiration was restricted at higher air VPD values (>1.5 kPa) registered during the dry period. Coffee transpired more per unit leaf area in full sun than under shade, an indication of higher environmental coffee stress in non shaded conditions. Nonetheless, coffee daily water consumption per hectare was generally higher under shade than in full sun due higher vegetative growth of shade-grown coffee plants. Minimum and maximum daily transpiration were 0.74 and 4.08 mm for coffee, 0.35 and 1.06 mm for E. deglupta, 0.70 and 2.10 mm for T. ivorensis and 0.13 and 0.79 mm for E. poeppigiana. Estimation of the annual combined water transpiration by coffee and shade trees was 20–250% higher than that of coffee grown in full sun. Nevertheless, there was no evidence that water use by associated trees decreased soil water availability for coffee and hence limited coffee transpiration in the dry season due to its relatively short length (3 months) and the high annual rainfall (over 3100 mm). In the sub-optimal, low altitude conditions of this experiment, E. deglupta was the optimum shade species as it maintained a more constant shade level throughout the year and ensured a better protection to coffee underneath than T. ivorensis and E. poeppigiana which underwent a complete defoliation during the adverse meteorological conditions of the dry period.     

STUDY ON GEOGRAPHIC VARIATION OF DAHURIAN LARCH IN CHINA

Based on the main characteristies of growth,ohenology and resistance at both seed-ling stage (one year old) and sampling stage (seven years old) of Dahurian larch (Larix gmelinii(Rupr.)Rupr.) from seventeen provenances in China ranging from 47°10′ to 52°25′N and 119° 57′to 130°25′E,the rules and patterns of infraspecific geolgraphic variation have been studied by　using variance analysis,corelation analysis,partial correlation analysis and regression analysis.The best provenance for local site and its neighborhood was selected according to the growth char-acteristies of seven years provenance test on Liangshui Experimental Forest Farm of NortheastForestry University.     

Timber harvest,damage to crop plants and yield reduction in two Costa Rican coffee plantations with Cordia alliodora shade trees

Both model and field estimations were made of the damage inflicted to coffee plants due to the harvest of timber shade trees (Cordia alliodora) in coffee plantations. Economic analyses were made for different coffee planting densities, yields, and both coffee and timber prices.Damage due to tree felling and log skidding should not be a major limitation to the use of timber shade trees in coffee plantations. The timber price that would balance all discounted losses and benefits to zero, for scenarios with and without trees ranged between 8–20 US $/m³ (current overbark log volume at the saw mill yard is US$ 66/m³). There will be lower margins for coffee damage in high yielding plantations, specially in years of good coffee prices. Nevertheless, the use of timber shade trees is recommended even in these scenarios.     

Water use in a Grevillea robusta–maize overstorey agroforestry system in semi-arid Kenya

Novel approaches involving a combination of sap flow measurements of transpiration and allometric estimates of biomass production were used to determine seasonal water use by trees and crops in agroforestry systems. The results were used to test the hypothesis that agroforestry may improve productivity by capturing a greater proportion of annual rainfall than annual crops. Grevillea robusta A. Cunn., which is reputed to have a deep rooting habit, was grown in semi-arid Kenya either as sole stands or in combination with maize (Zea mays L.). Water use by individual trees and maize plants was determined using constant temperature heat balance gauges and scaled to provide stand-level estimates of transpiration based on linear relationships (r²>0.70) between sap flow and leaf area across a range of tree ages and environmental conditions. Maximum stand-level transpiration rates for grevillea ranged from 2.6 to 4.0 mm per day, consistent with previous studies in similar environments. Biomass production by grevillea was closely correlated with stand-level transpiration (r²>0.69–0.74), suggesting that non-destructive estimates of biomass increments can be used to provide reliable estimates of seasonal transpiration. Cumulative water use by grevillea over the 4.5-year observation period was comparable in the sole tree and agroforestry treatments, reaching a maximum utilisation of annual rainfall of 64–68% 3–4 years after planting. Approximately 25% of the water transpired by the trees was used during the dry season, indicating that they were able to utilise off-season rainfall, comprising 16% of the total annual rainfall, and residual water remaining in the soil profile after the cropping period. During the 1995 long rains, when 221 mm of rain was received, transpiration by sole maize was <50% of precipitation, compared to ca. 85% by the trees in the sole grevillea and agroforestry treatments. These results confirm that agroforestry systems may greatly increase rainfall utilisation compared to annual cropping systems. However, careful consideration of the tradeoffs between the loss of crop production and the additional value provided by tree products is essential.     

Growth reduction in harvest-age,coniferous forests with residual trees in the western central Cascade Range of Oregon

To evaluate the relationship of overstory residual trees to the growth of unmanaged young-to-mature understory Douglas fir (Pseudotsuga menziesii (Mirb.) Franco) and western hemlock (Tsuga heterophylla (Raf.) Sarg.), the basal area and volume of 14 paired plots with and without residual trees were examined in the Willamette National Forest, Oregon. Residual trees were large survivors of the fires that initiated the understory between 55 and 121 yr ago. Understory stands were naturally regenerated and not managed in any way. High residual tree and understory densities were negatively associated with understory volume. The relation of density of residual trees to total understory and Douglas-fir basal areas and volumes was best described by a negative logarithmic function. The rate of decrease in total understory and Douglas-fir basal areas and volumes per individual residual tree became smaller with increasing residual-tree density. Predicted total understory volume reduction was 23% with five residual trees/ha and 47% with 50 residual trees/ha, averaging 4.6% and 0.9% per residual tree, respectively. After including the estimated volume growth of residual trees since initiation of the understory, stand volume was still 19% lower with five residual trees/ha and 41% lower with 50 residual trees/ha than in stands with no residual trees, averaging a reduction of 38% and 0.8% per residual tree, respectively. In mixed stands of Douglas fir and western hemlock, predicted Douglas-fir basal area and volume declined more rapidly than did total understory basal area and volume when residual-tree densities exceeded about 15 trees/ha. This difference was probably due to the relative shade-intolerance of Douglas fir. Predicted Douglas-fir volume reduction was 13% with five residual trees/ha and 75% with 50 residual trees/ha, averaging 2.6% and 1.5% per residual tree, respectively. The southern aspects had more than 150% the total understory basal area and volume and more than 200% the Douglas-fir volume and basal area of the northern aspects. Lower density and basal area of understory trees, particularly of dominant and codominant Douglas fir, were associated with increasing residual-tree densities. Given the same diameter at breast height (DBH), heights of Douglas fir were not related to residual trees. Regardless of understory age, understory volume was greatest in stands with the lowest understory densities. These results suggest that timber production in unthinned green-tree retention units may be reduced and may depend on the density of leave-trees. Thinning of understory trees is recommended to reduce growth loss from intraspecific competition.     

Effects of <Emphasis Type="Italic">Inga densiflora</Emphasis> on the microclimate of coffee (<Emphasis Type="Italic">Coffea arabica</Emphasis> L.) and overall biomass under optimal growing conditions in Costa Rica

The advantages of associating shade trees in coffee agroforestry systems (AFS) are generally thought to be restricted mostly to poor soil and sub-optimal ecological conditions for coffee cultivation whereas their role in optimal conditions remains controversial. Thus, the objective of this study was to investigate, under the optimal coffee cultivation conditions of the Central Valley of Costa Rica, the impact of Inga densiflora, a very common shade tree in Central America, on the microclimate, yield and vegetative development of shaded coffee in comparison to coffee monoculture (MC). Maximum temperature of shaded coffee leaves was reduced by up to 5°C relative to coffee leaf temperature in MC. The minimum air temperature at night was 0.5°C higher in AFS than air temperature in MC demonstrating the buffering effects of shade trees. As judged by the lower relative extractable water (REW) in the deep soil layers during the dry season, water use in AFS was higher than in MC. Nevertheless, competition for water between coffee and associated trees was assumed to be limited as REW in the 0–150 cm soil layer was always higher than 0.3 in shaded coffee compared to 0.4 in monoculture. Coffee production was quite similar in both systems during the establishment of shade trees, however a yield decrease of 30% was observed in AFS compared to MC with a decrease in radiation transmittance to less than 40% during the latter years in the absence of an adequate shade tree pruning. As a result of the high contribution (60%) of shade trees to overall biomass, permanent aerial biomass accumulation in AFS amounted to two times the biomass accumulated in MC after 7 years. Thus provided an adequate pruning, Inga-shaded plantations appeared more advantageous than MC in optimal conditions, especially considering the fact that coffee AFS provides high quality coffee, farmers’ revenue diversification and environmental benefits.     

Root competition for phosphorus between coconut, multipurpose trees and Kacholam (Kaempferia galanga L.) in Kerala, India

The magnitude of root competition 17 year-old coconut palms suffer from three year-old inter-planted multipurpose trees, Vateria indica L., Ailanthus triphysa (Dennst.) Alston. or Grevillea robusta A. Cunn. and kacholam (Kaempferia galanga L.), a herbaceous medicinal plant, was evaluated based on the extent of absorption of applied ³²P by the palms in sole and mixed crop situations. The multipurpose tree (MPT) species were grown under two planting geometries (single row and double row). The hypothesis that, when grown together, widespread root proliferation of coconut and multipurpose trees occurs in the well-fertilised kacholam beds was tested by root excavation. Interplanted MPTs substantially altered absorption of ³²P by coconut. Both Ailanthus and Vateria exerted a modest depressing effect, while Grevillea enhanced ³²P uptake by coconut. Single rows of MPTs also favoured ³²P recovery by coconut, presumably because of the increased root densities in the subsoil. Ailanthus, Vateria and Grevillea absorbed substantial ³²P. Overall, high ³²P absorption in the coconut-Grevillea plots indicates complementary root-level interactions between these species. ³²P absorption by MPTs was generally higher closer to the trees owing to the greater root concentration of the MPTs, which in turn suggests possible root interference between MPTs and coconut. Hence selection of tree species with low root competitiveness and/or trees with complementary root interaction is of strategic importance in agroforestry. Kacholam showed substantial ³²P content in its foliage. This ³²P appears to have been translocated by coconut into the kacholam beds where new coconut roots were abundant. This revised version was published online in June 2006 with corrections to the Cover Date.     

Three newly recorded species of the genus Diphascon (Tardigrada; Hypsibiidae) from China

This paper reports three newly recorded species, Diphascon modestum Binda, Pilato and Dastych, 1984, Diphascon nobilei Binda, 1969, and Diphascon triodon Maucci, 1996, of the genus Diphascon (Tardigrada; Hypsibiidae) from China. Five specimens of Diphascon modestum were collected from Taibai Mt. (34°18′N, 107°42tpE) at 2,500 m above the sea level (a.s.l.), two specimens of Diphascon nobilei were collected from Tonggu County, Jiangxi Province (28°31’N, 114°26′E) at 900 m a.s.l, and one specimen of Diphascon triodon was collected from Linzhi County, Tibet (29°40′N, 94°23′E) at 3,500 m a.s.l. All specimens are deposited at the College of Life Sciences, Shaanxi Normal University, China. A key to the Chinese species of Diphascon was also given. Foundation project: This study was supported by scientific research foundation project of Shaanxi Institute of Education (No. 07KJ37Q)     

Comparative performance of seventeen upperstorey tree species associated with crops in the highlands of Uganda

Trials were established at three sites in Uganda to test the suitability of multipurpose trees (MPTs) as upperstorey in crop lands to provide poles, small timber and fuelwood. The three sites were Kachwekano District Farm Institute (1°16 S, 29°57 E, 2000 m.a.s.l.) in Kabale District, Kabanyolo University Farm (0°28 N, 32°27 E, 1250 m.a.s.l.) in Mpigi District and Bushenyi District Farm Institute (0°34 S, 30°13 E, 1610 m.a.s.l.) in Bushenyi District. The MPTs were planted in single rows at intra spacing of 2 m and each plot contained seven or nine trees. On both sides of the tree row, crops were raised. Data on crop yields were collected every season, while data on the growth of the trees were collected four times each year.In terms of tree growth,Grevillea robusta, Casuarina cunninghamiana andAlnus acuminata performed well with height growth of 1.8–2.4 m per year at Kachwekano, while at BushenyiGrevillea robusta, Casuarina junghuhniana, Cupressus lusitanica andCedrela serrulata averaged 1.6–2.0 m height per year. At Kabanyolo,Melia azedarach, Cassia siamea, Jacaranda mimosifolia, Grevillea robusta andMaesopsis eminii had height increments ranging from 1.8–2.7 m annually. Crop yields were affected by the presence of the MPTs, withMaesopsis eminii being the most competitive (averaging 60% reduction, over five seasons). The crop rows nearest to the tree line were the most affected. OnlyAlnus acuminata seems to have had a positive effect on crop yields. The installation of a root mesh to reduce tree root competition for nutrients and water in four species increased yields in plots with MPTs by 5% (Melia azedarach) to 152% (Maesopsis eminii), but the control plot still had significantly higher bean yields, suggesting that shading could also be important. In the case of maize, suppression seems to be due mainly to root competition because after its elimination yields obtained thereafter did not differ significantly from those of the control except for the Maesopsis plots. The negative influence of the MPTs could, therefore, be minimized by periodically pruning the tree crowns and roots.     

Biomass Production and Chemical Composition of Moringa oleifera under Different Management Regimes in Nicaragua

The effects of different planting densities (250,000, 500,000 and 750,000 plants ha⁻¹) and cutting frequencies (45, 60 and 75 days) on the biomass production and chemical composition of Moringa oleifera was studied in a completely randomised split plot design with four blocks, in Managua, Nicaragua, located geographically at 12°08′15′′ N and 86°09′36′′ E. The 75 day cutting frequency produced the highest fresh matter yield, 100.7 and 57.4 Mg ha⁻¹ year⁻¹, and dry matter (DM) yield, 24.7 and 10.4 Mg ha⁻¹ year⁻¹, during the first and second year, respectively. All planting densities produced the highest DM yield at 75 day cutting frequency. In the first year, the density of 750,000 plants ha⁻¹ produced the highest fresh matter yield, 88.0 Mg ha⁻¹ and highest DM yield, 18.9 Mg ha⁻¹, but in the second year the density of 500,000 plants ha⁻¹ gave the highest yields, 46.2 Mg ha⁻¹ and 8.1 Mg ha⁻¹, respectively. During the first year, DM (22.8%), neutral detergent fibre (NDF) (30.8%) and ash (9.14%) contents were highest and in vitro DM digestibility (IVDMD) (68.2%) was lowest in the longest cutting interval, while contents of crude protein (CP) (22.8%) and acid detergent fibre (ADF) (22.8%) were not affected significantly by cutting frequency. In the second year, DM and CP contents and IVDMD were not significantly affected by cutting frequency, whereas NDF, ADF and ash contents were lowest in the 60 day cutting frequency. Planting density had no significant effect on chemical composition or IVDMD. These data suggest that Moringa forage could be an interesting protein supplement for ruminants.