Sward dynamics of a smooth-stalked meadowgrass dominantwhite clover sward rotationally grazed by cattle and/or sheep

This experiment was carried out to study the responses of sward components (particularly white clover, Trifolium repens ) to grazing management in a natural sward dominated by smooth-stalked meadowgrass ( Poa pratensis ) syn. Kentucky bluegrass. Treatments during two grazing seasons (1989–90) were: cattle grazing alone (C); cattle grazing followed by topping (CT); cattle grazing followed by sheep grazing (CS); and sheep grazing alone (S). Mean target pre- and post-grazing herbage masses were 2200 and 1100 kg DM ha⁻¹, estimated by single-probe electronic capacitance meter. Sward component dynamics were monitored using turf dissections, marked white clover stolons, and ring-toss white clover leaf counts. Component and sward data for the C, CT, CS and S treatments respectively, were: number of white clover leaves m⁻², 1295, 1384, 1408, 900 (s.e. ± 108); number of leaves per growing point, 3·2, 3·4, 3·0, 2·8 (s.e. ± 0·2); herbage accumulation (t DM ha⁻¹), 5·16, 5·02, 5·87, 8·28 (s.e. ±0 08); rejected herbage (% pasture area) 39·7, 7·7, 16·0, 0 (s.e. ± 75); and annual net herbage production (t DM ha⁻¹) 3·39, 4·35, 4·99, 8·28 (s.e. ± 0.07). Swards grazed by sheep alone contained less white clover, but regrew quicker and produced more herbage than other treatments. Close topping or grazing by sheep following dairy cattle grazing decreased sward rejection by cattle. These treatments maintained more of the pasture in better condition for subsequent cattle grazing, resulting in greater net herbage production than where no post-cattle grazing treatment was used.     

Seasonal changes in the crude protein concentration of mixed swards of white clover/perennial ryegrass grown without fertilizer N in an organic farming system in the United Kingdom

Changes in the crude protein (CP) concentration of white clover and perennial ryegrass herbage from a mixed sward were determined on six sampling dates from May to October in each of 2 years. The swards were grown without fertilizer N in an organic farming system and continuously grazed by dairy cows during the grazing season. The annual mean contents of white clover in the dry matter (DM) of the sward were 272·3 and 307·0 g kg⁻¹ in Years 1 and 2. The mean CP concentrations of the white clover and perennial ryegrass herbage were 251·6 and 151·9 g kg⁻¹ DM in Year 1 and 271·9 and 174·0 g kg⁻¹ DM in Year 2 respectively. The CP concentration of the white clover increased significantly during the grazing season from 220·0 to 284·1 g kg⁻¹ DM in Year 1 and from 269·0 to 315·5 g kg⁻¹ DM in Year 2. In the perennial ryegrass herbage the CP concentration increased from 112·2 to 172·6 g kg⁻¹ DM in Year 1 and from 142·7 to 239·5 g kg⁻¹ DM in Year 2. The rate of increase during the season in the CP concentration of the perennial ryegrass herbage was similar to the rate of increase recorded in the white clover herbage.     

The liveweight gain of Limousin × Friesian heifers grazing perennial ryegrass/white clover swards of different clover content and the effects of their grazing on sward botanical composition

An experiment was carried out in 1992 and 1993 to examine the effect of white clover content of perennial ryegrass/white clover swards on the performance of Limousin × Friesian heifers. Swards with low (L), medium (M) and high (H) white clover contents were established and managed by continuous variable stocking. A compressed sward height of 5·5 cm was maintained using a buffer fence to vary plot areas, with herbage surplus to grazing requirements cut, removed and yields measured. The mean white clover proportions for treatments L, M and H were 0·02, 0·19 and 0·18 in 1992 and 0·13, 0·16 and 0·31 in 1993 respectively. White clover contents of the swards reached a maximum in August and September, and differences between treatments diminished. There was no significant difference between treatments in the content of white clover in the swards in autumn 1993.
Liveweight gains of heifers increased asymptotically with increasing white clover content of the sward. Below a white clover herbage mass of 300 kg DM ha^–1, there was little effect on liveweight gain, which was 0·70 kg day^–1 over the grazing season. Between 400 and 450 kg DM ha^–1 white clover, liveweight gains were 0·85–0·90 kg day^–1. While clover content of the sward did not significantly affect utilized metabolizable energy output; the mean output over the grazing season in the two years from liveweight gain and herbage yield was 78 GJ ha^–1. It is suggested that, using this grazing system, white clover reached an equilibrium with a mean herbage mass of about 400 kg DM ha^–1 over the grazing season.     

Effect of long-term changes in relative resource availability on dietary preference of grazing sheep for perennial ryegrass and white clover

Three replicate paddocks, each of 0·235 ha, containing adjacent monocultures of perennial ryegrass or white clover [50:50 by ground area, 6 cm sward surface height (SSH) at start of experiment] were continuously stocked with three yearling and four mature non-lactating, non-pregnant Scottish halfbred ewes for 12 weeks. Herbage intake, grazing behaviour and dietary selection were measured on seven occasions. Clover SSH declined rapidly over the first 5 weeks then stabilized at 1·2–1·6 cm, whereas perennial ryegrass SSH rose slightly initially, then declined gradually. Animals initially included proportionately c . 0·6 white clover in their diet but, by the end of the experiment, this had fallen to 0·3. Total daily herbage intake declined over the 12 weeks from 1·8 kg dry matter (DM) day^–1 at the start to 1·0 kg DM day^–1. Total grazing time increased from 561 min day^–1 to 649 min day^–1 at the end of the experiment. The results suggest that, despite overall herbage depletion and a greater depletion of white clover than perennial ryegrass as a result of the initial partial preference for white clover, the animals traded-off a reduced total intake and an increased grazing time in an attempt to maintain their initial preferred dietary composition.     

Effects of application of nitrogen fertilizer on concentrations of P, K, S, Ca, Mg, Na, Cl, Mn, Fe, Cu and Zn in perennial ryegrass/white clover pastures in south-western Victoria, Australia

Effects of timing and rate of N fertilizer application on concentrations of P, K, S, Ca, Mg, Na, Cl, Mn, Fe, Cu and Zn in herbage from perennial ryegrass/white clover pastures were studied at two sites in south-western Victoria, Australia. Nitrogen fertilizer (0, 15, 25, 30, 45 and 60 kg ha^–1) was applied as urea in mid-April, early May, mid-May, early June and mid-June 1996 to pastures grazed by dairy cows. At Site 1, N fertilizer resulted in a linear increase in P, K, S, Mg and Cl concentrations in herbage and a linear decrease in Ca concentration. For all times of application, concentrations of P, K, Ca, Mg and Cl in herbage increased by 0·0048, 0·08, −0·010, 0·0013 and 0·053 g kg^–1 dry matter (DM) per kg N applied respectively. For S concentration, maximum responses occurred in mid-May (0·012 g kg^–1 DM per kg N applied). At Site 2, N fertilizer resulted in a linear increase in P, S and Na concentrations in herbage, a linear decrease in Ca concentration and a curvilinear increase in K and Cl concentration. The maximum responses for P, S and K concentrations in herbage occurred for the N application in mid-June and were 0·015, 0·008 and 0·47 g kg^–1 DM per kg N applied respectively. For Cl concentration, the maximum response occurred for the N application in early June and was 0·225 g kg^–1 DM per kg N applied. Overall, applications of N fertilizer up to 60 kg ha^–1 did not alter herbage mineral concentration to levels that might affect pasture growth or animal health.     

Effects of interval between harvests and application of fertilizer N in spring on the growth of perennial ryegrass in a grass/white clover sward

A perennial ryegrass/white clover sward, which had been grazed for over 2 years, was cut at 1-, 2-, 3- or 6-week intervals from 18 April to 28 November 1986. Two rates of fertilizer N application in spring, 0 and 66 kg N ha⁻¹, were compared. Perennial ryegrass growth was studied in three 6-week periods, beginning on 18 April, 18 July and 17 October. Clover growth was studied in the same three periods and described by Fisher and Wilman (1995) Grass and Forage Science , 50 , 162–171.
Applied N increased the number of ryegrass tillers m⁻², the rate of leaf extension and the weight of new leaf produced tiller⁻¹ and m⁻²per week. Increasing the interval between cuts reduced the number of ryegrass tillers m⁻² and increased the rate of leaf extension, weight tiller⁻¹and the weight of new leaf produced tiller⁻¹week⁻¹. Increasing the interval between cuts increased the weight of new ryegrass leaf produced m⁻² where N had recently been applied, but otherwise had little effect on the weight of new leaf produced m⁻². Applying N favoured the grass in competition with the clover in every respect, whereas increasing the interval between cuts only favoured the grass, compared with clover, where N had recently been applied; where N had not been applied, the ratio of ryegrass tillers to clover growing points in the sward was very little affected by the interval between cuts.     

Intake and behaviour responses by sheep to changes in sward characteristics under continuous stocking

Perennial ryegrass pastures were maintained at sward surface heights (SSH) of 30, 60, 90 and 120 mm by continuous variable stocking using lactating ewes and their twin lambs in spring and non-lactating (dry) ewes in autumn.
The effects of SSH on ingestive behaviour, herbage intake, animal performance and sward structure and production were measured in spring, and again in autumn, when an additional SSH of 20 mm was also established.
In spring, differences in sward structure were quickly established with mean tiller numbers of 41 000, 30 000, 21 000 and 19 000 m⁻² and leaf area indices (LAI) of 1·5, 2·2, 3·3 and 4·1 at SSH of 30 to 120, respectively. By autumn the mean tiller numbers had fallen to 26 000, 26 000, 23 000, 18 000 and 13 000 m⁻² and LAI to 1·0, 1·1, 1·5. 2·0 and 1·5 for treatments 20 to 120.
The mean stocking rates for the ewes in spring were 27, 20, 22, and 19 ewes ha⁻¹ and growth rates of their lambs were 208, 275, 250 and 263 g d⁻¹ for treatments 30 to 120. In autumn the stocking rates maintained for dry ewes were 22, 15, 9, 4 and 0 for treatments 20 to 120.
In spring, both grazing times (GT) and prehension biting rate (BR) were negatively correlated with SSH, while bite mass (BM) was positively correlated with SSH. However, mean daily intakes of organic matter (OM), were reduced only for animals grazing the 30 mm sward.
In autumn, intake rate was lower only for the animals grazing the 20mm sward and GT and BR were not significantly affected by SSH.
Relationships between SSH and, components of ingestive behaviour and intake, are presented and the control mechanisms involved are discussed. The results showed that the optimum sward surface height for continuously stocked swards, grazed by sheep, Was between 30 mm and 60 mm.     

Forage availability from a temperate pasture managed with intensive rotational grazing

Successful integration of rotational grazing into livestock production systems requires estimates of pasture growth rates for feed budgeting of daily animal intake. By matching livestock nutrient demand with forage availability, over-feeding of supplements can be minimized, which reduces feed costs and the need lo manage surplus nutrients, A three-year grazing study was carried out on a Kentucky bluegrass ( Poa pratensis L.)-dominant pasture to estimate the daily quantity of herbage available to cattle in an intensive, rotational grazing system. Herbage production, species composition, and forage quality were determined in each of the six grazing cycles in a year, from April until September. The average length of a grazing cycle was 28·6 d, with 2·7 d for duration of grazing on a paddock. Pre-grazing and post-grazing sward heights, measured with a plate meter, were 14 and 7 cm, and the corresponding herbage masses were 1955 and 775 kg DM ha⁻¹ respectively. Under adequate soil moisture during 1989, herbage available for daily intake was 53 kg ha⁻¹ from April until mid- August, declining to approximately 32 kg ha⁻¹ d⁻¹ by the end of September. Distribution of this herbage was fairly uniform until the end of August. However, a dry summer in 1991 reduced herbage availability to 15 kg ha⁻¹ d⁻¹. Bluegrass and white clover ( Trifolium repens L.) formed 70% of the herbage yield during the period April–June. Later in the season, dead matter and other species increased, reducing the contribution of bluegrass and clover to approximately 60% of total dry matter. While these pastures have the potential to provide significant amounts of forage for 5–6 months in a year, additional on-farm forage reserves are needed during periods of water stress.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK. 5. The effect of stocking rate and reduced levels of nitrogen fertilizer

The implications for UK upland sheep systems of reducing nitrogen fertilizer application to perennial ryegrass/white clover swards were studied over 3 years. Sward height (3·5–5·5 cm) was controlled for ewes with lambs until weaning using surplus pasture areas for silage; thereafter, ewes and weaned lambs were grazed on separate areas, and sward height was controlled by adjusting the size of the areas grazed and using surplus pasture areas for silage if necessary. Combinations from three stocking rates [10, 6 and 4 ewes ha⁻¹ on the total area (grazed and ensiled)] and four nitrogen fertilizer levels (150, 100, 50 and 0 kg ha⁻¹) provided six treatments that were replicated three times. Average white clover content was negatively correlated with level of nitrogen fertilizer. The proportion of white clover in the swards increased over the duration of the experiment. Control of sward height and the contribution from white clover resulted in similar levels of lamb liveweight gain on all treatments. All treatments provided adequate winter fodder as silage. It is concluded that the application of nitrogen fertilizer can be reduced or removed from upland sheep pastures without compromising individual animal performance provided that white clover content and sward height are maintained. Resting pastures from grazing by changing ensiled and grazed areas from year to year sustained white clover content over a 3-year period.     

The effects of regrowth and maintenance height on a grass sward with a high density of tillers

A perennial ryegrass sward was grazed by sheep in April 1993 to a target sward surface height (SSH) of 3 cm to create a high density of grass tillers. From 3 May, the sheep were removed and small plots were established on the sward, when the average tiller density (± s.e.) was 35 900 ± 420 live tillers m⁻². Different regrowth treatments were then imposed by allowing plots to regrow to target SSHs of 6 cm (18 g dry matter (DM) M⁻²), 9 cm (78 g DM m⁻²), 12 cm (132 g DM m⁻²) or 15 cm (197 g DM m⁻²). The plots were then maintained by cutting at either 6 or 9 cm SSH until the end of the experiment on 30 September. Live tiller density was reduced by regrowth beyond 8.9 cm (78 g DM m⁻², P <0.001) and leaf-stem ratio and in vitro organic matter digestibility were reduced by regrowth beyond 6.1 cm (18 g DM m⁻², P < 0.05). The effect on live tiller density was sustained through the remainder of the season. From the beginning of June to the end of September, maintenance of SSH at an average of 9.1 cm compared with 6.4 cm also resulted in lower live tiller density, live-dead tiller ratio and leaf-stem ratio and higher herbage mass (at least P < 0. 05). There were significant interactions between regrowth SSH and maintenance SSH, so that leaf-stem ratio, live-dead tiller ratio and live tiller density were reduced by regrowth to a SSH of 16–4 cm followed by maintenance at 9.1 cm, compared with regrowth to 6.1 cm and maintenance at 6.4 cm.     

The effect of herbage allowance on daily intake by Creole heifers tethered on natural Dichanthium spp. pasture

Two experiments were carried out in Guadeloupe to estimate the organic matter intake (OMI) and digestibility (OMD) of a Dichanthium spp. sward, grazed by tethered Creole heifers [mean live weight (LW) 202 ± 2·0 kg], at three daily herbage allowances. Experiment 1 examined herbage allowances of 16, 25 and 31 kg of dry matter (DM) d^–1 on a fertilized sward at 21 days of regrowth whereas, in experiment 2, lower allowances of 11, 15 and 19 kg DM d^–1 were examined on the same sward, which was unfertilized and grazed at 14 days of regrowth. In each experiment, the herbage was grazed with three groups of two heifers in a 3 × 3 Latin square design. Sward characteristics were described before grazing. OMI was calculated from total faecal output, and OMD was predicted from the crude protein (CP) content of the faeces. The amount of herbage defoliated by the heifers was also estimated on tillers selected at random.
Organic matter intakes were on average 26 g and 19 g OM kg^–1 LW, and OMD values were 0·740 and 0·665 for Experiments 1 and 2, respectively, and were not affected by allowance. In Experiment 1, the herbage quality was high [0·50 of leaf and 116 g CP kg^–1 organic matter (OM)] for a tropical forage, whereas in Experiment 2, the quality of the herbage (0·27 of leaf and 73 g CP kg^–1 OM) was lower. These differences were reflected in differences in intake and digestibility in the two experiments.
The experimental tropical Dichanthium spp. swards can have intake characteristics similar to those of a temperate sward.     

Cattle slurry as a source of nutrients for red clover: herbage production and clover contribution

An established sward of red clover cv. Hungaropoly sown pure received approx. 30 kg P ha^-1 and 200 kg K ha^-1 each year for 3 successive years. The P and K were applied either as cattle slurry, inorganic fertilizer or combinations of these. Treatments were applied either in spring or after the first harvest. There were a total of six treatments and these were harvested three times each year. The average yields of total herbage DM over all the treatments in the first, second and third years were 15·2, 14·2 and 14·2 t ha^-1 respectively and the various treatments had no significant effect on the overall yields.
Treatments had a significnt effect on red clover DM yields and percentage red clover in one harvest in each of the first 2 years and all three in the third year. Yields of red clover were lower and grass higher in treatments receiving cattle slurry only. On this treatment there was a total yield of 23·2 t ha^-1 red clover DM in the 3 years compared with 30·2 t ha^-1 on the inorganic fertilizer treatments. However, by applying P and K fertilizer in the spring, followed by cattle slurry after the first harvest, it was possible to maintain a high proportion of red clover in the sward and to produce yields of red clover DM similar to those on the inorganic fertilizer treatments.     

The intake and performance of cashmere goats grazing sown swards

An experiment was conducted to examine the relationships between sward surface height and the intake and liveweight gain of lactating female goats and their single kids and of 12-month-old castrated male goats continuously grazing grass pasture. Herbage intake was measured using the n -alkane marker technique. Goats grazed the experimental area from May to August at nominal sward heights of 3–4, 5–6, 7–8 and 9–10 cm. Sward heights achieved were variable but were consistently ranked in treatment order.
The herbage intake of females [57-140 g DM (kg LW^0.75)⁻¹ d⁻¹] and castrates [26-88 g DM (kg LW^0.75)⁻¹ d⁻¹] increased linearly with sward height over the range 3.2-11.0 cm. The liveweight change of females and castrates reflected the pattern of change in sward height.
The herbage intake of kids [17-41 g DM (kg LW^0·75)⁻¹ d⁻¹] was not related to sward height, but there was a consistent increase in liveweight gain with sward height from 98 to 129 g d⁻¹. The herbage intake of kids did not increase with age with means of 25–29 g DM (kg LW 0·75 d⁻¹ for kids aged 9–17 weeks.
There was evidence that all classes of goat selected green leafy material of high digestibility.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK. 1. Effects of two annual stocking rates in combination with two sward height profiles

The consequences of controlling sward height at two levels, around 5 cm (HS) and around 3·5 cm (LS), during spring and summer by adjusting weekly the proportion of the areas grazed in response to changes in rate of herbage production and utilizing the surplus pasture areas to conserve winter fodder were studied over three complete years for Greyface ewes with their lambs at 15ha⁻¹ (SR15) and 10ha⁻¹ (SR10). The rules used to control grazed sward height resulted in acceptable sward height control in three of four treatments and, by providing supplementary feed when sward height was below target, gave rise to similar levels of individual animal performance. Total output of lamb weaned was greater for the SR15 than for the SR10 flocks (607 vs 477 kg live weight ha⁻¹; P <0·001). Amounts of winter fodder produced were less for the SR15 than for the SR10 flocks [39 vs 213kg dry matter (DM) per ewe; P < 0·001]. Significantly more supplementary feed (10·0 vs 4·4kg DM per ewe; P < 0·01) and hay (13·7 vs 4·0kg DM per ewe; P7lt;0·05) were offered around mating to SR15 flocks than to SR10 flocks. It is concluded that, provided that sward height can be controlled between 3·5 and 5·0cm during the spring and summer and that supplementation is offered when the grazed sward is below 3·5 cm, flock performance will fall within acceptable and predictable limits.     

A comparison of diploid and tetraploid perennial ryegrass and tetraploid ryegrass/white clover swards under continuous sheep stocking at controlled sward heights. 1. Sward characteristics

Two 1·0 ha plots of a late-heading diploid perennial ryegrass (var. Contender) and a late-heading tetraploid ryegrass (var. Condesa), and two 1·4 ha plots of the tetraploid with Aberystwyth S184 small-leaved white clover, were direct sown in May 1987. Over the three years 1988–90 they were continuously stocked by Mule ewes with Suffolk-cross twin lambs, from early April to the end of August, at a target sward surface height (SSH) of 4–6 cm on one set of plots (constant swards) and, on the other set, al 4–6 cm rising after June to a target 6–8 cm (rising swards). The heights were achieved by variable stocking. Fertilizer N was applied only to the grass plots at the rate of 150- 180kgN ha^-1 annually.
SSH was mainly within the target 4–6 cm, after higher initial heights at turnout in 1988and 1990. Mean heights of the constant swards (April- August) averaged 5·53, 4·43 and 5·04cm in the three years. The rising swards (July-August) increased in height over the constant swards by an average of 0·88, 0·48 and 0·55 cm, in successive years.
Clover content of the herbage mass dry matter in the grass/clover swards increased over each grazing season to average 13·0, 26·5 and 21·2% in the three years, with a high mean stolon density of 130 in m^-2 in August 1990. Ryegrass tiller densities in year 3 were 23% higher in the diploid than in the tetraploid swards, which had 43% more than the 10000 tillers m^-2 of the tetraploid ryegrass/clover swards.
It is concluded that the combination of a densely stoloniferous small-leaved clover with the open growth habit of a tetraploid ryegrass can achieve swards of high clover content under continuous sheep stocking.     

Herbage intake and liveweight gain of bulls and steers continuously stocked at fixed sward heights during autumn and spring

Two experiments were conducted to examine the relationship between sward surface height, herbage intake and liveweight gain in beef cattle grazed on pasture. In Experiment 1, two 'animal types' (18 Charolais × Angus steers and 18 Friesian bulls) were continuously grazed for 22 days during the late autumn on replicated swards maintained at sward surface heights of 6, 10 and 15 cm. Herbage intakes, assessed from the faecal concentration of chromium delivered from an intraruminal controlled release capsule and the in vitro digestibility of hand-plucked herbage samples, were curvilinearly related to sward height (r = 0·76, p <0·0·01). Average liveweight gains were 0·02, 0·61 and 1·31 kg d^-1 ( P <0·05) and increased linearly ( r = 0·84, P <0·001) with sward height. The maintenance organic matter intakes of the steers and bulls, with initial mean (± s.e.) live weights of 225 ± 15 kg and 172 ± 15 kg respectively, were estimated to be 3·6 and 3·5 kg d^-1 respectively. In Experiment 2 (spring) 36 cattle, including 35 of those used in Experiment 1, were reallocated to sward heights of 5, 10 and 15 cm using the same design as for Experiment 1. Average liveweight gains were 0·94, 1·57 and 1·68kg d^-1 ( p 0·05) and were curvilinearly related to sward height ( r = 0·093, p <0·05). Maintenance intakes could not be reliably extimated for the cattle in Experiment 2 because few animals had liveweight gains close to zero. These trials confirm that liveweight gain in continuously grazed finishing steers and bulls increases with sward surface height to maximum of 8–10cm with spring ryegrass/white clover pastures while, in autumn, swards of 12–15cm height are required to achieve maximum performance.     

The effect of herbage allowance upon the herbage intake and performance of suckler cows and calves

Twenty-four Hereford × Friesian cows and their South Devon cross calves were allocated to three herbage allowances allotted daily for three periods of 8 weeks in a Latin square design. The daily allowances were 17, 34 and 51 g dry matter per kg cow plus calf live weight. Milk production was depressed by 0·2 and 1·2 kg d^-1 at the medium and low allowances. The corresponding falls in liveweight gain were for cows 0·26 and 0·25, and for calves 0·27 and 0·24 kg d^-1. Residual sward height after grazing gave a better indication of the animals' reaction to sward conditions and the management imposed than actual herbage allowance. The quantity per unit area and the composition of material present were important factors influencing intake. Calves were unable to compete with their dams to maintain herbage intake at the lower allowances and therefore are likely to benefit from additional feeding or creep grazing when residual sward height falls below 6cm for periods in excess of 1–2 weeks.     

The herbage intake and performance of set-stocked suckler cows and calves

Groups of eight Hereford × Friesian cows and their South Devon cross calves were set stocked over a 24-week grazing season at 3·23 (low), 3·21 (medium) or 4·24 (high) cows ha^-1 together with their calves. For the first 8 weeks only two-thirds of the total area was grazed. Increasing the stocking rate from low to medium reduced daily milk yield and cow and calf liveweight gains by 1·2, 0·24 and 0·29 kg d^-1 respectively, and from medium to high by 1·2, 0·24 and 0·23 kg d^-1. The main sward factor influencing faecal output and herbage intake was the quantity of organic matter on the pastures but the digestibility of the herbage selected also exerted a significant effect on the intake of cows. Major depressions in the herbage intake of cows occurred once the average sward height fell below 7 cm. Output of calf live weight was 628, 658 and 743 kg ha^-1 for the 3 stocking rates from low to high, and for cows 246, 179 and 30 kg ha^-1. It was concluded that decisions on pasture management should be taken in relation to the cow rather than the calf on set-stocked swards.     

Effect of regrowth interval on the productivity of swards defoliated by cutting and grazing

Over three grazing seasons (1984-1986) a sward of perennial ryegrass, cv. Talbot, which received a total of 336 kg N ha^-1 each season, was cut or grazed with ewes at 3- or 4-week intervals on a rotational basis.
Sward productivity was higher under cutting than under grazing irrespective of the interval between defoliations. Under cutting, mean herbage organic matter (OM) yields over both intervals were 8·66, 9·62 and 8·17 t ha-¹ in 1984, 1985 and 1986 respectively while under grazing the corresponding yields were 7·65, 8·63 and 7·50 t ha^-1. The mean annual yield of herbage defoliated at 3-week intervals was 7·50, 8·64 and 7 ·20 t OM ha^-1 compared with 8·80, 9·60 and 8·46 t OM ha^-1 for swards defoliated at 4-week intervals in the three years respectively.
The nitrogen (N) content of both the available and the residual herbage was consistently higher under grazing than under cutting. Available herbage contained 31·3 and 27·7 g N kg OM^-1 and residual herbage 26·1 and 22·7 g N kg OM^-1 under grazing and cutting respectively.
The mean yield of N under cutting was 284 kg ha^-1 compared with 304 kg ha^-1 under grazing. Defoliation interval had no effect on N yield, the overall mean yield being 294 kg ha^-1 under both 3- and 4-week defoliation intervals. The effect of the treatments on tiller population was slight and inconclusive.
The process of grazing reduced yield probably as a result of damage to the sward through trampling; the positive effect of excretal N on yield was minimal on account of the short grazing periods.     

Growth, colonization and productivity of two white clover cultivars strip-seeded into an upland Festuca-Agrostis sward

Small plots of a Festuca-Agrostis upland sward on a peaty gley podsol were strip-seeded during late June 1986 with white clover cvs Aberystwyth S184 or Menna at 4 kg ha⁻¹ and defoliated early (20 August) or late (3 September) and then frequently or infrequently (every 2 weeks or 4 weeks) until the end of September. All plots were defoliated in early November, at 3-weekly intervals during the growing season in 1987 and then grazed rotationally during 1988.
Satisfactory seedling establishment, representing 46% emergence, was achieved 5 weeks after sowing. The differential defoliation regimes had no persistent significant effects on clover development. S184 soon produced more leaves per seedling than Menna and a smaller proportion of its leaf number and weight were removed at each defoliation. Following large losses of leaves over the 1986–87 winter, SI84 had significantly more leaves per stolon than Menna; subsequently it also colonized the sward at a quicker rate. During 1987 amounts of herbage harvested (6.1 t ha⁻¹) were similar with the two clover cultivars, with S184 contributing 47% and Menna 44% of this respectively. SI84 made a larger contribution to yield during May and June but Menna was more productive during September and October. During 1988 clover populations were maintained with rotational grazing without additional fertilizer inputs.
The results show that, despite initial soil and climatic contraints, both small and medium-leaved clovers can be strip-seeded into upland swards with large subsequent benefits to yield and herbage quality. However, they also indicate the need for further experiments to determine the influence of sward morphology and defoliation regime on stolon branching rates and accumulation of growing points which, in turn, govern sward colonization.