Bat response to shelterwood harvests and forest structure in oak-hickory forests

Forest management practices, such as shelterwood harvesting, can greatly impact bat habitat relationships. Such practices can alter the amount of structural volume within a forest, which can influence bat foraging patterns. We determined the effects of shelterwood harvests of different retention levels (50% and 70% of full stocking) on bat activity patterns in oak-hickory forests located in southern Ohio. We used the Anabat system to monitor activity during May-September 2006. Our objectives were to quantify the effects of harvesting on structural volume and use the results to explain variations in bat activity. Because harvesting alters vertical structure as well as the total amount of volume within a forest, we also determined the height within the vertical profile where changes in structural volume begin to influence overall and species-specific activity. Overall bat activity did not differ significantly between shelterwood harvest levels, but was significantly different between harvested and control sites, with more passes detected within the harvested sites. Lasiurus borealis (red bat), Eptesicus fuscus (big brown bat), and Lasionycteris noctivagans (silver-haired bat) activity was significantly greater in harvested versus control sites, but did not differ between shelterwood harvest levels. Myotis spp. (Myotis lucifugus (little brown bat) and Myotis septentrionalis (northern Myotis)) and Perimyotis subflavus (tri-colored bat) activity did not vary between shelterwood harvest levels or between harvested and control sites. The greatest reductions in structural volume occurred in the understory to mid-canopy of the shelterwood harvests. Overall activity was most influenced by the amount of volume within 3-6 m above the forest floor, and declined as volume within that height strata increased. Mean bat passes declined by 50% when volume within 3-6 m exceeded 17 m³/ha. Estimated use by L. borealis decreased by 50% at volumes exceeding 1750 m³/ha in the understory to mid-canopy (0-12 m), while E. fuscus and L. noctivagans estimated use was the highest when volumes within 3-6 m were less than 63 m³/ha. Our results suggest that forest management practices that reduce the amount of structural volume in the understory to mid-canopy provide suitable habitat for foraging bats. Quantifying the amount of structural volume at various heights within the vertical profile of the forest can lend valuable insights into overall and species-specific bat activity patterns.     

Use of modified water sources by bats in a managed pine landscape

In eastern North Carolina, USA, intensively managed pine (Pinus spp.) forests, a primary forest type in the Southeastern United States, often are established where water levels are maintained via ditching. In these landscapes, water occurs in numerous linear ditches and several small ponds (heliponds), used by helicopters for the suppression of forest fires. Changes to the physical structure of water sources may be important to bats that use these sources for drinking and foraging. Therefore, we examined bat activity in a managed pine landscape and an adjacent natural wetland in relation to water source type (heliponds, ditches, and natural wetland) and insect abundance. We sampled bat activity using remote acoustic detectors; we sampled insects using passive traps. Insect abundance did not differ among water source types. Bat activity was most concentrated at modified water sources within the managed pine landscape (heliponds and ditches). Feeding activity was highest at heliponds. Heliponds and ditches were important sources of open water and insect prey for bats in the managed pine forest landscape. We suggest management for bats in similar forest landscapes should include maintenance of open and accessible water sources such as heliponds.     

Response of bat species to sylvo-pastoral abandonment

We assessed the effect of abandonment of sylvo-pastoral practices in chestnut orchards (Castanea sativa) on bats in southern Switzerland to determine practical recommendations for bat conservation. We compared bat species richness and foraging activities between traditionally managed and unmanaged chestnut orchards, testing the hypothesis that managed orchards provide better foraging opportunities and harbour more bat species. Echolocation calls of foraging bats were sampled simultaneously at paired sites of managed and unmanaged orchards using custom made recorders. Vegetation structure and aerial insect availability were sampled at the recording sites and used as explanatory variables in the model. In a paired sampling design, we found twice the number of bat species (12) and five times higher total foraging activity in the managed chestnut orchards compared to the unmanaged ones. Bat species with low flight manoeuvrability were 14 times more common in managed orchards, whereas bats with medium to high manoeuvrability were only 5 times more common than in abandoned orchards. The vegetation structure was less dense in managed orchards. However, management did not affect relative insect abundance. Bats primarily visited the most open orchards, free of undergrowth. As a result of restricted access into the overgrown forests, the abandonment of chestnut orchards leads to a decline in bat species richness and foraging activities. Continued management of chestnut orchards to maintain an open structure is important for the conservation of endangered bat species in the southern Swiss Alps.     

Vegetation community responses to different establishment regimes in loblolly pine (Pinus taeda) plantations in southern MS,USA

Management treatments involving multiple herbicide applications are standard procedure on most industrial pine plantation sites in the southern USA, raising concerns about biodiversity impairment. Management decisions impact not only plant communities but also the habitat potential they create for wildlife. We tested the effects of five intensities of stand establishment treatments on vegetation communities in loblolly pine plantations (n = 4) to age 5 in the Middle Coastal Plain (MCP) of Mississippi. Measurements were species richness, diversity, coverage, and community composition. Treatments were combinations of mechanical site preparation (MSP), chemical site preparation (CSP), and herbaceous weed control (HWC) both banded or broadcast for 1 or 2 years using the same herbicide mixtures. Tree richness and diversity were reduced by increasing treatment intensity; tree coverage, which included crop and non-crop trees, was less in moderate-intensity treatments. Vine richness and coverage were less in more intensive treatments, but 2 diversity indices differed on whether vine diversity was likewise affected. Richness and coverage of forbs and graminoids was lessened by broadcast HWC, with effects mostly limited to the year of application. Plant communities differed in all 5 years, with CSP acting as the primary factor for years 2–5. Early seral communities were favored by CSP, but broadcast HWC suppressed resulting herbaceous plants. Though CSP may somewhat reduce stand-level plant diversity, it may increase overall biodiversity within plantation-dominated landscapes by creating early succession plant communities that enhance wildlife habitat.     

Ground-dwelling invertebrates in reforested conifer plantations

Plantation forests are an important part of the forest estate in many countries. In Ireland, they cover around 9% of the land area and many that are commercially mature are now being felled and reforested. The potential biodiversity value of such second rotation forests has yet to be determined, yet this may be particularly significant in Ireland where cover of semi-natural woodland is only 1%. Invertebrates are a vital component of forest biodiversity, functioning as decomposers and pollinators, herbivores, predators and prey. Spiders and Carabid beetles are often used in biodiversity assessment as they are easily captured using pitfall traps, are taxonomically well known and respond to changes in habitat structure. This study aimed to examine spider and Carabid beetle diversity in second rotation Sitka spruce (Picea sitchensis) plantations at different stages of the forest cycle (5, 8–12, 20–30, 35–50 years), and compare the spiders captured in second rotation forests with those from first rotation. Spider and beetle diversity was influenced by stand structural development in second rotation plantations with numbers of forest-associated species increasing over the forest cycle. Overall, spider richness declined over the forest cycle and this was related to decreasing cover of field layer vegetation and fewer open-associated species. In contrast, total beetle richness increased and became more specialised over the forest cycle which may be related to slower colonisation of disturbed areas by beetles in comparison with spiders, and fewer open specialists at the early stages of second rotation. Spider assemblages were distinguished between rotations. This may be related to differing habitat conditions in second rotation forests including dryer soils with lower pH, differing vegetation complexity and presence of brush piles. Few of the forest species accumulated during first rotation were retained and the early stages of second rotation forest cycle was characterised by a generalist open fauna. Nonetheless, as the forest cycle progressed the spider assemblages between rotations became more similar. Current forest policy supports retaining over-mature trees and creating a mosaic of different aged stands within a plantation. Such measures may provide refuge for forest species after clearfell. In countries where forest fragments exist in a landscape dominated by agriculture, consideration should be given to the capacity of mature forest adjacent to felled stands to support forest species, and to the configuration of over-mature areas retained after felling.     

Objectives and considerations for wildland fuel treatment in forested ecosystems of the interior western United States

Many natural resource agencies and organizations recognize the importance of fuel treatments as tools for reducing fire hazards and restoring ecosystems. However, there continues to be confusion and misconception about fuel treatments and their implementation and effects in fire-prone landscapes across the United States. This paper (1) summarizes objectives, methods, and expected outcomes of fuel treatments in forests of the Interior West, (2) highlights common misunderstandings and areas of disagreement, and (3) synthesizes relevant literature to establish a common ground for future discussion and planning. It is important to understand the strengths and limitations of fuel treatments to evaluate their potential to achieve an objective, develop sensible fire management policies, and plan for their effective use. We suggest that, while the potential of fuel treatment to reduce wildfire occurrence or enhance suppression capability is uncertain, it has an important role in mitigating negative wildfire effects, increasing ecosystem resilience and making wildfire more acceptable.     

Goshawk breeding densities in relation to mature forest in southeastern Norway

Forest harvesting practices have been assumed to be the main reason for the decrease in Goshawk Accipiter gentilis populations in Fennoscandia during the late 1900s, by reducing prey density, especially grouse and Red Squirrel (Sciurus vulgaris), and preferred hunting habitats (mature forest). In Norway, the highest breeding densities are now found in areas with a high proportion of farmland and urban areas where corvids and pigeons are important prey. To assess the importance of mature forest for Goshawk populations in southeastern Norway, we compared breeding densities from 20 municipalities with different percentages of farmland and mature forest. In multiple regression models, regional breeding densities were positively related to both the percentage of mature forest, and the percentage of farmland and urban areas. We conclude that the recommendation of establishing buffer zones around nest sites will not be sufficient to maintain Goshawk populations if the proportion of mature forest is reduced at a landscape level.     

Habitat use of large ungulates in northeastern Germany in relation to forest management

Estimating large herbivore density has been a major area of research in recent decades. Previous studies monitoring ungulate density, however, focused mostly on determining animal abundance, and did not interpret animal distribution in relation to habitat parameters. We surveyed large ungulates in the Biodiversity Exploratory Schorfheide-Chorin using faecal pellet group counts. This allowed us to explore the link between relative ungulate abundance, habitat use, and browsing damage on trees in a region with several types of forest, including unharvested and age-class beech forests, as well as age-class pine forests. Our results demonstrate that roe deer and fallow deer relative abundance is negatively correlated with large tree cover, and positively correlated with the cover of small shrubs (Rubus spec., Vaccinium spec.), and winter food supply. Habitat use of roe deer and fallow deer, as estimated by counting faecal pellet groups, revealed a preference for mature pine forests, and avoidance of deciduous forests. This differential habitat use is explained by different distributions of high quality food resources during winter. The response of deer to understory cover differed between roe deer and fallow deer at high cover percentages. The amount of browsing damage we observed on coniferous trees was not consistent with the relative deer abundance. Browsing damage was consistently higher on most deciduous trees, except for beech saplings which sustained less damage when roe deer density was low. Because roe deer is a highly selective feeder, it was reported to affect tree diversity by feeding only on trees with high nutritional value. Consequently, we propose that managing the number of all deer species by hunting is necessary to allow successful forest regeneration. Such an adjustment to deer numbers would need to account for both current tree diversity and alternative food resources. Our findings may be applicable to other forest landscapes in northeastern Germany including mature pine stands and differently harvested deciduous forests.     

Selective logging in public pine forests of the central Iberian Peninsula: Effects of the recovery process on ant assemblages

Pine forests on the northern plateau of the Iberian Peninsula have a long history of use, exploitation, and management. Current management practices consist of selective logging with relatively short periods of time between logging events. The main objective of this study is to detect changes in ant assemblages in the short time periods between selective logging activities. Ants were sampled at 44 sites considering three grouping categories of time periods after the last timber extraction: short (<4 years), medium (4-8 years) and long (>8 years). After selective logging the number of ant species increases as the forest recovers. A look at the differences between the assemblages when the analysis shifts from the species-specific level to functional groups showed differences between the short and long categories. This indicates that in certain circumstances the functional groups may be more informative of the functional restructuring of the ant assemblages in a disturbed habitat. Ant species from three functional groups display significant indicative values (Opportunist, Hot Climate Specialists and Subordinate Camponotini) in the medium- and long-time-after-logging categories: Messor capitatus (HCS) for medium-time category sites; and Aphaenogaster iberica (Op), Camponotus cruentatus (SC) and, Cataglyphis velox (HCS) for long-time category sites. No indicator species were found for the early stages of recovery. This information may also be of interest to managers because it reduces the number of data elements of the recovery status of these forests, and can be translated into monitoring protocols. The continued exploitation of these forests leads to an ant fauna that reflects this change. The results show that ant assemblages need at least 8 years to recover since only after that much time there is an emergence of Subordinate Camponotinae, a behaviorally dominant and low stress-tolerant functional group. This selective logging maintains the relative diversity and structure of ant assemblages.     

Response of <Emphasis Type="Italic">Pinus pseudostrobus</Emphasis> Lindl. to fertile growing medium and tephra-layer depth under greenhouse conditions

Major disturbances that remove vegetation cover create conditions that differ greatly from those in adjacent forests. In volcanic areas, tephra deposition can be a major disturbance that eliminates vegetation cover and creates a barrier for plant establishment. The eruption of the Paricutín volcano, in Michoacán, México, from 1943 to 1952, created large tephra deposits adjacent to forest stands. Many of these deposits still lack native vegetation and understanding of the limiting factors for establishment of native tree species is necessary to restore these areas. Pinus pseudostrobus early growth in response to the amount of fertile growing medium on top of a variable-depth tephra layer was evaluated in a greenhouse experiment. Increasing amounts of fertile growing medium increased height of 28-week-old plants (no growing medium = 5.5 cm tall ±1.8 cm, 3 cm of growing medium = 21.2 ± 4.6 cm and 6 cm of growing medium 24.8 ± 4.4 cm). Other variables showed the same trend (diameter, number of fascicles, belowground biomass and aboveground biomass). Our results suggest that P. pseudostrobus establishment in tephra covered areas will depend on the accumulation of a fertile layer of more than 3 cm in thickness.     

Identifying cost-effective indicators to assess the conservation status of forested habitats in Natura 2000 sites

The Natura 2000 (N2000) network represents one of the most important actions for biodiversity conservation in Europe; its main aim is to assure the favourable conservation status of Europe's natural habitats and wild species. Monitoring of N2000 sites is required under the Habitats Directive, and Member States are required to report monitoring results to the European Commission. However, there is a widespread lack of understanding about precisely what should be monitored and which methods should be used. This research aimed to identify potential indicators and evaluate their suitability for evaluating the conservation status of forested habitats in N2000 sites. In addition, the cost-effectiveness of monitoring methods was assessed. Three monitoring methods were assessed in two N2000 sites, the Foresta del Cansiglio (IT) and the New Forest (UK): (i) sample plots, (ii) a point-transect method and (iii) a visual assessment method. Indicators were selected on the basis of a literature review, and related to forest structure and composition, dead wood volume, tree regeneration and ground flora composition. Results suggested that mean values of indicators did not differ between the plot and the point-transect-based methods (P > 0.05; paired t-test and Wilcoxon signed rank test). However, the survey method employed influenced the extent of variation within the indicators (as indicated by Z scores), a measure of their sensitivity. Correlations between indicators differed between the two sites. Of the three methods considered, the point-transect method was the most efficient in terms of set-up and total time required, but required the longest time per indicator and surveyed the smallest area (P < 0.001, Kruskal–Wallis test). The visual assessment method assessed the largest forest area; however, total scores of the reference values obtained from the plot-based and point-transect methods were poorly correlated with those obtained from this subjective approach. The contrasting results obtained between different methods and between different sites suggest that a general approach to monitoring N2000 sites may be difficult to develop; rather, a framework that can be adapted to the specific needs and characteristics of individual sites may be required.     

Simulating age-related changes in carbon storage and allocation in a Chinese fir plantation growing in southern China using the 3-PG model

Chinese fir [(Cunninghamia lanceolata (Lamb.) Hook (Taxodiaceae)] plantations are helping to meet China's increasing demands for timber, while, at the same time, sequestering carbon (C) above and belowground. The latter function is important as a means of slowing the rate that CO₂ is increasing in the atmosphere. Available data are limited, however, and even if extensive, would necessitate consideration of future changes in climatic conditions and management practices. To evaluate the contribution of Chinese fir plantations under a range of changing conditions a dynamic model is required. In this paper, we report successful outcome in parameterizing a process-based model (3-PG) and validating its predictions with recent and long-term field measurements acquired from different ages of Chinese fir plantations at the Huitong National Forest Ecosystem Research Station. Once parameterized, the model performed well when simulating leaf area index (LAI), net primary productivity (NPP), biomass of stems (W_S), foliage (W_F) and roots (W_R), litterfall, and shifts in allocation over a period of time. Although the model does not specifically include heterotrophic respiration, we made some attempts to estimate changes in root C storage and decomposition rates in the litterfall pool as well as in the total soil respiration. Total C stored in biomass increased rapidly, peaking at age 21 years in unthinned stands. The predicted averaged above and belowground NNP (13.81 t ha⁻¹ a⁻¹) of the Chinese fir plantations between the modeling period (from 4 to 21-year-old) is much higher than that of Chinese forests (4.8–6.22 t ha⁻¹ a⁻¹), indicating that Chinese fir is a suitable tree species to grow for timber while processing the potential to act as a C sequestration sink. Taking into account that maximum LAI occurs at the age of 15 years, intermediate thinning and nutrient supplements should, according to model predictions, further increase growth and C storage in Chinese fir stands. Predicted future increases (approximately 0–2 °C) in temperature due to global warming may increase plantation growth and reduce the time required to complete a rotation, but further increases (approximately 2–6 °C) may reduce the growth rate and prolong the rotational age.     

提高集体林权制度改革后森林经营管理水平的探讨

针对辽东山区集体林权制度改革后的实际情况,从建立林业合作经济组织、建立林业评估及风险防范体系、加大中幼林抚育财政补贴、提高森林抚育经营技术、进行采伐限额制度改革五个方面,详细阐述了提高集体林权制度改革后森林经营管理水平的几项措施,为林改后山区农民森林经营管理提供切实可行的指导意见。     

Bird community responses to a gradient of site preparation intensities in pine plantations in the Coastal Plain of North Carolina

Although intensively managed pine forests are common in the southeastern US, few studies describe how combinations of mechanical (MSP) and chemical site preparation (CSP) and herbaceous weed control (HWC) techniques affect bird communities that use early successional habitats within young pine forests. Therefore, we examined effects of six treatments of increasing management intensity via combinations of MSP (strip-shear and wide spacing or roller chop and narrow spacing) and CSP (application or no application) treatments with banded or broadcast HWC on bird communities in six loblolly pine (Pinus taeda) plantations in the Coastal Plain of North Carolina, USA, for 8 years following site preparation. Wide pine spacing and strip-shear MSP increased bird abundance and species richness over narrow spacing and chopped MSP for 6 years after planting. Chemical SP reduced bird abundance in year 2, increased bird abundance in year 6, had no effect on abundance after year 7, and did not affect species richness in any year. Total bird abundance and species richness were similar between banded and broadcast HWC. Site preparation and HWC had no effect on bird diversity and bird communities were most similar in treatments of similar intensity. Site preparation and HWC had few or no effects on birds based upon migratory status, habitat association, or conservation value. The addition of chemical site preparation or HWC had little effect on birds beyond pine spacing, and bird abundance was not proportional to management intensity. Although we observed treatment effects, all treatments provided habitat used by a variety of bird species, and pine plantations may play an increasingly important role in bird conservation as forests become fragmented and converted to other land uses and as natural processes that create early successional habitat, such as fire, are suppressed.     

Plant community responses to a gradient of site preparation intensities in pine plantations in the Coastal Plain of North Carolina

Intensively managed loblolly pine (Pinus taeda) forests are common in the southeastern United States and offer opportunities for conservation of biologic diversity. Within intensively managed landscapes, stand establishment relies on combinations of mechanical and chemical site preparation and herbaceous weed control (HWC) to manage competing vegetation and increase pine production. However, few long-term studies have described relationships between intensity of stand establishment and effects on plant communities. Therefore, we examined effects of 6 treatments that varied in intensity via combinations of mechanical (wide spacing and strip shear or narrow spacing and roller chop) and chemical (application or no application) site preparation treatments with HWC (broadcast or banded) from 1 to 8 years after site preparation on plant communities in loblolly pine plantations (n = 6) in the Coastal Plain of North Carolina, USA. All treatments resulted in abundant and diverse plant communities. Chemical site preparation had short lived (?4 year) effects on the herbaceous community but long-term effects on woody plants and pine cover. Increasing management intensity by including broadcast HWC or roller chop/narrow spacing did not additively reduce woody vegetation cover or species richness. However, broadcast HWC reduced grass, vine, and forb cover in the first year post-treatment. Average Morista community similarity values ranged from 0.69 to 0.89 among treatments and plots receiving the same chemical site preparation contained the most similar plant communities. Banded HWC can be paired with wide spacing to maximize herbaceous plant growth important for many wildlife species, particularly in the first few years after site preparation. Site preparation techniques should be tailored to local site conditions, plant communities, and management objectives.     

浅析云南森林植被恢复和保护技术

对云南省“天保”工程县 (局 )级实施方案编制中森林分类区划经营和管理技术进行分析 ,提出一些值得探讨的问题和建议 ,为云南森林植被恢复和保护及云南“天保”工程的顺利实施提供多方位的技术论证     

Root carbohydrates and aspen regeneration in relation to season of harvest and machine traffic

Season of harvest has often been suggested as a driver for the erratic success of aspen (Populus tremuloides) sucker regeneration, partially due to root carbohydrate reserves and soil conditions at the time of harvest. A field experiment in western Manitoba, Canada, assessed root suckering and root carbohydrates of aspen in response to season of harvest and machine traffic. Six sites (120 m × 120 m) were selected within two large mature aspen stands slated for summer harvest. Plots (50 m × 50 m) were hand-felled (without machine traffic) in mid-summer, late summer, winter, and one plot was left uncut as a control. Season of cut with no traffic had no effect on sucker density, height or leaf dry mass per sucker. During the dormant season, root starch reserves were highest in the winter cut plots, however, just prior to suckering, this difference in carbohydrate reserves among the three seasons of harvest disappeared and by the end of the first growing season root reserves in all three seasons of cut had recovered to near control levels. Adjacent plots that were conventionally harvested in the summer and impacted by logging traffic had similar sucker densities but had 19% less height growth of suckers and 29% less leaf dry mass per sucker compared to suckers in plots harvested at the same time without traffic. After one growing season, root carbohydrate levels were similar whether or not machine traffic was used; however, the reduction in leaf dry mass in plots with machine traffic could have negative implications for carbohydrate accumulation and growth. The study suggests that the phenological state of the mature aspen plays a very small role in aspen regeneration and that harvesting practices and site conditions are likely the main drivers of aspen regeneration success.     

Soil nitrogen status of larch plantations in comparison with secondary broad-leaved forest

IntroductionSoildegradationofsouthernChinesefir(CunnhaamialanceolaIe)pIantationshavebeenCon-tinuaIlyreiorted(ShenWeitongl992;Che11BingI1aol992;FangQil992).lnthenortI1eastforestregionsofChina,larch(LarirohensisandLa)`ixgInelinii)planta-tionsmade7opercentoftheman-madeforests,andthechangesinsoilpropertiesundertI1econiferouspIanta-tionshavealsodrawnmuchattentionofmanyforesters.OurobjectivewastoexaminethedifferenceinsoilnitrogenstatusbetweenLarchp1antatiol1sandsecon-darybroad-Ieavedforests(…