Post-fire tree mortality in mixed forests of central Portugal

Wildfires are a recurrent disturbance in the Mediterranean Basin. However, managers from this region are confronted with a lack of information on the effects of fire on most woody species, which is required for defining sustainable forest management strategies. Following a large wildfire in central Portugal (2003), we surveyed the area during the first year and assessed the vegetative condition of 1040 burned trees from 11 different species. Among those trees, 755 individuals were selected and monitored annually for 4 years. At the end of the study, almost all the broadleaved trees survived, while most coniferous died. In spite of the low mortality observed in broadleaves, most were top-killed and regenerated only from basal resprouts, which implies a slow recovering process. Quercus suber, however, showed vigorous post-fire crown resprouting and was the most resilient species. We fitted logistic regression models to predict the probability of individual tree mortality and top-kill from fire injury indicators and tree characteristics. Besides the differences between the two main functional groups (coniferous, broadleaved), bole char height and crown volume scorched or consumed were important predictors of tree responses. Additionally, the main factor determining crown mortality on broadleaved species was bark thickness. The selected models performed well when tested with independent data obtained on four other wildfires. These models have several potential applications and can be useful to managers making pre-fire or post-fire decisions in mixed forest stands in the western Mediterranean Basin.     

Fire cycle and fire adaptation of main tree species in Daxing’anling forest region,China

Based on the fire statistics, the Daxing’anling forest area were classified into three fire cycle regions: northern coniferous virgin forest region with a fire cycle ofl 10–120 years, middle conifer-broad-leaved mixed forest region with a fire cycle of 30–40 years, and southern broad-leaved secondary forest region with a fire cycle of 15–20. The percentage of conifers and broad-leaved trees, forest age and natural mature period of main tree species in different fire cycle regions were discussed in concern with fire occurrence. The characteristics of fire adaptation and fire resistance of main tree species, such as sexual and asexual reproduction, were discussed and evaluation of the synthetical fire adaptation was made.     

Competition-density effect of tree organs in Acacia auriculiformis stands

The competition-density (C-D) effects for mean mass for tree, stem, branch and leaf were analyzed in Acacia auriculiformis stands. Mean tree mass-density and mean organ mass-density were well explained by the C-D equation of tree and the C-D equation of tree organ, respectively. An equation describing the relationship between mean leaf area u and density was formulated that fit the u-data well. The relationship between mean tree mass w and the ratio of each organ to mean tree mass (wo/ w) was examined. With increasing w, the stem mass ratio wS/w increased, whereas the branch mass ratio wB/w and the leaf mass ratio wL/w decreased. The yield difference between the lowest-density stand and the high-density stand became greater with stand growth. However, the yield of the mid-density stand was slightly lower than the yield of the high-density stand during the experimental period. To produce the most desirable combination of demanding individual-tree size and relative high stem yield, the mid-density is recommended as proper planting density for future management of A. auriculiformis stands.     

南方型杨树人工林直径和断面积预估模型的研究

对南方型杨树的主栽无性系I-63杨、I-69杨、I-72杨的直径生长过程进行了差异性检验,结果3个无性系的平均相对误差分别为0.34%、-3.89%和4.23%,直径生长过程表现出良好的相似性。采用不分无性系的混合建模方法建立了南方型杨树人工林断面积直径预估模型-Dg=0.968 1H01.361 4N-0.032H00.540 2,从而得到林分断面积预估模型G=0.000 073 607N02.722 8N1-0.064H00.540 2。     

Impacts of thinning on structure,growth and risk of crown fire in a Pinus sylvestris L. plantation in northern Spain

We studied the combined effects of thinning on stand structure, growth, and fire risk for a Scots pine thinning trial in northern Spain 4 years following treatment. The thinning treatments were: no thinning, heavy thinning (32–46% of basal area removed) and very heavy thinning (51–57% of basal area removed). Thinning was achieved via a combination of systematic and selective methods by removing every seventh row of trees and then by cutting suppressed and subdominant trees in the remaining rows (i.e., thinning from below). Four years after thinning, mean values and probability density distributions of stand structural indices showed that the heavier the thinning, the stronger the tendency towards random tree spatial positions. Height and diameter differentiation were initially low for these plantations and decreased after the 4-year period in both control and thinned plots. Mark variograms indicated low spatial autocorrelation in tree diameters at short distances. Diameter increment was significantly correlated with the inter-tree competition indices, and also with the mean directional stand structural index. Two mixed models were proposed for estimating diameter increment using a spatial index based on basal area of larger trees (BALMOD) in one model versus spatial competition index by Bella in the other model. As well, a model to estimate canopy bulk density (CBD) was developed, as this variable is important for fire risk assessment. Both heavy and very heavy thinning resulted in a decrease of crown fire risk over no thinning, because of the reduction in CBD. However, thinning had no effect on the height to crown base and thus on the flame length for torching. Overall, although thinning did not increase size differentiation between trees in the short term, the increase in diameter increment following thinning and the reduction of crown fire risks support the use of thinning. Also, thinning is a necessary first step towards converting Scots pine plantations to more natural mixed broadleaved woodlands. In particular, the very heavy thinning treatment could be considered a first step towards conversion of overstocked stands.     

福州地区人工杉木林分蓄积量表的研究

本文提出以林分构成因子——林分胸高总断面积及林分平均高，估计林分蓄积量的办法，选取９种方程求解参数，配合了最优林分蓄积量经验方程，制定出《福州地区人工杉木林分蓄积量表》，对完善我省林业数表的种类、形式和内容，进行了初步尝试。     

Natural mortality in thinning and fertilisation experiments with pine and spruce in Sweden

Natural mortality in a 30-year period was examined in thinning and fertilisation experiments with 48 blocks in Scots pine (Pinus sylvstris L.) and 23 blocks in Norway spruce (Picea abies (L.) Karst.) with up to 12 different treatments. Of about 90,000 living trees at start of the experiments 18.7% were registered as dead by natural mortality in the 30-year observation period. In non-thinned stands about 20% of the volume growth disappeared by natural mortality, in thinned stands about 10%. In normally thinned pine stands (repeated thinning from below with moderate intensity) the annual mortality of the basal area at start of an average 7-year period was 0.34%. In spruce stands, on more fertile sites, the corresponding figure was about 0.6%. In an effort to model the mortality, severe damage not leading to final felling was identified in 1.7% of the observation periods. It was assumed that this part of the mortality, representing 24% of the total volume mortality, could be recovered by active thinning. The probability for severe damage increased sharply with stand top height, as shown in a logistic regression. The more sparse mortality was expressed as a function of site fertility, stand density, disturbance by thinning and form of treatment (thinned from above or below or non-thinned). The naturally dead trees were approximately of mean size in normally thinned stands while the self-thinning in non-thinned stands tended to occur amongst smaller than average trees. Diagrams were presented for basal area development and stem number reduction in the non-thinned stands.     

Integration of LiDAR and QuickBird imagery for mapping riparian biophysical parameters and land cover types in Australian tropical savannas

Riparian zones are exposed to increasing pressures because of disturbance from agricultural and urban expansion and overgrazing. Accurate and cost-effective mapping of riparian environments is important for baseline inventories and monitoring and managing their functions associated with water quality, biodiversity, and wildlife habitats. In this study, we integrate remotely sensed light detection and ranging (LiDAR) data and high spatial resolution satellite imagery (QuickBird-2) to estimate riparian biophysical parameters and land cover types in the Fitzroy catchment in Queensland, Australia. An object based image analysis (OBIA) was adopted for the study. A digital terrain model (DTM), a tree canopy model (TCM) and a plant projective cover (PPC) map were first derived from the LiDAR data. A map of the streambed was then produced using the DTM information. Finally, all the LiDAR-derived biophysical map products and the QuickBird image bands were combined in an OBIA to (1) map the following land cover types: riparian vegetation, streambed, bare ground, woodlands and rangelands; (2) determine the distribution of overhang vegetation within the streambed; and (3) measure the width of both the riparian zone and the streambed. The combined use of both datasets allowed accurate land cover mapping, with an overall accuracy of 85.6%. The estimated widths of the riparian zone and the streambed showed strong correlation with the actual field measurements (r = 0.82 and 0.98 respectively). Our results show that the combined use of LiDAR and high spatial resolution imagery can potentially be used for the assessment of the riparian condition in a tropical savanna woodland riparian environment. This work also shows the capacity of OBIA to assist in the assessment of the composition of the riparian environment from multiple image datasets.     

Prescribed fire effects on bark beetle activity and tree mortality in southwestern ponderosa pine forests

Prescribed fire is an important tool in the management of ponderosa pine (Pinus ponderosa Dougl. ex Laws.) forests, yet effects on bark beetle (Coleoptera: Curculionidae, Scolytinae) activity and tree mortality are poorly understood in the southwestern U.S. We compared bark beetle attacks and tree mortality between paired prescribed-burned and unburned stands at each of four sites in Arizona and New Mexico for three growing seasons after burning (2004–2006). Prescribed burns increased bark beetle attacks on ponderosa pine over the first three post-fire years from 1.5 to 13% of all trees, increased successful, lethal attacks on ponderosa pine from 0.4 to 7.6%, increased mortality of ponderosa pine from all causes from 0.6 to 8.4%, and increased mortality of all tree species with diameter at breast height >13 cm from 0.6 to 9.6%. On a per year basis, prescribed burns increased ponderosa pine mortality from 0.2% per year in unburned stands to 2.8% per year in burned stands. Mortality of ponderosa pine 3 years after burning was best described by a logistic regression model with total crown damage (crown scorch + crown consumption) and bark beetle attack rating (no, partial, or mass attack by bark beetles) as independent variables. Attacks by Dendroctonus spp. did not differ significantly over bole heights, whereas attacks by Ips spp. were greater on the upper bole compared with the lower bole. Three previously published logistic regression models of tree mortality, developed from fires in 1995–1996 in northern Arizona, were moderately successful in predicting broad patterns of tree mortality in our data. The influence of bark beetle attack rating on tree mortality was stronger for our data than for data from the 1995–1996 fires. Our results highlight canopy damage from fire as a strong and consistent predictor of post-fire mortality of ponderosa pine, and bark beetle attacks and bole char rating as less consistent predictors because of temporal variability in their relationship to mortality. The small increase in tree mortality and bark beetle attacks caused by prescribed burning should be acceptable to many forest managers and the public given the resulting reduction in surface fuel and risk of severe wildfire.     

Effects of species and tree size diversity on recruitment in the Alaskan boreal forest: A geospatial approach

This study empirically evaluates and maps the relationships between recruitment and species and tree size diversity, as measured with the Shannon’s index, within mixed poplar/birch and mixed spruce stands across the boreal forest of Alaska. Data were collected from 438 permanent sample plots re-measured at a 5-year interval. Significant explanatory factors of recruitment, including species and tree size diversity were first identified using hierarchical partitioning. The effects of tree diversity on recruitment were then studied using generalized linear models and universal kriging to account for non-spatial factors and for spatial autocorrelation. We found a consistent positive relationship between recruitment and species diversity and a general negative relationship between recruitment and tree size diversity, indicating a tradeoff between species diversity and tree size diversity in affecting recruitment. These relationships however were not uniform across the landscape, presumably because they were subject to strong spatial autocorrelation attributable to natural disturbances and environmental stressors. In general, diversity had least effect on recruitment in stressful environments where stress, rather than competition, most likely governed recruitment.     

不同强度林火干扰对红花尔基樟子松天然林更新的影响

在依据火烧迹地林木烧伤程度确定林火强度基础上,对内蒙古红花尔基樟子松林国家级自然保护区不同林火强度下的幼苗更新状况进行调查和分析,以期探明自然条件下林火干扰对樟子松天然林更新的影响,为今后开展林火促进更新方面提供科学依据。结果表明:不同火烧强度下更新树种均为樟子松;樟子松天然更新株数介于2 500~6 500株·hm-2,平均值为4 542株·hm-2;樟子松天然更新频度介于50%~100%,平均值为89%;表明林火能够促进樟子松天然更新。随着火烧程度增加,樟子松天然更新株数和频度显著降低,轻度和中度火烧更有利于促进樟子松天然更新,而重度和极重度火烧下樟子松仍能天然更新。     

The impact of fire and density-dependent mortality on the spatial patterns of a pine forest in the Hulun Buir sandland,Inner Mongolia,China

Wildland fire, especially surface fire, is one of the major disturbances in forest ecosystems such as the Mongolian pine forest of the Hulun Buir sandland. However, little is known about the impact of fire on the spatial patterns at the stand level and its consequences for successional dynamics. To fill this gap we mapped three plots of Mongolian pine forest comprising sites of the same age that were affected by fire in 2006 and 1994, and a site without fire. We explored the stand structure using diameter at breast height (DBH) distributions and basal area, and used point pattern analysis to quantify the observed spatial patterns. Null models included homogeneous Poisson and Thomas cluster processes for univariate analyses, toroidal shift to test for independence, and random labelling for analyzing mortality. Large stems showed at all three stands a tendency to regularity, whereas small stems were mostly clustered. However, small stems did not become more regular at plots with earlier fire, and only the “overstocked” 2006 pre-fire stand showed negative association between large and small trees. Mortality was non-random, showing clustering of dead (or surviving) trees and clear density dependence where stems with more neighbors had a higher risk to die. Fire killed mostly smaller trees and after prolonged fire-free periods the stand approached the pre-fire patterns structurally and spatially. Our study showed that surface fire causes strong thinning in later succession stages and pushes the forest into maturity. It may also enhance resistance of our forest to more severe fire under the relatively harsh environmental conditions. A novel test statistic for random labeling allowed directly testing for density-dependent mortality and could be widely applied in point pattern analysis.     

Spatial variability in stand structure and density-dependent mortality in newly established post-fire stands of a California closed-cone pine forest

Fire is an important process in California closed-cone pine forests; however spatial variability in post-fire stand dynamics of these forests is poorly understood. The 1995 Vision Fire in Point Reyes National Seashore burned over 5000 ha, initiating vigorous Pinus muricata (bishop pine) regeneration in areas that were forested prior to the fire but also serving as a catalyst for forest expansion into other locales. We examined the post-fire stand structure of P. muricata forest 14 years after fire in newly established stands where the forest has expanded across the burn landscape to determine the important factors driving variability in density, basal area, tree size, and mortality. Additionally, we estimated the self-thinning line at this point in stand development and compared the size-density relationship in this forest to the theorized (−1.605) log-log slope of Reineke’s Rule, which relates maximum stand density to average tree size. Following the fire, post-fire P. muricata density in the expanded forest ranged from 500 to 8900 live stems ha⁻¹ (median density = 1800 ha⁻¹). Post-fire tree density and basal area declined with increasing distance to individual pre-fire trees, but showed little variation with other environmental covariates. Self-thinning (density-dependent mortality) was observed in nearly all stands with post-fire density >1800 stems ha⁻¹, and post-fire P. muricata stands conformed to the size-density relationship predicted by Reineke’s Rule. This study demonstrates broad spatial variability in forest development following stand-replacing fires in California closed-cone pine forests, and highlights the importance of isolated pre-fire trees as drivers of stand establishment and development in serotinous conifers.     

The effects of thinning and burning treatments on within-canopy variation of leaf traits in hardwood forests of southern Ohio

Leaf nitrogen content (N_mass, %) and leaf mass per area (LMA, g m⁻²) are two important features that are closely linked to the photosynthetic performance of plants and, thus, the NPP of forest ecosystems. Forest management practices, such as burning and thinning, change stand structure and soil dynamics, which may result in changes in N_mass and LMA. The objective of this study was to understand how N_mass and LMA of seven canopy tree species/genus (Quercus alba, Q. coccinea, Q. prinus, Q. velutina, Carya spp., Acer rubrum, and Liriodendron tulipifera) responded to (i) thinning and/or burning treatments and to (ii) different landscape and soil properties in southern Ohio. We collected leaves from the top, and bottom, of five individuals of each taxa in each treatment unit. Leave traits (N_mass and LMA) were compared using analysis of variance followed by orthogonal contrasts. To further understand the factors that influence the canopy leaf traits, we used regression tree analysis (RTA) to examine how variations of LMA and N_mass were linked to thinning and/or burning treatments, soil, and landscape variables. Finally, we assessed the potential ramifications of changes in these traits on canopy carbon budgets using a PnET-Day model, which is a daily time-step canopy carbon exchange model. We found significant effects of thinning, burning, and their interactions on LMA at the bottom of the crown while none of the treatments showed significant effects on LMA at the top of the crown. N_mass responded significantly to only burning treatment. RTA results exhibited minor effects of landscape features and soil properties on N_mass and LMA. Interspecific differences accounted for most variations of both leaf traits. Sensitivity analysis of PnET-Day model suggested these subcanopy changes in LMA increased the annual net primary production (NPP) by 8%. In summary, our results suggest that forest management can substantially influence canopy leaf traits such as N_mass and LMA and that alteration of these traits can influence forest NPP. Given the role of forests as global carbon sinks, the potential influence of thinning and burning on canopy traits, and thus NPP, is an important consideration for forest management.     

Effects of prescribed fire and season of burn on direct and indirect levels of tree mortality in Ponderosa and Jeffrey Pine Forests in California,USA

Many forests that historically experienced frequent low-intensity wildfires have undergone extensive alterations during the past century. Prescribed fire is now commonly used to restore these fire-adapted forest ecosystems. In this study, we examined the influence of prescribed burn season on levels of tree mortality attributed to prescribed fire effects (direct mortality) and bark beetles (Coleoptera: Curculionidae, Scolytinae) (indirect mortality) in ponderosa pine, Pinusponderosa Dougl. ex Laws., and Jeffrey pine, Pinusjeffreyi Grev. and Balf., forests in California, USA. A total of 816 trees (9.9% of all trees) died during this 3-yr study. Significantly higher levels of tree mortality (all sources) occurred following early and late season burns compared to the untreated control, but no significant difference was observed between burn treatments. The majority (461 trees) of tree deaths were attributed to direct mortality from prescribed burns and was strongly concentrated (391 trees) in the smallest diameter class (<20.2 cm diameter at breast height, dbh). For the largest trees (>50.7 cm dbh), significantly higher levels of tree mortality occurred on early season burns than the untreated control, most of which resulted from indirect mortality attributed to bark beetle attacks, specifically western pine beetle, Dendroctonus brevicomis LeConte, and mountain pine beetle, D. ponderosae Hopkins. Red turpentine beetle, D. valens LeConte, was the most common bark beetle species found colonizing trees, but tree mortality was not attributed to this species. A total of 355 trees (4.3% of all trees) were killed by bark beetles. Dendroctonus brevicomis (67 trees, 18.9%) and D. ponderosae (56 trees, 15.8%), were found colonizing P. ponderosa; and Jeffrey pine beetle, D. jeffreyi Hopkins, was found colonizing P. jeffreyi (seven trees, 2.0%). We also found pine engraver, Ips pini (Say) (137 trees, 38.6%), and, to a much lesser extent, Orthotomicus (=Ips) latidens (LeConte) (85 trees, 23.9%) and emarginate ips, I. emarginatus (LeConte) (3 trees, 0.8%) colonizing P. ponderosa and P. jeffreyi. Few meaningful differences in levels of indirect tree mortality attributed to bark beetle attack were observed between early and late season burns. The incidence of root and root collar pathogens (Leptographium and Sporothrix spp.), including species known to be vectored by bark beetles, was low (18% of trees sampled). The implications of these and other results to management of P. ponderosa and P. jeffreyi forests are discussed in detail.     

冀北山地油松、蒙古栎林土壤水分—物理性质研究

土壤是森林生态系统的组成成分和物质基础。以冀北山地油松林、蒙古栎林、油松蒙古栎混交林作为研究对象,分析和比较了林下土壤水分—物理性质,结果表明:土壤容重随土层深度的增加而增加,由大到小排序为:油松林(1.30g/cm~3)、蒙古栎林(1.14g/cm~3)、油松蒙古栎混交林(1.10g/cm~3);土壤总孔隙度由大到小排序为:蒙古栎林(48.04%)、油松蒙古栎混交林(42.58%)、油松林(33.58%);土壤最大持水量由大到小排序为:蒙古栎林(40.32%)、油松蒙古栎混交林(38.81%)、油松林(33.58%);地表土壤稳渗速率由大到小排序为:油松林、油松蒙古栎混交林、蒙古栎林,土壤入渗速率与入渗时间存在明显的幂函数关系。     

卫星林火监测在西南地区应用状况及存在问题

西南地区卫星林火监测分中心于1997年投入运行,对林火监测具有连续性、高时效性和监测范围广等特性,与森林航空监测方式结合,可有效地提高飞行效益．分析现在还存在的系统建设不足、人员素质不高,定位精度不高,地面信息反馈不及时等问题,并相应地提出了解决问题的对策和建议．     

Predicting Potential Density and Basal Area Development in the Social Forestry Plantations of Acacia nilotica in Gujarat State of India

ABSTRACT

Growth modelling is an important and effective tool for evaluating the effects of a particular management action on the future development of a forest ecosystem. However, such necessary growth models are not available for many indigenous tree species in India. Acacia nilotica is an important multipurpose tree species found in India and growth models are required for proper management of the species in the region. This paper presents equations for estimating potential stand density and predicting basal area in pure even-aged stands of A. nilotica in Gujarat State of India. Although no thinning was suggested, decrease in the number of trees in the stands was observed because of mortality due to overcrowding and some biotic factors. Relationships between quadratic mean diameter and stems per hectare were developed, which was used to establish the limiting density line. Eight different stand level models, belonging to the path invariant algebraic difference form of a non-linear growth function, were compared for projecting basal area. They can be used to predict future basal area as a function of stand variables like dominant height and stem number per hectare and are crucial for evaluating different silvicultural treatment options. The performance of the models was evaluated using different statistical criteria to recommend the suitable model for projecting the basal area in A. nilotica stands.     

杉松人工林伐区调查设计中树高曲线图的绘制与应用

为提高罗源县杉、松人工林树高曲线图绘制的准确性,利用广泛拟合择优选取的方法确定了能很好描述罗源县杉、松人工林胸径(D)树高(H)的一元二次函数模型,同时利用Excel电子表格设计绘制树高曲线图的程序。研究结果具有较大的应用价值。     

大岗山林区水体、空气、噪声环境背景值研究*

研究表明大岗山林区水体的微量元素环境背景值均不超过世界天然水的环境背景值，水理化性质及微量元素含量均达国内外规定的生活饮用水水质标准；ＳＯ２、ＮＯｘ、总悬浮颗粒物含量均达到《中国大气环境质量标准》一级标准；噪声也不超过国际国内标准，整个林区环境质量优良，生物与环境的生态关系协调并相互适应。