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1.
墨西哥柏幼苗生长和光合生理对氮沉降的响应   总被引:2,自引:0,他引:2  
选取2年生墨西哥柏幼苗为试验对象,以NH4NO3为外加氮源,通过模拟氮沉降试验,研究其生长和光合生理对氮增加的响应.结果表明:幼苗在高氮水平处理下其生长不断受到抑制,而在中氮水平处理下,幼苗生长不断得到促进.随着施氮浓度增加,墨西哥柏幼苗的根质量和根长均降低,表明氮处理抑制根的生长,分配到根部分的生物量下降.幼苗的净光合速率呈现出随氮处理浓度增加而先增加后降低的趋势,即中氮组的净光合速率最高,而高氮组开始逐渐下降.幼苗的水分利用效率变化趋势与净光合速率一致.幼苗叶片光合色素含量随氮处理水平增加而增加.说明短期内中氮水平对幼苗生长和光合作用影响最显著,而高氮水平对幼苗光合色素影响最显著.  相似文献   

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3.
To determine the effects of irrigation and coppicing on above- and below-ground growth dynamics, a plantation of Populus × euramericana cv. ‘Eugenei’ and Populus tristis × Populus balsamifera cv. ‘Tristis 1’ was established in May 1984 on a level site in East Lansing, MI, USA. Supplemental water in the form of drip irrigation was applied to half the trees beginning the first growing season. All trees were cut down in March 1988 and the stumps allowed to coppice. Pre- and post-coppice height and diameter growth of ‘Eugenei’ always exceeded that of ‘Tristis’, and the former clone showed a positive response to irrigation, whereas the latter did not. The greater growth of ‘Eugenei’ was primarily due to its full use of the growing season. Post-coppice rates of photosynthesis were not affected by irrigation in either clone, but stomatal conductances were reduced in non-irrigated trees. Analysis of microvideo images taken in minirhizotron tubes buried in the soil close to the trees showed that ‘Tristis’ produced a greater length and number of fine roots in the top 30 cm of soil than ‘Eugenei’, regardless of treatment. Irrigated trees consistently produced more fine roots at 0–30 cm soil depth than non-irrigated trees only in ‘Eugenei’, but both clones showed greater fine-root production in non-irrigated trees at 30–100 cm. Both clones also showed substantial fine-root production in the spring immediately following coppicing, with no evidence of a shock-induced dieback of roots. The root systems of these two poplar genotypes apparently contain sufficient carbon and nitrogen reserves to fuel a spring flush of fine-root growth, even though the tops have been severed during the dormant season.  相似文献   

4.
文章研究了土壤不同水分条件下200、400kg/(株.a),不同氮沉降0、45和135g/(株.a)对文冠果结实(包括果实产量、种子千粒重、种仁含油率、果实纵横径及其比值等)的影响。结果表明:文冠果结实具有明显的变化特征。氮沉降在一定程度上提高了果实产量和果实横径。氮沉降对种子千粒重、种仁含油率的影响与土壤水分条件有关。低水分条件下,前者随着氮沉降的增加而增加,后者变化不明显。高水分条件下,前者变化不明显,后者则升高。2种水分条件下,果实的纵径、纵横径比均随着氮沉降的增加而降低。  相似文献   

5.
Biomass, leaf area, canopy photosynthesis, photosynthetic nitrogen-use efficiency (PNUE), nitrogen-partitioning ratio (NPR: ratio of nitrogen taken up by jack pine relative to two different competitor species), and nitrogen uptake (NU) of jack pine (Pinus banksiana Lamb.) competing with large-leaved aster (Aster macrophyllus L.) and Canada blue-joint grass (Calamagrostis canadensis (Michx.) Beauv.) were examined at three nitrogen levels in a controlled-environment growth chamber. When grown with large-leaved aster, jack pine biomass, photosynthesis and PNUE (p<0.001) increased as nitrogen level increased. Jack pine biomass, photosynthesis and NPR (p<0.001) decreased as nitrogen level increased when grown with Canada blue-joint grass. At the lowest nitrogen supply level, jack pine photosynthesis decreased as competitor PNUE increased (r2=0.84, p<0.001). Jack pine photosynthesis decreased as NU of large-leaved aster (37.5 mg N l−1: r2=0.75, p<0.001; 100 mg N l−1: r2=0.86, p<0.001) and Canada blue-joint grass (37.5 mg N l−1: r2=0.96, p<0.001; 100 mg N l−1: r2=0.84, p<0.001) increased. NU and PNUE may play an important role in the outcome of interactions between jack pine seedlings and competing forest vegetation in newly planted stands.  相似文献   

6.
We compared shade tolerance of maple, ash and beech in the saplingstage from two sites with rich soils differing in water supply,growing in dense thickets underneath a beech shelterwood ofvarying canopy densities. Shade tolerance was described by twocomponents: mortality in shade and height growth in high light.At low light, beech showed the least mortality, maple the highestand ash in between on both sites. The decline with increasinglight was steepest in beech and more gradual with ash and maple.At 15 per cent above canopy light, all three species approachedzero mortality. Beech as the most shade-tolerant species hadthe highest survival rate under low light and the least lengthgrowth rate under high light (>17 per cent). Ash had a lowersurvival rate at low light than beech and a highest growth rateat high light. Maple showed a bit weaker trade-off with thelowest survival rate but a growth rate inferior to ash. On thebetter water-supplied site, height growth was significantlysuperior in all three species only under high light. On thebasis of these results, silvicultural conclusions are drawnwith respect to appropriate light levels and cutting types.  相似文献   

7.
Samuelson  L.J. 《New Forests》2000,19(1):95-107
Leaf physiology and fractional dry weight allocation were examined in four open-pollinated families of loblolly pine (Pinus taeda L.) and slash pine (Pinus elliottii Englm. var. elliottii) in response to growth under low and high N supply for six months. Nitrogen greatly influenced dry weight allocation, total mass, leaf net photosynthesis and leaf conductance in seedlings of both species. Family variation in fine root allocation was observed under low but not high N treatment, but for the majority of physiological and growth traits, family variation was stable under varying N treatment. Family rankings based on juvenile height, diameter and biomass accretion were similar to rankings based on field performance at 22 years in slash pine but not loblolly pine. Lower leaf maintenance respiration rates were associated with families exhibiting the most rapid juvenile growth.  相似文献   

8.
The northern and central Appalachian forests are subject to high levels of atmospheric acid deposition (AD), which has been shown in some forests to negatively impact forest growth as well as predispose the forest system to damage from secondary stresses. The purpose of this study was to evaluate the possible contribution of AD to changes in composition and productivity of the Monongahela National Forest, and to evaluate soil-based indicators of acidification that might be useful for detecting AD-related forest changes. Soils adjacent to 30 Forest Inventory and Analysis (FIA) sites were sampled and analyzed for a suite of acidity indicators. These indicators were correlated with the periodic mean annual volume increment (PMAVI) of the forest stands on FIA plots for the 10-yr period 1989–2000. PMAVI ranged from −9.5 to 11.8 m3 ha−1 yr−1, with lower-than-expected growth (<3 m3 ha−1 yr−1) on two-thirds of the sites. In the surface horizon, effective base saturation, Ca2+ concentration, base saturation, K+ concentration, Ca/Al molar ratio, and Mg/Al molar ratio, were positively correlated with PMAVI and Fe concentration was negatively correlated with PMAVI (p ≤ 0.1). In the subsurface horizon pH(w) and effective base saturation were positively correlated and Al3− concentration and K+ concentration were negatively correlated with PMAVI. We hypothesized that NO3-N/NH4-N ratio would also be correlated with PMAVI, but it was not. Correlations between soil chemical indicators and PMAVI suggest that AD may contribute, in part, to the lower-than-expected forest growth on the Monongahela National Forest.  相似文献   

9.
After many years of increasing nitrogen deposition, the deposition rates are now decreasing. A major question is whether this will result in the expected positive effects on plant species diversity. Long-term experiments that investigate the effects of decreasing deposition are not available. Model simulations may yield insight into the possible effects of decreasing nitrogen deposition on the vegetation. Therefore we developed the vegetation succession model SUMO which is closely linked to the soil model SMART2. In SUMO, the biomass development of five functional plant types is simulated as a function of nitrogen availability, light interception and management. The model simulates the change in biomass distribution over functional types during the succession from almost bare soil via grassland or heathland to various forest types.The model was validated on three sites in The Netherlands and one site in the UK. The aboveground biomass of two grassland vegetation types was well simulated, as well as the aboveground biomass of heathlands during succession of sod removal. Some of the stages of forest succession were simulated less well, but the calculated biomass in the older stages agreed with measured values.To explore the long-term effect of a decrease in nitrogen deposition, we applied the model to a heathland and a pine stand. In the heathland a major change was predicted as a result of decreasing nitrogen deposition in combination with turf stripping. The dominance of grasses changed into a dominance of dwarf shrubs, whereas at continuing high levels of nitrogen deposition grasses remained dominant. In contrast, the simulations indicated only very small effects of a decreasing N deposition in pine forests. This difference is due to the removal of excess nitrogen by management (turf stripping) in the heathland, whereas the more extensive management in the forest hardly removes any nitrogen from the system. The main conclusion from these examples is that a decrease of nitrogen deposition may retard succession, and consequently increase biodiversity in heathland but probably not in forest. The effects of declining N deposition depend on the amount of N that is removed from the system as a consequence of the various management regimes.  相似文献   

10.
Sanding is a common practice required in order to prepare wood surfaces for coating. Little literature is available regarding the effect of sanding parameters on the quality of surfaces. Sugar maple wood surfaces were evaluated in samples that had been sanded using two types of abrasive minerals, three grit sizes and four feed speeds. Roughness, wetting properties and cell damage were used to assess surface quality. Both abrasives decreased roughness and cell damage from 100- to 120-grit sanding stage. Addition of a 150-grit stage did not further reduce the roughness, whereas the cell damage continued to decrease. Increasing feed speed led to rougher surfaces due to higher fibrillation. Surfaces produced by silicon carbide were smoother and less damaged than those obtained with aluminum oxide. However, the surfaces sanded with aluminum oxide were more wettable and showed no significant difference in wetting time as a function of grit size. For these surfaces, the wetting time was reduced as feed speed increased.An erratum to this article is available at .An erratum to this article can be found at  相似文献   

11.
论述了光合作用与光能利用率的关系,提出经营密度的控制范围。  相似文献   

12.
Nitrogen-fixing plant species may respond more positively to elevated atmospheric carbon dioxide concentrations ([CO2]) than other species because of their ability to maintain a high internal nutrient supply. A key factor in the growth response of trees to elevated [CO2] is the availability of nitrogen, although how elevated [CO2] influences the rate of N2-fixation of nodulated trees growing under field conditions is unclear. To elucidate this relationship, we measured total biomass, relative growth rate, net assimilation rate (NAR), leaf area and net photosynthetic rate of N2-fixing Alnus glutinosa (L.) Gaertn. (common alder) trees grown for 3 years in open-top chambers in the presence of either ambient or elevated atmospheric [CO2] and two soil N regimes: full nutrient solution or no fertilizer. Nitrogen fixation by Frankia spp. in the root nodules of unfertilized trees was assessed by the acetylene reduction method. We hypothesized that unfertilized trees would show similar positive growth and physiological responses to elevated [CO2] as the fertilized trees. Growth in elevated [CO2] stimulated (relative) net photosynthesis and (absolute) total biomass accumulation. Relative total biomass increased, and leaf nitrogen remained stable, only during the first year of the experiment. Toward the end of the experiment, signs of photosynthetic acclimation occurred, i.e., down-regulation of the photosynthetic apparatus. Relative growth rate was not significantly affected by elevated [CO2] because although NAR was increased, the effect on relative growth rate was negated by a reduction in leaf area ratio. Neither leaf area nor leaf P concentration was affected by growth in elevated [CO2]. Nodule mass increased on roots of unfertilized trees exposed to elevated [CO2] compared with fertilized trees exposed to ambient [CO2]. There was also a biologically significant, although not statistically significant, stimulation of nitrogenase activity in nodules exposed to elevated [CO2]. Root nodules of trees exposed to elevated [CO2] were smaller and more evenly spaced than root nodules of trees exposed to ambient [CO2]. The lack of an interaction between nutrient and [CO2] effects on growth, biomass and photosynthesis indicates that the unfertilized trees maintained similar CO2-induced growth and photosynthetic enhancements as the fertilized trees. This implies that alder trees growing in natural conditions, which are often limited by soil N availability, should nevertheless benefit from increasing atmospheric [CO2].  相似文献   

13.
Four years of severe drought from 1999 through 2003 led to unprecedented bark beetle activity in ponderosa and Jeffrey pine in the San Bernardino and San Jacinto Mountains of southern California. Pines in the San Bernardino Mountains also were heavily impacted by ozone and nitrogenous pollutants originating from urban and agricultural areas in the Los Angeles basin. We studied bark beetle activity and bark beetle associated tree mortality in pines at two drought-impacted sites in the San Bernardino Mountains, one receiving high levels of atmospheric pollutants, and one with more moderate atmospheric input. We also investigated the effects of nitrogen addition treatments of 0, 50 and 150 kg N ha−1 year−1 at each site. Tree mortality and beetle activity were significantly higher at the high pollution site. Differences in beetle activity between sites were significantly associated with ozone injury to pines, while differences in tree mortality between sites were significantly associated with both ozone injury and fertilization level. Tree mortality was 9% higher and beetle activity 50% higher for unfertilized trees at the high pollution site compared to the low pollution site. Tree mortality increased 8% and beetle activity increased 20% under the highest rates of nitrogen additions at the low pollution site. The strong response in beetle activity to nitrogen additions at the low pollution site suggests that atmospheric nitrogen deposition increased tree susceptibility to beetle attack at the high deposition site. While drought conditions throughout the region were a major factor in decreased tree resistance, it appears that both ozone exposure and atmospheric nitrogen deposition further increased pine susceptibility to beetle attack.  相似文献   

14.
Nitrogen (N) is usually the most limiting nutrient in degraded agricultural soils and affects the growth and ecological function of poplar (Populus spp.) plantations. We hypothesized that application of organic mulch would improve soil nitrogen availability and increase tree growth, while the quality of mulching materials would alter the supply of essential nutrients. In this study, poplar plantations were established in 2004 and two experiments were established in the field. The fresh above-ground biomass of cogongrass (Imperata cylindrica (L.) P. Beauv.), oak (Quercus fabri Hance), Chinese coriaria (Coriaria nepalensis Wallich) and brackenfern (Pteridium aquilinum (L.) Kuhn. var. latiusculum (Desv.) Underw. ex Heller) in the current year was selected as mulching materials, and mulches were annually applied at the rate of 5.0 kg/m2 in the N mineralization experiment and 20,000 kg/ha in growth and nutrient availability experiment. Additions of fresh biomass significantly improved annual net N-mineralization estimates and the mulching treatments increased the cumulative N mineralized over the incubation period by 22–30%. However, a significant difference in the cumulative N mineralized was not observed among the mulched treatments, even if the cumulative N mineralized in the treatment mulched with oak (QF) was 5.9, 3.3 and 2.2% greater than that of treatments with brackenfern (PA), Chinese coriaria (CN) and cogongrass (IC), respectively. Application of fresh biomass mulch markedly affected soil available N contents and growth performance of poplar plantations. The mulching treatments with QF, IC, PA and CN increased annual means of available N concentration by 39.0, 29.0, 29.6 and 39.7% respectively. At the end of the fifth growing season, the average height of poplar plantations in treatments mulched with QF, IC, PA and CN was 46.8, 14.9, 42.6 and 57.4% greater than that in Check (CK-no mulch), while mean DBH increased by 35.4, 12.5, 33.3 and 52.1%, respectively. Overall, the productivity was enhanced in mulched plots at the end of the third growing season. Compared with CK, the treatments mulched with QF, IC, PA and CN increased total biomass by 97.4, 96.4, 63.1 and 81.6%, respectively. Based on the results of this study, annual application of 20,000 kg/ha fresh woody biomass would be recommended for soil improvement in young poplar plantations growing on a degraded agricultural soil.  相似文献   

15.
Townend J 《Tree physiology》1993,13(4):389-399
Two-year-old Sitka spruce (Picea sitchensis (Bong.) Carr.) plants from four clones were grown in naturally lit growth chambers for 6 months at either ambient (350 ppm) or ambient + 250 ppm (600 ppm) CO(2) concentration. Plants were grown in large boxes filled with peat, in a system that allowed the roots of individual plants to be harvested easily at the end of the growing season. Half of the boxes were kept well watered and half were allowed to dry out slowly over the summer. Plants growing in elevated CO(2) showed a 6.9% increase in mean relative growth rate compared to controls in the drought treatment and a 9.8% increase compared to controls in the well-watered treatment, though there was considerable variation in response among the different clones and water treatments. Rates of net CO(2) assimilation were higher and stomatal conductances were lower in plants grown in elevated CO(2) than in ambient CO(2) in both the well-watered and drought treatments. Both of these factors contributed to the doubling of instantaneous water use efficiency. The partitioning of biomass to roots was unaffected by elevated CO(2), but the ratio of needle mass/stems + branches mass decreased. Together with reduced stomatal conductance, this probably caused the observed increases in xylem pressure potentials with elevated CO(2).  相似文献   

16.
In Southwestern Spain, multifunctional silvopastoral systems consisting of pastureland and open oak woodlands are known as Dehesas. These, and other similar systems of the Mediterranean basin, are currently threatened by increasing intensive land use. As a consequence, oak regeneration is declining and is in need of adequate management and active restoration. Traditional restoration practices outplant one-year-old, nursery-produced oak seedlings grown in 250–350 cm3 containers, but establishment and growth results are typically poor. This work examines holm oak (Quercus ilex L. subsp. ballota (Desf.) Samp.) grown in a non-conventional container size (24 l) and age (6–7 years) with three mulch treatments (control, stone, and straw). In an open Dehesa of SE Spain, 106 oak saplings were planted in March 2010, and survival, diameter at breast height (DBH), and leaf-nutrient concentrations were analyzed. Forty months after planting, all treatments showed high survival (81 %) but only straw-mulched saplings differed significantly (94 %) from control (74 %). DBH increased over time but showed no significant differences among mulch treatments. Saplings with high initial DBH showed the greatest growth and change in DBH at the end of the study period. Leaf-nutrient concentrations changed significantly in the year following outplanting. Bi-monthly foliar nutrient concentration data show decreases in P, K, Zn, and B and sharp increases in Ca and Fe. In this work, we provide some evidence concerning the viability of non-conventional oak size for restoring, regenerating, or building up new agroforestry or silvopastoral systems. A combination of saplings with more than 10 mm of DBH and straw mulch is recommended.  相似文献   

17.
The influence of forest fragmentation on population and community dynamics of woody plants has been well established worldwide, but rarely at the level of an individual plant. We evaluated the influence of fragmentation on juvenile stem morphology of Acer saccharum Marsh. (sugar maple), while also examining light levels and considering possible confounding effects attributed to elevation gradients in temperate forests of northeastern Ohio, USA. At two sites, plant stem dimensions, canopy openness, and relative ground level elevation were measured using randomly positioned plots in forest edge and interior habitats that were within 25 and 60–100 m from a forest edge, respectively. Ratios of stem length to stem basal diameter were greater in forest interiors than near forest edges. These differences in stem morphology between habitats were likely a result of stem elongation in relation to a shade avoidance response in forest interiors that were consistently darker than forest edge areas across study sites. By contrast, such morphological differences were likely not related to variation in relative ground level elevation since a subtle elevation gradient was detected at only one site. We encourage experimentation to identify mechanisms that affect plant stem morphology of young individuals and its influence, in turn, on plant population dynamics in fragmented forests.  相似文献   

18.
[目的]探讨基质配方对红豆树容器苗生长和生理的影响,筛选适宜红豆树容器苗生长、提高苗木生理质量的基质配方,为培育红豆树优质苗木提供技术指导.[方法]以泥炭、椰糠、珍珠岩和蛭石为基质原料,按体积比配制成9种育苗基质,以无纺布袋作为容器进行红豆树容器育苗试验,分析基质配方的理化性质,测定红豆树容器苗苗高、地径、生物量和根系...  相似文献   

19.
【目的】讨论不同产区木荷种源在 NH4+-N或 NO3--N沉降下的生长表现和响应差异,揭示不同形态氮素对木荷生长发育的影响,为在大气氮沉降环境背景下,选育营养高效利用的木荷速生优质新品种提供理论依据。【方法】以木荷北缘种源区-浙江杭州种源、中部种源区-福建建瓯种源和中部靠南边缘种源区-江西信丰种源3个有代表性的木荷种源作为试验材料,模拟不同形态氮沉降( NH4+和NO3-)增加对不同土壤磷素处理下木荷幼苗生长和叶片氮、磷元素含量的影响。盆栽试验设置土壤低磷(1.1 mg·kg -1)处理和高磷(25 mg·kg -1)对照,以人为喷施 NH4 Cl和 NaNO3溶液进行氮沉降模拟,分别设置3个氮沉降量水平:0,80和200 kg·N·hm -2·a -1,试验按完全随机区组设计,每种源每处理重复12株苗。2013年11月收获,测定苗高、地径等生长指标,并分别测定根、茎、叶各部分干物质量和磷、氮含量。【结果】不同形态氮沉降对木荷苗木生长影响差异显著,磷素可提高种源间对氮素的响应差异。在低磷环境下,不同氮处理下木荷植株生物量和根冠比变异系数较大,这为氮沉降下木荷耐受型植株的选择提供了可能。低磷环境下,NO3--N 对木荷苗木生长促进作用显著,苗高、地径和生物量分别较 NH4+-N处理高4.5%,17.8%和75.2%,叶片氮、磷含量提高,叶片 N:P 比下降。NH4+-N 对木荷植株的生长抑制作用较强,导致叶片磷含量下降,N:P升高,植株受到磷胁迫增强。而在高磷环境下,NH4+-N的促进作用增强,苗高、地径和生物量分别较 NO3--N处理高13.5%,10.4%和25.4%。无论土壤在高磷还是低磷环境下,NO3--N 降低叶片N/P比,而 NH4+-N增加叶片 N/P比。木荷种源间对不同形态氮沉降响应差异显著,在土壤低磷环境下,NH4+-N处理抑制了福建建瓯种源和江西信丰种源生长,生物量下降,而杭州种源却在 NH4+-N80处理下,苗高和地径生长较对照分别增加19%和20%。【结论】在低磷环境下,NO3--N对木荷不同种源幼苗生长促进作用更强,而当土壤磷含量提高时,NH4+-N的促进作用增强,同时苗木生长差异增大。浙江杭州种源对 NH4+-N 的适应性更强,而福建建瓯和江西信丰种源则对 NO3--N适应性更强。  相似文献   

20.
This paper characterizes the growth and crown morphology of young balsam fir saplings naturally regenerated under a gradient of understory light environments and intraspecific competition densities for two size classes (50–100 cm and 100–200 cm). Most growth and crown morphological parameters investigated were strongly related to the natural light gradient investigated (3–83% full sunlight), but the relationship tended to plateau at around 25% full sunlight. The relationships were generally better for the larger size class. Intraspecific competition did not significantly affect growth and crown morphology of saplings receiving less than 25% full sunlight, but it affected relative height growth, relative radial growth and the apical dominance ratio for those receiving more than 25% full sunlight (R2=0.506; p<0.001; R2=0.403; p<0.002; R2=0.348; p<0.001, respectively). These results suggest that live crown ratio, apical dominance ratio and the number of internodal branches can provide, alone or in combination, useful indicators of vigour for understory fir. Such a study provides the basic data inputs required for the development of empirically-derived mechanistic models that can predict understory tree growth and survival.  相似文献   

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