Ground-dwelling invertebrates in reforested conifer plantations

Plantation forests are an important part of the forest estate in many countries. In Ireland, they cover around 9% of the land area and many that are commercially mature are now being felled and reforested. The potential biodiversity value of such second rotation forests has yet to be determined, yet this may be particularly significant in Ireland where cover of semi-natural woodland is only 1%. Invertebrates are a vital component of forest biodiversity, functioning as decomposers and pollinators, herbivores, predators and prey. Spiders and Carabid beetles are often used in biodiversity assessment as they are easily captured using pitfall traps, are taxonomically well known and respond to changes in habitat structure. This study aimed to examine spider and Carabid beetle diversity in second rotation Sitka spruce (Picea sitchensis) plantations at different stages of the forest cycle (5, 8–12, 20–30, 35–50 years), and compare the spiders captured in second rotation forests with those from first rotation. Spider and beetle diversity was influenced by stand structural development in second rotation plantations with numbers of forest-associated species increasing over the forest cycle. Overall, spider richness declined over the forest cycle and this was related to decreasing cover of field layer vegetation and fewer open-associated species. In contrast, total beetle richness increased and became more specialised over the forest cycle which may be related to slower colonisation of disturbed areas by beetles in comparison with spiders, and fewer open specialists at the early stages of second rotation. Spider assemblages were distinguished between rotations. This may be related to differing habitat conditions in second rotation forests including dryer soils with lower pH, differing vegetation complexity and presence of brush piles. Few of the forest species accumulated during first rotation were retained and the early stages of second rotation forest cycle was characterised by a generalist open fauna. Nonetheless, as the forest cycle progressed the spider assemblages between rotations became more similar. Current forest policy supports retaining over-mature trees and creating a mosaic of different aged stands within a plantation. Such measures may provide refuge for forest species after clearfell. In countries where forest fragments exist in a landscape dominated by agriculture, consideration should be given to the capacity of mature forest adjacent to felled stands to support forest species, and to the configuration of over-mature areas retained after felling.     

Developing and implementing participatory and ecosystem based multiple use forest management planning approach (ETÇAP): Yalnızçam case study

This paper explains the conceptual framework of ecosystem based multiple use forest management (ETÇAP) approach and its implementation in a case study area. The new management philosophy has four important pillars; integration of biodiversity conservation into forest management process, characterization and accommodation of multiple forest values, effective participation of stakeholders and the use of advanced information technologies and management science techniques. These components comprise a sound framework of a forest management planning system in Turkey as the ownership, land use policy, social structure and forest ecosystems are unique to the country. Among few implementation sites, some experiences from the case study area of Yanlızçam planning unit were documented to realize the performance of the concept. The liaison between the government institutions and major stakeholders is found necessary, and the effective use of geographic information system (GIS) and remote sensing (RS) have been realized critically important to better communicate among stakeholders and generate precise database and the plan. The case study indicated that structured participation is an effective communication tool and has a better possibly to promote multiple use forest management rather than using participation as solo information gathering process. Primary challenges relate to the effectiveness of national conservation program, availability of coherent biodiversity data and adequacy of institutional capacity that involve awareness, training and common understanding of biodiversity and protected area concept; coordination among the related institutions and stakeholders, and willingness and enthusiasm of authorities to accept and implement the concept.     

An examination of stocking and early growth in the Warra silvicultural systems trial confirms the importance of a burnt seedbed for vigorous regeneration in Eucalyptus obliqua forest

Clearfelling of wet eucalypt forest followed by high intensity burning and aerial sowing, a silvicultural system designed to mimic the natural dynamic of sporadic regeneration following cataclysmic disturbance, has attracted criticism for not maintaining the structural diversity that is associated with natural disturbance. A silvicultural systems trial was established at the Warra Long-Term Ecological Research site in southern Tasmania to explore alternatives to clearfelling in tall wet eucalypt forest. Stocking, density and growth of the seedling regeneration were monitored for up to 3 years after harvesting and regeneration treatments were applied from 1998 to 2007. The treatments were clearfell with understorey islands, a patchfell, stripfell, dispersed retention, aggregated retention, and single-tree/small-group selection. High intensity burning, low intensity burning and no burning were variously applied as part of these treatments.     

Regeneration of Populus nine years after variable retention harvest in boreal mixedwood forests

Aspen and balsam poplar regeneration from root suckers were assessed in boreal mixedwood forests nine years after logging in a variable retention experiment (EMEND Project—Ecosystem Management Emulating Natural Disturbance) located north of Peace River, Alberta, Canada. Five levels of retention of mature trees (2%, 10%, 20%, 50% or 75% of the original basal area) were applied in stands dominated by aspen, white spruce or mixtures of the two species. Basal area of aspen (or that of aspen plus balsam poplar combined) prior to logging strongly influenced sucker density of aspen (or aspen + balsam poplar combined) and in some cases their growth. Nine years after harvest there was a decline in sucker density and volume ha⁻¹ with increasing retention levels of aspen (or both poplars combined); sucker density declined by 50% when only 20% of the original basal area was left in the stand. Retaining mature spruce trees in the stand had little influence on the number of suckers but did affect their total volume ha⁻¹. Thus, we suggest that by knowing stand aspen and balsam poplar density prior to logging and varying levels of retention of aspen and balsam poplar or conifers at harvest, the density of Populus regeneration can be predicted by managers, thereby allowing them to create a range of mixedwood conditions.     

Alternative silvicultural practices with variable retention to improve understory plant diversity conservation in southern Patagonian forests

Understory plants could can act as indicators of temperate forest sustainability, health and conservation status due to their importance in ecosystem function. Harvesting impacts on understory plant diversity depends on their intensity. Variable retention has been proposed to mitigate the harmful effects of timber harvesting, but its effectiveness remains unknown in southern Patagonian Nothofagus pumilio forests. The objectives of this study were to: (i) define a baseline of understory plant diversity in old-growth forests along a site quality gradient and under canopy gaps; (ii) evaluate stands with three different variable retention treatments compared to old-growth forests; and (iii) assess temporal changes during 4 years after harvesting (YAH). A 61 ha N. pumilio forest was selected. Understory plant (Dicotyledonae, Monocotyledonae and Pteridophyta) richness, cover (including woody debris and bare forest floor) and aboveground dry biomass were characterized in summer for 5 years. Before harvesting, baseline samples were conducted along a site quality gradient and outside/inside canopy gaps. Analyzed treatments include a control of old-growth forest (OGF) and three different harvesting treatments with variable retention: (i) dispersed retention (DR) of 30 m² ha⁻¹ (20-30% retention); (ii) aggregated retention (AR) with one aggregate per hectare and clear-cuts (28% retention); and (iii) combined dispersed and aggregated retention (DAR) with one aggregate per hectare and dispersed retention of 10-15 m² ha⁻¹ (40-50% retention). Data analyses included parametric and permutational ANOVAs, multivariate classification and ordinations.Before harvesting, 31 plant species were found, where richness, cover and biomass were directly related to site quality. The presence of canopy gaps did not have a significant impact on the measured variables. After harvesting, 20 new species appeared from adjacent associated environments (two from N. antarctica forests and 18 from grasslands and peatlands). At the stand level, understory values were higher in AR > DR > DAR > OGF. Most (81-95%) plant richness at baseline conditions was conserved in all treatments, where inside the aggregates understory remained similar to OGF. Combination of aggregated and dispersed retention (DAR) better limited exotic species introduction and protected sensitive species, improving conservation in harvested stands. Changes in understory variables were observed after the first YAH in all treatments; greater changes were observed in the harvested areas than in aggregates. Changes stabilized at the fourth YAH. As a conclusion, the location of retention aggregates should be selected to preserve species understory diversity of more speciose and diverse habitats or particularly uncommon stands. Implementation of different kinds (patterns and levels) of retention for improvement of biodiversity conservation in harvested forests should be included in timber and forest management planning.     

Effects of clearfelling on forest floor accumulation and litter decomposition in a radiata pine plantation

The short-term effects of clearfelling a Pinus radiata D. Don. stand in Catalonia (NE Spain) under Mediterranean conditions were assessed. Harvesting had little effect on the surface soil distribution and accumulation. However, after clearfelling the range of soil temperatures increased, the moisture content of the L layer decreased, and the decomposition of brown needles was retarded (mass loss rate for the first year of the incubation 0.47 year⁻¹ vs. 0.29 year⁻¹). Green needles incorporated as a result of harvesting operations showed the same rates of mass loss as brown needles. Brown needles released substantial amounts of all major nutrients, especially at the beginning of the incubation. In both clearfelled and undisturbed stands, this sudden nutrient flush may have important implications in the nutrient economy of the vegetation. In contrast, green needles behaved as a sink of nitrogen and calcium, therefore representing a temporary reservoir after clearfelling.     

Early responses of birds to clearfelling and its alternatives in lowland wet eucalypt forest in Tasmania,Australia

The impacts on bird assemblages of silvicultural alternatives to clearfelling in lowland wet eucalypt forest were studied over a decade at Warra, Tasmania. Using a multiple-visit point-count procedure and a before-after-control-impact design, indices of relative incidence were derived for the 44 bird species recorded at 177 survey-points, and used in both univariate and multivariate analyses. The majority of species had strong associations with either mature forest or young regeneration post-harvest, with very few more generalist species present. Bird assemblages in the unharvested mature forest were not static over this period, but this could not readily be explained by the expansion of forest harvesting in the surrounding landscape. However, the shift was subtle compared to the responses to harvesting, which induced a fundamental change in assemblage composition irrespective of the silvicultural system applied. The ability of the different systems to retain elements of the original mature forest avifauna varied markedly. Dispersed retention silviculture was no better than clearfelling, with or without unharvested understorey islands in the latter system. By contrast, both the unharvested parts of coupes subjected to stripfelling, and the retained aggregates in coupes subjected to aggregated retention silviculture, maintained mature forest bird assemblages. The long-term value of stripfelling for mature forest avifauna is compromised because the unharvested strips will be harvested mid-rotation. Aggregated retention silviculture, however, is designed for long-term retention of mature forest in aggregates, and our results suggest that this system may offer a means of sustaining mature forest bird assemblages at the coupe-level. However, individual mature forest species varied in their degree of tolerance, and we suggest that no system is completely resilient to harvesting effects.     

森林生态系统的生物多样性保护问题

生物多样性的保护和可持续的利用对森林的保护与可持续经营具有特别重要的意义,生物多样性保护在森林生态系统管理中也越来越显示其重要性。讨论全球森林生态系统生物多样性保护的发展趋势及森林生态系统生物多样性保护的理论和对策,以及生物多样性指数在评价生物多样性管理的应用,提出我国生物多样性保护研究应该加强的方向。     

汶川地震灾区森林植被恢复问题及其生态对策

2008年5月12日在四川汶川境内发生的里氏8.0级大地震是我国大陆历史上发生的破坏性最严重的地震灾害干扰事件。不仅造成重大经济损失,也使区域生态功能衰退显著。本文在分析地震对森林植被和生物多样性破坏的基础上,总结了震后3年来在森林植被恢复中存在的问题,提出了灾后森林植被恢复生态对策。     

Large-scale interdisciplinary experiments inform current and future forestry management options in the U.S. Pacific Northwest

Over the last 20 years, changing public values and increased ecological understanding have led to a paradigm shift in forestry from timber management to sustainable ecosystem management on U.S. federal lands. Forest managers are now seeking alternative management approaches that simultaneously meet socio-cultural, ecological and economic goals. Consequently, many field experiments have become increasingly interdisciplinary and larger in scale or scope. Individually and collectively, these studies in western Washington and Oregon represent major investments by research and land management organizations to enhance the science and understanding for sustainable forest management under increasing public scrutiny and demands for safeguarding healthy environments, conserving biological diversity and providing some level of economic prosperity. They also help to facilitate the transfer of scientific results into practical applications and to realize a more effective interface between science and policy. Questions addressed in this paper include (i) what do we mean by large-scale experiments, (ii) who is investing in these kinds of experiments and why, (iii) where is this information being put to use, and (iv) what does the future hold for these studies?     

Threshold effects of variable retention harvesting on understory plant communities in the boreal mixedwood forest

We studied the effects of six levels of dispersed green-tree retention (GTR) harvesting (clearcut (0%), 10%, 20%, 50%, and 75%, and unharvested reference (100%)) on understory plant communities in the 8th growing season post-harvest in the mixedwood boreal forest in northwestern Alberta. For the partial harvest treatments (10%, 20%, 50%, 75%) sample plots were located in the partially harvested (retention) strips as well as in the intervening machine corridors used by the harvesting equipment. The understory plant community was significantly influenced by the gradient of retention level. The cover of understory vegetation, especially graminoids, increased with increasing harvesting intensity for the retention strips and overall considering both plots types. Species richness was unaffected by retention level but did decrease as tree density increased. Lower levels of retention lead to increased abundance of early successional, shade-intolerant species. The results suggest a threshold in understory response to GTR harvesting between the 10% and 20% retention treatments. In terms of understory cover and composition, machine corridors within partially harvested forests resembled clearcuts. The results suggest that retaining more than 10% during GTR harvesting could have significant benefits in terms of maintaining understory plant communities more similar to unharvested reference forest.