Long-Term Effects of Weed Control With Picloram Along a Gradient of Spotted Knapweed Invasion

Broadleaf herbicides are commonly used in rangelands to suppress exotic weeds and release native communities from negative impacts of invasion. However, few studies have comprehensively evaluated treatment effects on differing community components across a gradient of initial invasion levels. We conducted a 6-yr experiment within grasslands of western Montana to measure local-scale effects of a broadcast application of picloram on 1) cover of the target invader, spotted knapweed (Centaurea stoebe L.), 2) prevalence of native functional groups, and 3) the secondary invader cheatgrass (Bromus tectorum L.) at differing initial levels of knapweed invasion. Treatment effectively suppressed knapweed, with cover in treated vs. control plots reduced by >60% in the sixth posttreatment year. Treatment also appeared to alleviate knapweed’s impacts on native perennial grasses, but only at the highest initial level of invasion, where cover of this group increased by >30% in treated vs. control plots to equal levels associated with noninvaded plots. In some cases, treatment appeared to exacerbate knapweed’s impacts on native forbs. At the no-invasion level, perennial forb cover declined by >20% in treated vs. control plots to match values associated with moderate or high levels of invasion, but these treatment effects were minimal at the latter invasion levels. Across initial invasion levels, species richness of perennial and/or annual forbs declined by >20% in treated vs. control plots. Treatment also promoted increases in cheatgrass cover, although differences between treated and control plots were relatively small by the sixth posttreatment year. Overall, effects of picloram application depended on initial levels of knapweed invasion, largely due to the varying strength of release effects. Selective treatment of invaded patches vs. broadcast applications would reduce side effects of broadleaf herbicide application and increase compatibility with other management measures designed to improve rangeland conditions and restore grassland communities.     

Responses of Chaparral and Oak Woodland Plant Communities to Fuel-Reduction Thinning in Southwestern Oregon

Fire suppression has led to large fuel accumulations in many regions of the United States. In response to concerns about associated wildfire hazards, land managers in the western United States are carrying out extensive fuel-reduction thinning programs. Although reductions in cover by woody vegetation seem likely to cause changes in herbaceous communities, few published studies have reported on consequences of such treatments for native or exotic plant species. We compared vegetation and abiotic characteristics between paired thinned and unthinned chaparral and oak woodland communities of southwestern Oregon 4–7 yr posttreatment and contrasted impacts of manual vs. mechanical treatments. Herbaceous cover increased on thinned sites, but species richness did not change. Herbaceous communities at thinned sites had an early postdisturbance type of composition dominated by native annual forbs and exotic annual grasses; cover by annuals was nearly twice as high on treated as on untreated sites. Absolute and proportional cover of native annual forbs increased more than any other trait group, whereas exotic annual forbs and native perennial forbs declined. Exotic annual grass cover (absolute and proportional) increased, whereas cover by native perennial grasses did not. Shrub reestablishment was sparse after thinning, probably because of a lack of fire-stimulated germination. Manual and mechanical treatment impacts on abiotic site conditions differed, but differences in vegetation impacts were not statistically significant. Fuel-reduction thinning may have some unintended negative impacts, including expansion of exotic grasses, reductions in native perennial species cover, persistent domination by annuals, and increased surface fuels. Coupled with sparse tree or shrub regeneration, these alterations suggest that ecological-state changes may occur in treated communities. Such changes might be mitigated by retaining more woody cover than is currently retained, seeding with native perennials after treatment, or other practices; further research is needed to inform management in these ecosystems.     

Land-Use Legacies and Vegetation Recovery 90 Years After Cultivation in Great Basin Sagebrush Ecosystems

Agricultural land use is known to alter ecological processes, and native plant communities can require decades to centuries to recover from the disturbance of cultivation. “Recovery” is typically measured by comparison to undisturbed adjacent sites as a control. Recovery following cultivation in sagebrush ecosystems of the Great Basin remains largely unexamined even though nearly a half million hectares of land were dry-farmed and abandoned in the early 1900s. We tested the hypothesis that the native vegetation has not recovered from this exotic disturbance by evaluating differences in canopy cover of shrubs, grasses, and forbs between paired sets of historically dry-farmed land and adjacent never-cultivated areas. Paired sites were located in three ecological sites in northwestern Utah. We found that vegetation recovery from cultivation is variable by growth form, species, and ecological site. Shrub recovery was different among sagebrush (Artemisia) species. Yellow rabbitbrush (Chrysothamnus viscidiflorus [Hook.] Nutt.) and black greasewood (Sarcobatus vermiculatus [Hook.] Torr.), which often increase following disturbance, maintained higher cover inside old fields. At one of the paired sets, shrub composition was altered from a mix of four species to dominance of mainly Wyoming big sagebrush (Artemisia tridentata Nutt. subsp. wyomingensis Beetle & Young). Total forb cover was generally lower in cultivated areas and some species, such as spiny phlox (Phlox hoodii Richardson), had not recovered. The most common grass species encountered across all ecological sites, bottlebrush squirreltail (Elymus elymoides [Raf.] Swezey), had higher cover in cultivated areas. Surprisingly, exotic annual species, such as cheatgrass (Bromus tectorum L.), did not dominate these sites as they have for decades after cultivation in other areas of the Great Basin. This study demonstrates that the land-use legacy of dry farming on vegetation remains nearly a century after cultivation has ceased, and has direct implications for describing ecological site conditions.     

Native Vegetation Composition in Crested Wheatgrass in Northwestern Great Basin

Crested wheatgrass, an introduced perennial bunchgrass, has been seeded extensively on the rangelands of western North America. There is a perception that this species is very competitive and that it forms monoculture or low diversity stands where successfully seeded. However, there is limited information on species composition in sites previously seeded to crested wheatgrass. We measured native vegetation and environmental characteristics in areas seeded with crested wheatgrass across the northwestern Great Basin. Plant community composition within these crested wheatgrass stands was variable, from seedings that were near monocultures of crested wheatgrass to those that contained more diverse assemblages of native vegetation, especially shrubs. Environmental factors explained a range of functional group variability from 0% of annual grass density to 56% of large native bunchgrass density. Soil texture appeared to be an important environmental characteristic in explaining vegetation cover and density. Native vegetation was, for all functional groups, positively correlated with soils lower in sand content. Our results suggest environmental differences explain some of the variability of native vegetation in crested wheatgrass stands, and this information will be useful in assessing the potential for native vegetation to co-occupy sites seeded with crested wheatgrass. This research also suggests that crested wheatgrass seedings do not always remain in near monoculture vegetation states as seedings substantially varied in native vegetation composition and abundance with some seeded areas having a more diverse assemblage of native vegetation. In half the sites, there were five or more perennial herbaceous species and 63% of sites contained Wyoming big sagebrush. Although not exclusively true, species most commonly encountered in crested wheatgrass seedings are those that are able to minimize competition with crested wheatgrass via temporal (i.e., Sandberg bluegrass, annual forbs, annual grasses) or spatial (i.e., shrubs) differentiation in resource use.     

Evaluation of Herbicide and Disking to Control Invasive Bluestems in a South Texas Coastal Prairie

Conservation and restoration efforts of native grasslands are being hindered by invasive, exotic plants. Exotic bluestem grasses (Bothriochloa and Dichanthium spp.) have become increasingly invasive throughout the rangelands of the central and southern Great Plains, United States. Accordingly, the aim of this study was to evaluate the efficacy of glyphosate, imazapyr, and imazapyr + glyphosate treatments with or without disking to remove exotic bluestems from a south Texas coastal prairie. We evaluated three different control regimens: 1) herbicide treatments only, 2) herbicide treatments followed by two diskings (H + D), and 3) disking followed by herbicide treatments (D + H). Percent exotic bluestem, native grass, and forb cover were visually estimated at 0 (pre-treatment: May 2006), 20, 52, and 104 wk after treatment (WAT). The herbicide-only and H + D regimens were ineffective at controlling exotic bluestems. However, exotic bluestem cover in herbicide-treated plots of the D + H regimen was significantly lower (P ≤ 0.05) compared to control plots and most treatment plots of the herbicide-only and H + D regimens up to 52 WAT. Control regimens did not notably facilitate an increase in native grass cover from pre-treatment levels, but native grass cover remained the highest, and increased the most, in some imazapyr-treated plots of the herbicide-only and D + H regimens, respectively. In the H + D and D + H regimens, disking resulted in a flush of forb cover (up to 50%) at 52 WAT; yet forb cover was ≤ 5% in these plots by 104 WAT. Exotic bluestem cover recovered back to, or was greater than, pre-treatment levels among most treatment plots across all three control regimens at 104 WAT. This study suggests that follow-up control measures are needed to suppress the re-invasion of exotic bluestems after initial control efforts. Additional studies are needed to evaluate other strategies to control exotic bluestems in rangelands of the central and southern United States.     

Lack of Native Vegetation Recovery Following Biological Control of Leafy Spurge

Leafy spurge (Euphorbia esula L.) is an aggressive exotic species that has been successfully suppressed in a variety of situations using classical biological control (flea beetles; Aphthona spp.). This 9-yr study investigated patterns of vegetation responses following significant reductions in leafy spurge cover and density by flea beetles in southeastern Montana. We hypothesized that the vegetation following leafy spurge suppression would be dominated by species and plant functional groups able to persist through heavy infestations. Flea beetles were first released in 1998, and by 2006 leafy spurge foliar cover was reduced 80% to 90% compared to 1998 values on both release and nonrelease plots. Although total cover of the resident vegetation, excluding leafy spurge, increased 72% to 88%, relative cover of the functional groups (native forbs, native sedges, native grasses, and non-native species) was similar among years and between release and nonrelease plots. Mean diversity and mean species richness values did not differ among years or between release and nonrelease plots (P < 0.05), but mean diversity on both release and nonrelease plots was significantly less than noninfested plots, although richness was similar (P < 0.05). Indicator species analysis revealed that non-native Poa spp. replaced leafy spurge as the dominant species on release and nonrelease plots. Conversely, noninfested plots contained a variety of native species with high indicator values. Although total abundance of the resident vegetation in 2006 was significantly greater than 1998, plant species composition and relative cover showed little change for the duration of the study. Failure of the native vegetation to recover to a community that approached nearby noninfested conditions may be attributed to a variety of interacting scenarios, some of which may be ameliorated by treating infestations as soon as possible to avoid long-term residual effects.     

Long-Term Effects of Aeration and Fire on Invasion of Exotic Grasses in Mixed-Brush Plant Communities

Invasion of exotic grasses into grasslands dominated by native plants changes fire cycles and reduces biodiversity. Brush management practices that create soil disturbance, such as aeration, may potentially result in invasion of exotic grasses and replacement of native vegetation. We tested the hypothesis that a long-term effect of aeration and prescribed burning is an increase in exotic grasses. The study was conducted at the Chaparral Wildlife Management Area in the western south Texas plains where four treatments were evaluated: aeration, warm-season burn, aeration followed by a warm-season burn, and no treatment (control). The experimental design was a randomized complete block with four replicates. We estimated percentage canopy cover of exotic grasses, native grasses, forbs, litter, bare ground, and woody and succulent plants in 2007. There was a multivariate main effect among treatments for the dependent variables absolute canopy cover of exotic grasses, native grasses, forbs, litter, and bare ground (Wilks's Lambda F_15,179.84 = 2.78, P = 0.001). Variables that contributed to the significant overall effect included litter (F_3,69 = 4.32, P = 0.008) and native grasses (F_3,69 = 6.11, P = 0.001). The multivariate main effect of treatment was significant (Wilks's Lambda F_9,180.25 = 2.04, P = 0.038) for the relative canopy cover of herbaceous species. Relative cover of exotic grasses was 31% higher (P = 0.024) in control than in the prescribed burn treatment. Native grasses relative cover was 30% higher (P = 0.003) in prescribed burn than in the control treatment. We did not detect differences among treatments in the percentage of total woody and succulent plants canopy cover (P = 0.083). Under the environmental conditions at the time of the study, aeration and/or prescribed burning do not increase exotic grasses.     

A Method For Landscape-Scale Vegetation Assessment: Application to Great Basin Rangeland Ecosystems

The growth of landscape-scale land management necessitates the development of methods for large-scale vegetation assessment. Field data collection and analysis methods used to assess ecological condition for the 47 165-h North Spring Valley watershed are presented. Vegetation cover data were collected in a stratified random design within 6 Great Basin vegetation types, and the probability of detecting change in native herbaceous cover was calculated using power analyses. Methods for using these quantitative assessment data are presented to calculate a departure index based on reference condition information from LANDFIRE (an interagency effort to map and model fire regimes and other biophysical characteristics at a mid-scale for the entire United States) Biophysical Setting models for the mountain big sagebrush (Artemisia tridentata Nutt. subsp. vaseyana [Rydb.] Beetle) vegetation type. For mountain big sagebrush in the North Spring Valley landscape, we found that the earliest successional classes were underrepresented and that mountain big sagebrush moderately invaded by conifers was more abundant than predicted by the LANDFIRE reference based on the historic range of variability. Classes that were most similar to the reference were mountain big sagebrush with the highest conifer cover and late development mountain big sagebrush with perennial grasses. Overall, results suggested that restoration or approximation of the historic fire regime is needed. This method provides a cost-effective procedure to assess important indicators, including native herbaceous cover, extent of woody encroachment, and ground cover. However, the method lacks the spatial information that would allow managers to comprehensively assess spatial patterns of vegetation condition across the mosaics that occur within each major vegetation type. The development of a method that integrates field measurements of key indicators with remotely sensed data is the next critical need for landscape-scale assessment.     

Effect of Pinyon–Juniper Tree Cover on the Soil Seed Bank

As pinyon–juniper (specifically, Pinus monophylla and Juniperus osteosperma) woodlands in the western United States increase in distribution and density, understory growth declines and the occurrence of crown fires increases, leaving mountainsides open to both soil erosion and invasion by exotic species. We examined if the loss in understory cover that occurred with increasing tree cover was reflected in the density and diversity of the seed bank. Seed banks in stands with low, medium, and high tree cover were measured in late October for 2 yr. Multivariate analyses indicated that cover and diversity of standing vegetation changed as tree cover increased. However, the seed bank did not differ in overall seed density or species diversity because seeds of the 13 species that comprised 86% of the seed bank occurred in similar density across the tree-cover groups. Sixty-three percent of the species that were in the seed bank were absent from the vegetation (mostly annual forbs). In addition, 49% of the species that occurred in the standing vegetation were not in the seed bank (mostly perennial forbs and shrubs). Only Artemisia tridentata, Bromus tectorum, and Collinsia parviflora displayed positive Spearman rank correlations between percent cover in the vegetation and density in the seed bank. Thus, much of the standing vegetation was not represented in the seed bank, and the few species that dominated the seed bank occurred across varying covers of pinyon–juniper.     

Mowing Wyoming Big Sagebrush Communities With Degraded Herbaceous Understories: Has a Threshold Been Crossed?

Wyoming big sagebrush (Artemisia tridentata ssp. wyomingensis &lsqb;Beetle & A. Young] S.L. Welsh) plant communities with degraded native herbaceous understories occupy vast expanses of the western United States. Restoring the native herbaceous understory in these communities is needed to provide higher-quality wildlife habitat, decrease the risk of exotic plant invasion, and increase forage for livestock. Though mowing is commonly applied in sagebrush communities with the objective of increasing native herbaceous vegetation, vegetation response to this treatment in degraded Wyoming big sagebrush communities is largely unknown. We compared mowed and untreated control plots in five Wyoming big sagebrush plant communities with degraded herbaceous understories in eastern Oregon for 3 yr posttreatment. Native perennial herbaceous vegetation did not respond to mowing, but exotic annuals increased with mowing. Density of cheatgrass (Bromus tectorum L.), a problematic exotic annual grass, was 3.3-fold greater in the mowed than untreated control treatment in the third year posttreatment. Annual forb cover, largely consisting of exotic species, was 1.8-fold greater in the mowed treatment compared to the untreated control in the third year posttreatment. Large perennial grass cover was not influenced by mowing and remained below 2%. Mowing does not appear to promote native herbaceous vegetation in degraded Wyoming big sagebrush plant communities and may facilitate the conversion of shrublands to exotic annual grasslands. The results of this study suggest that mowing, as a stand-alone treatment, does not restore the herbaceous understory in degraded Wyoming big sagebrush plant communities. We recommend that mowing not be applied in Wyoming big sagebrush plant communities with degraded understories without additional treatments to limit exotic annuals and promote perennial herbaceous vegetation.     

Sage Grouse Groceries: Forb Response to Piñon-Juniper Treatments

Juniper and piñon coniferous woodlands have increased 2- to 10-fold in nine ecoregions spanning the Intermountain Region of the western United States. Control of piñon-juniper woodlands by mechanical treatments and prescribed fire are commonly applied to recover sagebrush steppe rangelands. Recently, the Sage Grouse Initiative has made conifer removal a major part of its program to reestablish sagebrush habitat for sage grouse (Centrocercus urophasianus) and other species. We analyzed data sets from previous and ongoing studies across the Great Basin characterizing cover response of perennial and annual forbs that are consumed by sage grouse to mechanical, prescribed fire, and low-disturbance fuel reduction treatments. There were 11 sites in western juniper (Juniperus occidentalis Hook.) woodlands, 3 sites in singleleaf piñon (Pinus monophylla Torr. & Frém.) and Utah juniper (Juniperus osteosperma [Torr.] Little), 2 sites in Utah juniper, and 2 sites in Utah juniper and Colorado piñon (Pinus edulis Engelm). Western juniper sites were located in mountain big sagebrush (A. tridentata ssp. vaseyana) steppe associations, and the other woodlands were located in Wyoming big sagebrush (A. tridentata ssp. wyomingensis) associations. Site potential appears to be a major determinant for increasing perennial forbs consumed by sage grouse following conifer control. The cover response of perennial forbs, whether increasing (1.5- to 6-fold) or exhibiting no change, was similar regardless of conifer treatment. Annual forbs favored by sage grouse benefitted most from prescribed fire treatments with smaller increases following mechanical and fuel reduction treatments. Though forb abundance may not consistently be enhanced, mechanical and fuel reduction conifer treatments remain good preventative measures, especially in phase 1 and 2 woodlands, which, at minimum, maintain forbs on the landscape. In addition, these two conifer control measures, in the short term, are superior to prescribed fire for maintaining the essential habitat characteristics of sagebrush steppe for sage grouse.     

Livestock as Ecosystem Engineers for Grassland Bird Habitat in the Western Great Plains of North America

Domestic livestock have the potential to function as ecosystem engineers in semiarid rangelands, but prevailing management practices largely emphasize livestock production and uniform use of vegetation. As a result, variation in vegetation structure might not occur at appropriate spatial and temporal scales to achieve some contemporary conservation objectives. Here, we introduce the utility of livestock as ecosystem engineers and address potential benefits and consequences associated with heterogeneity-based management practices for conservation grazing in the semiarid rangelands of the western North American Great Plains. To illustrate the potential value of this approach, we provide specific examples where engineering effects of livestock could alter vegetation heterogeneity at within-pasture (< 100 ha) and among-pasture (～100 ha to thousands of hectares) scales to improve habitat for declining native grassland birds. Experimental evaluations of the efficacy of livestock to achieve desired modifications to vegetation structure are needed, along with the economic aspects associated with implementing heterogeneity-based management practices. Using livestock as ecosystem engineers to alter vegetation structure for grassland bird habitat is feasible in terms of application by land managers within the context of current livestock operations, and provides land managers important tools to achieve desired contemporary objectives and outcomes in semiarid rangelands of the western North American Great Plains.     

A Hierarchical Perspective to Woody Plant Encroachment for Conservation of Prairie-Chickens

Encroachment of Great Plains grasslands by fire-sensitive woody plants is a large-scale, regional process that fragments grassland landscapes. Using prairie grouse (Tympanuchus spp.) of conservation concern, we apply hierarchy theory to demonstrate how regional processes constrain lower-level processes and reduce the success of local management. For example, fire and grazing management may be locally important to conservation, but the application of fire and grazing disturbances rarely cause irreversible fragmentation of grasslands in the Great Plains. These disturbance processes cause short-term alterations in vegetation conditions that can be positive or negative, but from a long-term perspective fire maintains large tracts of continuous rangelands by limiting woody plant encroachment. Conservation efforts for prairie grouse should be focused on landscape processes that contribute to landscape fragmentation, such as increased dominance of trees or conversion to other land uses. In fact, reliance on local management (e.g., maintaining vegetation structure) to alter prairie grouse vital rates is less important to grouse population persistence given contemporary landscape level changes. Changing grass height, litter depth, or increasing the cover of forbs may impact a few remaining prairie-chickens, but it will not create useable space at a scale relevant to the historic conditions that existed before land conversion and fire suppression.     

Extent of Coterminous US Rangelands: Quantifying Implications of Differing Agency Perspectives

Rangeland extent is an important factor for evaluating critical indicators of rangeland sustainability. Rangeland areal extent was determined for the coterminous United States in a geospatial framework by evaluating spatially explicit data from the Landscape Fire and Resource Management Planning Tools (LANDFIRE) project describing historic and current vegetative composition, average height, and average cover through the viewpoints of the Natural Resources Inventory (NRI) administered by the Natural Resources Conservation Service and the Forest Inventory and Analysis (FIA) program administered by the US Forest Service. Three types of rangelands were differentiated using the NRI definition encompassing rangelands, afforested rangelands, and transitory rangelands. Limitations in the FIA definition permitted characterization of only two rangeland types: rangeland and rangeland vegetation with a small patch size. These classes were similar to those from the NRI definition but differed in tree canopy cover threshold requirements. Estimated rangeland area resulting from the NRI- and FIA-LANDFIRE models were 268 and 207 Mha, respectively. In addition, the NRI-LANDFIRE model identified 19 Mha of afforested rangelands due principally to encroachment and increased density by species classified as trees belonging to the genera Quercus, Prosopis, and Juniperus. The biggest discrepancies between acreage estimates derived from NRI- and FIA-LANDFIRE models occurred in oak, pinyon-juniper, and mesquite woodlands. The differences in area estimates between the NRI and FIA perspectives demonstrate the need for development of unified, objective methods for determining rangeland extent that can be applied consistently to all rangelands regardless of ownership or jurisdiction. While the models and geospatial information developed here are useful for national-scale estimates of rangeland extent, they are subject to the limitations of the LANDFIRE data products.     

Performance of Quantitative Vegetation Sampling Methods Across Gradients of Cover in Great Basin Plant Communities

Resource managers and scientists need efficient, reliable methods for quantifying vegetation to conduct basic research, evaluate land management actions, and monitor trends in habitat conditions. We examined three methods for quantifying vegetation in 1-ha plots among different plant communities in the northern Great Basin: photography-based grid-point intercept (GPI), line-point intercept (LPI), and point-quarter (PQ). We also evaluated each method for within-plot subsampling adequacy and effort requirements relative to information gain. We found that, for most functional groups, percent cover measurements collected with the use of LPI, GPI, and PQ methods were strongly correlated. These correlations were even stronger when we used data from the upper canopy only (i.e., top “hit” of pin flags) in LPI to estimate cover. PQ was best at quantifying cover of sparse plants such as shrubs in early successional habitats. As cover of a given functional group decreased within plots, the variance of the cover estimate increased substantially, which required more subsamples per plot (i.e., transect lines, quadrats) to achieve reliable precision. For GPI, we found that that six–nine quadrats per hectare were sufficient to characterize the vegetation in most of the plant communities sampled. All three methods reasonably characterized the vegetation in our plots, and each has advantages depending on characteristics of the vegetation, such as cover or heterogeneity, study goals, precision of measurements required, and efficiency needed.     

Impact of Cultivation Legacies on Rehabilitation Seedings and Native Species Re-Establishment in Great Basin Shrublands

Little is known about how cultivation legacies affect the outcome of rehabilitation seedings in the Great Basin, even though both frequently co-occur on the same lands. Similarly, there is little known about how these legacies affect native species re-establishment into these seedings. We examined these legacy effects by comparing areas historically cultivated and seeded to adjacent areas that were seeded but never cultivated, for density of seeded crested wheatgrass (Agropyron cristatum [L.] Gaertn.) and native perennial grasses, vegetation cover, and ground cover. At half of the sites, historically cultivated areas had lower crested wheatgrass density (P < 0.05), and only one site had a higher density of crested wheatgrass (P < 0.05). Likewise, the native shrub Wyoming big sagebrush (Artemisia tridentata Nutt. subsp. wyomingensis Beetle & Young) had lower cover (P < 0.05) in historically cultivated areas at half the sites. Sandberg bluegrass (Poa secunda J. Presl.) density was consistently lower in historically cultivated areas relative to those seeded-only. At sites where black greasewood (Sarcobatus vermiculatus [Hook.] Torr.) and bottlebrush squirreltail (Elymus elymoides [Raf.] Swezey) were encountered, there was either no difference or a higher density and cover within historically cultivated areas (P < 0.05). Likewise, cover of exotic forbs, especially halogeton (Halogeton glomeratus [M. Bieb.] C. A. Mey.), was either not different or higher in historically cultivated areas (P < 0.05). Bare ground was greater in historically cultivated areas at three sites (P < 0.05). These results suggest that cultivation legacies can affect seeding success and re-establishment of native vegetation, and therefore should not be overlooked when selecting research sites or planning land treatments that include seeding and or management to achieve greater native species diversity.     

Using Unmanned Helicopters to Assess Vegetation Cover in Sagebrush Steppe Ecosystems

Evaluating vegetation cover is an important factor in understanding the sustainability of many ecosystems. Remote sensing methods with sufficient accuracy could dramatically alter how biotic resources are monitored on both public and private lands. Idaho National Laboratory (INL), in conjunction with the University of Idaho, evaluated whether unmanned aerial vehicles (UAVs) are sufficiently accurate and more efficient than the point-frame field method for monitoring vegetative cover and bare ground in sagebrush steppe ecosystems. These values are of interest to land managers because typically there are limited natural resource scientists and funding for comprehensive ground evaluations. In this project, unmanned helicopters were used to collect still-frame imagery to determine vegetation cover during June and July 2005. The images were used to estimate percent cover for six vegetative cover classes (shrub, dead shrub, grass, forbs, litter, and bare ground). Field plots used to collect imagery and on-the-ground measurements were located on the INL site west of Idaho Falls, Idaho. Ocular assessments of digital imagery were performed using SamplePoint, and the results were compared with field measurements collected using a point-frame method. The helicopter imagery evaluation showed a high degree of agreement with field cover class values for grass, litter, and bare ground and reasonable agreement for dead shrubs. Shrub cover was often overestimated, and forbs were generally underestimated. The helicopter method took 45% less time than the field method. This study demonstrates that UAV technology provides a viable method for monitoring selective types of cover on rangelands and could save time and resources.     

Basalt Milkvetch Responses to Novel Restoration Treatments in the Great Basin

The restoration of native forbs in the Great Basin and similar dryland ecosystems remains a great challenge for land managers. Variable soil water, precocious germination and emergence, and the presence of soil fungal pathogens often reduce plant establishment. Novel restoration treatments that increase soil water during early forb life stages, delay germination and emergence to coincide with favorable spring conditions, and reduce mortality from soil fungal pathogens may benefit native forb restoration. We compared the efficacy of three novel treatments—snow fencing, plant protection fabric, and seed coatings—to improve establishment of a forb species native to the Great Basin, basalt milkvetch (Astragalus filipes Torr. ex A. Gray). We replicated treatments plus a no-treatment control at three sites in the Great Basin in a randomized complete block design over 2 yr. To evaluate the efficacy of our restoration treatments, we measured germination, seedling emergence, establishment, and second-yr survival. The effects of snow fencing and plant protection fabric varied by site and life stage and were likely influenced by the wet spring conditions across sites, particularly the two northern sites. Snow fencing increased establishment and second-yr survival at one site. Plant protection fabric did not benefit any life stage at any site. Seed coatings increased seedling emergence at all sites but did not affect establishment or survival. Overall, second-yr survival remained low across all sites and treatments. Our study indicates that our treatments can positively affect basalt milkvetch emergence, establishment, and survival, but additional management actions are needed to improve long-term restoration success of native forbs.     

Are Early Summer Wildfires an Opportunity to Revegetate Exotic Annual Grass–Invaded Plant Communities?

Medusahead (Taeniatherum caput-medusae [L.] Nevski) is an exotic annual grass invading western rangelands. Successful revegetation of invaded-plant communities can be prohibitively expensive because it often requires iterative applications of integrated control and revegetation treatments. Prescribed burning has been used to control medusahead and prepare seedbeds for revegetation, but burning has been constrained by liability concerns and has produced widely varying results. Capitalizing on naturally occurring wildfires could reduce revegetation costs and alleviate liability concerns. Thus, our objective was to determine if early summer wildfires and fall drill seeding could be used as a treatment combination to decrease medusahead and increase perennial and native vegetation. Treatments were evaluated pretreatment and for 3 yr postfire at six sites and included 1) an early summer wildfire combined with a seeding treatment (burn and seed) and 2) a nontreated (no burn, no seed) control. Perennial grass density was 4.6- to 10.0-fold greater in the burn-and-seed treatment compared to the control in the first 3 yr posttreatment (P < 0.05). Exotic annual grass density and cover in the third year posttreatment were lower in the burn-and-seed treatment than in the control (P < 0.05). However, exotic annual grass density was still > 130 individuals · m^?2 in the burn-and-seed treatment. The density of exotic annual grass is of concern because over time medusahead may displace perennial grasses and annual forbs that increased with the burn-and-seed treatment. Though not directly tested in this study, we suggest that, based on other research, the burn-and-seed treatment may need to incorporate a preemergent herbicide application to further suppress medusahead and increase the establishment of seeded vegetation. However, it appears that early summer wildfires may provide an opportunity to reduce the cost of integrated programs to revegetate medusahead-invaded plant communities.     

Short-Term Response of Two Beneficial Invertebrate Groups to Wildfire in an Arid Grassland System,United States

Rangeland invertebrates contribute greatly to biodiversity and provide important services including pollination, pest control, and nutrient cycling. As wildfire frequency increases across these areas of the United States, it is imperative to understand how these disturbances affect beneficial invertebrate communities. We examined bee (Hymenoptera), spider (Araneae), and vegetative communities 1 yr before and 1 yr after a large wildfire swept across an intact grassland in eastern Oregon. Several sites were left unburned after the fire, and a before-after-control-impact study design was used to assess changes within the communities. Fire had no effect on bee or spider abundance, or spider diversity or richness; however, fire significantly increased native bee diversity and richness. In addition, composition of both native bee and spider communities differed significantly between burned and unburned areas 1 yr after the fire. Sheet web spiders (Linyphiidae) and several bee species (primarily large, generalist species) were associated with burned sites. Invasive annual grass and biological soil crust cover decreased significantly in burned sites, but maximum vegetation height and litter cover did not differ significantly among treatments. Forb abundance increased in burned sites; however, species richness of forbs in burned and unburned sites did not differ significantly 1 yr after the fire. Several forbs were indicative of burned areas including non-native species, such as Douglas’ knotweed (Polygonum douglasii) and Russian thistle (Salsola tragus), and native species such as Canadian horseweed (Conyza canadensis), hoary tansyaster (Machaeranthera canescens), and tall willowherb (Epilobium brachycarpum). This study demonstrates that both invertebrate and plant communities show strong short-term responses to wildfire, and our results can be used to inform management of rare habitat and biodiversity in rangelands impacted by wildfire in arid grasslands.