The relationship between level of sward height reduction in a rotationally grazed sward and short-term intake rates of dairy cows

The objective of this study was to investigate the relationship between level of sward height reduction (SHR) and short-term intake of herbage by lactating dairy cows offered swards differing in initial bulk density (BD). Three experiments were carried out in which cows were presented with swards representing different levels of SHR (nominally described as ungrazed, low, moderate and high). Experiments 1 and 2 differed with respect to initial sward BD [ungrazed sward 1·7 vs. 2·5 kg dry matter (DM) m^?3 respectively]. Experiment 3 investigated the interaction between BD and SHR. In each experiment, sixteen Holstein/Friesian cows (fitted with excreta collection bags) grazed for a 1-h period in 200-m² plots that had been grazed on the previous day to predetermined sward surface heights (SSH) (= levels of SHR). Herbage intake rates were assessed from changes in live weight, with a correction for insensible weight loss (IWL). Biting rates were recorded from visual observation. High levels of SHR were associated with a significant reduction in SSH, herbage mass and leaf fraction, and a significant increase in sward BD, stem and dead fraction, and DM concentration. Herbage intake, expressed either as DM intake per bite or DM intake per hour, declined as level of SHR increased from low to high. The level of SHR generally had no effect on biting rate. Intake rates varied from 1·9 to 4·4 kg DM h^?1, whereas DM intake bite^?1 ranged from 0·5 to 1·3 g. Pooled regression analysis identified SSH (P < 0·001; r² = 0·94) as the principal determinant of DM intake bite^?1. The regression equation was not significantly improved by the addition of terms for leaf fraction, BD, or herbage mass. In Experiment 3, a significant interaction between level of SHR and sward BD was observed. It is concluded that the principal factor controlling intake (g DM bite^?1 or kg DM h^?1), as swards are progressively grazed down, is SSH, but at a high level of SHR, sward BD also influences intake bite^?1.     

Pattern of herbage intake rate and bite dimensions of rotationally grazed dairy cows as sward height declines

To allow improved prediction of daily herbage intake of dairy cows in rotational grazing systems, intake behaviour was assessed throughout the day in 24‐h paddocks. Herbage intake in 16 lactating Holstein–Friesian cows was assessed using the short‐term (1‐h) weight gain method at four predetermined natural meal times throughout the day (early morning, T1; late morning, T2; mid‐afternoon, T3; and early evening, T4). The study comprised two 4‐day experiments, each with a cross‐over design of four blocks. In both experiments, cows grazed a 24‐h paddock daily, and the effect of the immediately previous grazing experience on intake behaviour was investigated throughout the day, taking account of daily fluctuations in the short‐term physiological condition of the cows. Experiment 1 was carried out to investigate overall grazing behaviour during meals as a sward is progressively depleted during the day, with intake being assessed within the paddock and, hence, on a depleted sward. Experiment 2 similarly investigated the effect of sward depletion and physiological condition throughout the day on intake, but cows were removed to fresh, undefoliated swards during intake measurement periods; thus, intake rate was not influenced by differences in sward condition. Intake behaviour from both experiments was compared to establish the effect on herbage intake of changes in sward state and non‐sward factors. In Experiment 1, sward surface height, available herbage mass, proportion of leaf and green leaf mass declined as the day progressed. Bite mass declined with sward depletion, and mean intake rate was 1·64 kg dry matter (DM) h^–1, which was significantly lower at T3 (P < 0·01) than during other meals. In Experiment 2, plot sward conditions did not change throughout the day, and intake behaviour also remained constant, with a mean intake rate of 2·11 kg DM h^–1. Mean bite depth as a proportion of pregrazing extended tiller height was constant throughout the day (mean 0·32). The results show that, although cows grazed throughout the day on progressively depleted swards, indicative of rotationally grazed paddocks (Experiment 1), bite mass declined linearly and intake behaviour was variable. However, where intake was assessed on high‐quality, undefoliated swards (Experiment 2), intake behaviour was similar regardless of the time of day and the immediately previous experience. There was some indication of an interaction between the effects of the sward and the physiological condition of the animal on herbage intake.     

Sward dynamics of a smooth-stalked meadowgrass dominantwhite clover sward rotationally grazed by cattle and/or sheep

This experiment was carried out to study the responses of sward components (particularly white clover, Trifolium repens ) to grazing management in a natural sward dominated by smooth-stalked meadowgrass ( Poa pratensis ) syn. Kentucky bluegrass. Treatments during two grazing seasons (1989–90) were: cattle grazing alone (C); cattle grazing followed by topping (CT); cattle grazing followed by sheep grazing (CS); and sheep grazing alone (S). Mean target pre- and post-grazing herbage masses were 2200 and 1100 kg DM ha⁻¹, estimated by single-probe electronic capacitance meter. Sward component dynamics were monitored using turf dissections, marked white clover stolons, and ring-toss white clover leaf counts. Component and sward data for the C, CT, CS and S treatments respectively, were: number of white clover leaves m⁻², 1295, 1384, 1408, 900 (s.e. ± 108); number of leaves per growing point, 3·2, 3·4, 3·0, 2·8 (s.e. ± 0·2); herbage accumulation (t DM ha⁻¹), 5·16, 5·02, 5·87, 8·28 (s.e. ±0 08); rejected herbage (% pasture area) 39·7, 7·7, 16·0, 0 (s.e. ± 75); and annual net herbage production (t DM ha⁻¹) 3·39, 4·35, 4·99, 8·28 (s.e. ± 0.07). Swards grazed by sheep alone contained less white clover, but regrew quicker and produced more herbage than other treatments. Close topping or grazing by sheep following dairy cattle grazing decreased sward rejection by cattle. These treatments maintained more of the pasture in better condition for subsequent cattle grazing, resulting in greater net herbage production than where no post-cattle grazing treatment was used.     

The effect of grazed sward height and stocking rate on animal performance and output from beef cow systems

During three consecutive summers, forty spring-calving beef cows and their calves grazed perennial ryegrass-dominant swards receiving 250 kg N ha^?1 at one of two annual stocking rates [2.0 (SR 2.0) or 2.5 (SR 2.5) cows ha^?1] and one of two sward heights [4–5 (LS) or 7–8 (HS) cm] in a 2 × 2-factorial experiment, replicated twice. Sward heights were maintained from turn-out in spring by weekly adjustment of the area grazed and herbage was cut for silage in June and again in August from the areas not grazed. After the second cut of silage there was no control of sward height. Calves were weaned in early October and cows removed from pasture and housed when sward heights fell to 4 cm in autumn. Cows were fed in groups a variable but measured quantity of silage during winter to achieve a body condition score of 2.0–2.25 at turn-out the following spring. During the period of sward height control the cows on the HS treatment gained more live weight than those on the LS treatment (0.841 vs. 0.496 kg day^?1; P<0.01) as did the calves (1.167 vs. 1.105; P<0.05). but the stocking rate treatment had no effect. From the time of second-cut silage to the time of weaning and housing respectively, calves and cows gained more live weight on the SR 2.0 treatment because sward heights were higher. Reproductive performance of cows was not affected by treatment. The quantity of silage produced and consumed per cow was not affected by sward height treatment, but the SR 2.0 treatment produced significantly (P<0.001) more silage (1559 kg dry matter per cow) than the SR 2.5 treatment (833 kg dry matter per cow) and had higher winter silage requirements (1249 vs. 1153 kg dry matter per cow: P<0.05). The overall mean stocking rate at which winter fodder production and requirements would be in balance was calculated as 2.25 cows ha^?1 but values were 1.86, 2.60 and 2.28 in each of the three years of the experiment. The results showed that it was possible to control sward height in temperate beef cow systems by adjusting the area available for grazing. Body condition score can be used as a means of determining the feeding levels required to manipulate body condition of cows over winter to achieve prescribed levels of body condition. The experimental approach allows the identification of the stocking rate at which self-sufficiency in winter fodder can be achieved and the year-to-year variation associated with that stocking rate. This approach could be generalized if information on herbage growth rate were available, either from direct measurement or from predictive models.     

The effect of nitrogenous fertilizer on the photosynthesis of leaves of a ryegrass sward

The addition of nitrogenous fertilizer to a newly cut, N-deficient ryegrass sward resulted in an increased photosynthetic capacity in the next leaf to expand, showing that greater photosynthesis per unit leaf area as well as greater leaf area contributes to the increased growth of fertilized swards. Later in the 4-week growth period there was little difference in photosynthetic capacity between fertilized and unfertilized swards.     

Effects of duration and combination of drought and flood conditions on leaf photosynthesis,growth and sugar content in sugarcane

Global climate change will result in extreme environments, such as droughts and floods. We investigated the individual and combined effects of droughts and floods of varying duration on sugarcane (Saccharum spp.) growth using a pot experiment under glasshouse conditions with the following six treatments: drought for 15 d, prolonged drought for 30 d, flood for 15 d, prolonged flood for 30 d, short flood followed by prolonged drought, and prolonged flood followed by prolonged drought. Plants that were subjected to drought conditions, including drought after a flood, had reduced CO₂ assimilation (through stomatal closure) and leaf areas, whereas flood conditions showed no effect. During flooding, some roots died, and adventitious roots with well-developed aerenchyma appeared from the submerged nodes. At the time of harvest, there were no significant differences in stem fresh weight, sucrose content, or sugar yield between the treatments. However, ion content analysis revealed that flood conditions caused an accumulation of sodium in the bottom of stems and adventitious roots. Therefore, under flood conditions, plants may develop adventitious roots, which may offset the negative effects of root death, helping them to maintain their growth and yield.     

White clover growth patterns during the grazing season in a rotationally grazed dairy pasture in New York

White clover (Trifolium repens L.) is an important stoloniferous pasture legume in the Great Lakes region of the United States, but it often has limited persistence. Researchers in New Zealand and Wales have found that in spring, compared with other seasons, white clover plants have reduced branching complexity and have the fewest buds that produce leaves. They therefore suggested that in spring the plants are most vulnerable to grazing and climatic stress. Because of severe winter and cool, wet spring weather in New York State, it was hypothesized that white clover plants would also be of low branching complexity, smaller and have low axillary bud activity in spring compared with later in the grazing season. To test this, growth of white clover was monitored in an orchard grass (Dactylis glomerata L.)/white clover pasture in New York that was rotationally grazed with dairy cows during the 1993 and 1994 grazing seasons. Three sets of plants were sampled. The first set consisted of forty random plants sampled before each grazing event. Stolon branching order, number of each stolon branching type and area the plant occupied were determined. Approximately each month before one grazing event, a separate set of 32 random plants was measured in the field to determine the area they occupied; these plants were then removed to the laboratory for the measurement of stolon order, number of each stolon type, stolon lengths, total number of growing points, number of taproots and adventitious roots, root position and above-ground dry matter. Once a month, 12 additional plants were removed to measure axillary bud activity at each node. Leaf development from nodes tended to increase from spring to summer. However, the stolon branching order of white clover plants was not simpler in spring compared with summer or autumn. In 1994 during and after a dry and hot period, white clover plants were smaller, of lower stolon branching order and had fewer roots. Climate and associated soil organism activity appear to explain the different white clover growth patterns observed in New York and New Zealand. Severe winters in New York limit earthworm activity and stolon burial, which is important in contributing to stolon/plant breakdown in New Zealand. During the years of this study in New York, a hot and dry period had the most negative effect on the growth pattern of white clover.     

The effects of sward height and nitrogen fertilizer application on changes in sward composition, white clover growth and the stock carrying capacity of an upland perennial ryegrass/white clover sward grazed by sheep for four years

Perennial ryegrass/white clover pastures grazed by sheep and receiving either no fertilizer N (No) or 120 kg N ha^?1 year^?1 (N₀) were maintained with surface heights of 2·5, 3·5 and 5·0 cm for over four years. The treatments were replicated. The white clover (WC) population was greatest in the N₀treatment, and declined during the study. Between-year variation in WC was negatively related to rainfall and positively related to temperature, WC as a proportion of the total plant population decreased during the summer in the Nl treatment. The perennial ryegrass (PRG) population was greater in the Nl treatment, declined during the study and both within and between years was positively related to temperature. The population density of the unsown grasses was highest in the N-fertilized treatment and in the swards maintained at the lowest heights (these treatments also had the highest stocking rate); it increased during the study, within-years being positively related to temperature and between-years being positively related to rainfall. The WC stolon extension rate was largely unaffected by N fertilizer application and was greatest in the taller swards. Leaf appearance rate was unaffected by N fertilizer application and sward height; it was positively related to temperature and negatively related to rainfall. Branching rate was greater in the N₀ treatment with significant sward height effects confined to a negative relationship with local sward height within treatment plots on one occasion; it was negatively related to rainfall. The ground level red:far red light ratio was negatively related to local sward height. The total live weight of sheep carried in the No treatments was 0·7 of that in the N₁ treatments. Expected photomorphogenic responses by we were confined to stolon extension. It was concluded that on the poorly drained clay-loam soil used in this study the effects of sheep, in interaction with climatic factors, had an overriding effect on clover branching rate and the ultimate species composition.     

The effects of regrowth and maintenance height on a grass sward with a high density of tillers

A perennial ryegrass sward was grazed by sheep in April 1993 to a target sward surface height (SSH) of 3 cm to create a high density of grass tillers. From 3 May, the sheep were removed and small plots were established on the sward, when the average tiller density (± s.e.) was 35 900 ± 420 live tillers m⁻². Different regrowth treatments were then imposed by allowing plots to regrow to target SSHs of 6 cm (18 g dry matter (DM) M⁻²), 9 cm (78 g DM m⁻²), 12 cm (132 g DM m⁻²) or 15 cm (197 g DM m⁻²). The plots were then maintained by cutting at either 6 or 9 cm SSH until the end of the experiment on 30 September. Live tiller density was reduced by regrowth beyond 8.9 cm (78 g DM m⁻², P <0.001) and leaf-stem ratio and in vitro organic matter digestibility were reduced by regrowth beyond 6.1 cm (18 g DM m⁻², P < 0.05). The effect on live tiller density was sustained through the remainder of the season. From the beginning of June to the end of September, maintenance of SSH at an average of 9.1 cm compared with 6.4 cm also resulted in lower live tiller density, live-dead tiller ratio and leaf-stem ratio and higher herbage mass (at least P < 0. 05). There were significant interactions between regrowth SSH and maintenance SSH, so that leaf-stem ratio, live-dead tiller ratio and live tiller density were reduced by regrowth to a SSH of 16–4 cm followed by maintenance at 9.1 cm, compared with regrowth to 6.1 cm and maintenance at 6.4 cm.     

Factors affecting sonic sward stick measurements: the effect of different leaf characteristics and the area of sward sampled

Laboratory experimentation found smooth and pubescent leaf surfaces to behave as specular (mirror-like) reflectors of ultrasound, whilst ribbed leaves behaved as an array of reflectors. It was concluded that swards composed of species with smooth, erectophile leaves would reflect ultrasound poorly and that this may explain the low sward height measurements returned by the sonic sward stick in the field when compared with the HFRO sward stick. The likely effect of species with planophile leaves such as clover is discussed. The sward surface influencing a sonic sward stick measurement was calculated to subtend a solid angle of about 43° at the transducer, equivalent to an area of about 1000 cm² with the transducer mounted at a height of 50 cm. However, measurements made over turves of varying sizes or over turves with cores of varying size removed found the sensitivity decreased very rapidly beyond an inner area subtending a solid angle of about 7° this is equivalent to an area of about 30 cm².     

Diet selection by sheep and goats and sward composition changes in a ryegrass/white clover sward previously grazed by cattle, sheep or goats

An experiment was conducted to examine how variation in the composition and structure of mixed grass/white clover swards affected diet selection by sheep and goats. Sward composition in a mixed perennial ryegrass/white clover sward was manipulated by continuous grazing from 28 May to 28 July (pre-experimental phase) with cattle, sheep or goats, and then from 29 July to 2 September (experimental phase) with sheep or goats in a factorial design replicated twice. Sward surface height was maintained at 6 cm by regular adjustment of stocking density. Grazing by different sequences of animal species resulted in significant differences in the proportions of white clover in the sward, and especially in the proportion of clover lamina and petiole. Grazing by goats in the pre-experimental phase led to greater proportions of clover lamina and petiole in the whole sward and the sward surface. The proportion of white clover in the diet selected by sheep in the experimental phase was consistently higher than that in the sward as a whole, but was closely related to that near to the sward surface (approximately the top 2 cm). For goats there was no significant relationship between the proportion of clover in the diet and in the whole sward, and they generally selected a diet with a lower proportion of white clover than was present in approximately the top 2 cm of the sward. It is concluded that on mixed grass/white clover swards goats do not graze as deeply into the sward as sheep and that this results in a lower proportion of white clover in their diet and therefore allows higher proportions of white clover to develop under grazing by goats than by sheep.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK. 5. The effect of stocking rate and reduced levels of nitrogen fertilizer

The implications for UK upland sheep systems of reducing nitrogen fertilizer application to perennial ryegrass/white clover swards were studied over 3 years. Sward height (3·5–5·5 cm) was controlled for ewes with lambs until weaning using surplus pasture areas for silage; thereafter, ewes and weaned lambs were grazed on separate areas, and sward height was controlled by adjusting the size of the areas grazed and using surplus pasture areas for silage if necessary. Combinations from three stocking rates [10, 6 and 4 ewes ha⁻¹ on the total area (grazed and ensiled)] and four nitrogen fertilizer levels (150, 100, 50 and 0 kg ha⁻¹) provided six treatments that were replicated three times. Average white clover content was negatively correlated with level of nitrogen fertilizer. The proportion of white clover in the swards increased over the duration of the experiment. Control of sward height and the contribution from white clover resulted in similar levels of lamb liveweight gain on all treatments. All treatments provided adequate winter fodder as silage. It is concluded that the application of nitrogen fertilizer can be reduced or removed from upland sheep pastures without compromising individual animal performance provided that white clover content and sward height are maintained. Resting pastures from grazing by changing ensiled and grazed areas from year to year sustained white clover content over a 3-year period.     

The effects of grazing with and without excretal returns on the accumulation of nitrogen by ryegrass in a continuously grazed upland sward

Rates of N accumulation were studied on sheep grazed grass swards maintained at a constant height of 5 cm for two growing seasons (1985 and 1986) and receiving no N fertilizer using a tiller tissue turnover technique. Grazing with normal excretal returns resulted in an 85-105% increase in the estimated rates of N accumulation by laminae compared with similarly grazed swards where excretal returns were prevented. Generally, increases in tiller numbers in plots receiving excreta were mainly responsible for the increased rates of N accumulation, rather than increased rates of N accumulation per tiller. Rates of N remobilization from laminae and senescence (flux of N to standing-dead litter pool) per tiller, and rates of leaf extension and leaf appearance/disappearance were unaffected by excretal returns, in urine patches increased tiller numbers and increased rates of accumulation per tiller both resulted in greater estimates of N accumulation compared with tillers not visibly affected by excreta. However, at any one time only about 11% of the plot area was affected by excreta and increases in N accumulation in excreta patches could not account for the overall plot increases in rates of N accumulation.
The increased tillering in plots receiving excreta may have increased exploitation of the soil N. With an average stocking rate of 15 sheep ha⁻¹, soil nitrate-N, but not total N (Kjeldahl) nor bulk density, was increased during the winter of the second year as a result of excretal returns.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK: 6. The effect of reducing stocking rate and application of fertilizer nitrogen in Wales

Abstract The implications for the agricultural productivity of the UK upland sheep systems of reducing nitrogen fertilizer application and lowering stocking rates on perennial ryegrass/white clover swards were studied over 4 years at a site in Wales. The system involved grazing ewes and lambs from birth to weaning on swards maintained at a constant height with surplus herbage made into silage, thereafter ewes and weaned lambs grazed on separate areas until the onset of winter with adjustments to the size of the areas grazed and utilizing surplus pasture areas for silage. Four stocking rates [SR 18, 15, 12 and 9 ewes ha^?1 on the total area (grazed and ensiled)] and two levels of annual nitrogen fertilizer application (N 200 and 50 kg ha^?1) were studied in five treatments (N200/SR18, N200/SR15, N50/SR15, N50/SR12 and N50/SR9). Average white clover content was negatively correlated with the level of annual nitrogen fertilizer application. White clover content of the swards was maintained over the duration of the experiment with an increasing proportion of clover in the swards receiving 50 kg N ha^?1. Control of sward height and the contribution from white clover resulted in similar levels of lamb liveweight gain from birth to weaning in all treatments but fewer lambs reached the slaughter live weight by September at the higher stocking rates and with the lower level of fertilizer application. Three of the five treatments provided adequate winter fodder as silage (N200/SR15, N50/SR12 and N50/SR9). Because of the failure to make adequate winter fodder and the failure of white clover to fully compensate for reduction in nitrogen fertilizer application, it is concluded that nitrogen fertilizer can only be reduced on upland sheep pastures if accompanied by reduced stocking rates.     

Frequency of stocking rate adjustments required on contrasting upland perennial ryegrass pastures continuously grazed to a sward height criterion from May to July

The use of sward height as a criterion for determining the time and extent of stocking-rate changes on continuously grazed swards was investigated over a 2-year period (1985–86) in a sheep production experiment. Swards of three contrasting perennial ryegrass ( Lolium perenne L.) varieties were established with and without Aberystwyth S184 small-leaved white clover ( Trifolium repens L.) at an upland site (310–363 m) in mid-Wales. From spring (late April) until weaning (mid-July) the pastures were continuously stocked with Beulah Speckled Face ewes and Suffolk cross lambs. During this period sward heights of 4 ± 0.5 cm were obtained and maintained by regular adjustment of animal number on individual paddocks. Grass-only swards received 160 and 200 kg N ha⁻¹ and the grass clover swards were given 80 and 75 kg N ha⁻¹ in 1985 and 1986 respectively.
Differences were observed between the treatments in sward height profiles over the season necessitating contrasting adjustments to stocking rates. Mean stocking rate necessary on early flowering Aurora (22 6 ewes ha⁻¹) was respectively 27% and 17% higher than on late-flowering Aberystwyth S23 and Meltra (tetraploid) ryegrasses; mean stocking rate on grass-only swards was 19% higher than on the grass-clover pastures.
It is concluded that sward height is a useful criterion on which to make adjustments to stocking rates to compare the potential performance of contrasting swards, under continuous grazing. The infrequent adjustments required to maintain a constant sward height, especially on the late flowering diploid perennial ryegrass variety on which many upland pastures are based, suggest that the criterion of sward height could be successfully employed on farms as an aid to efficient grazing management.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK. 4. The effect of seasonal pattern of nitrogen fertilizer application and annual stocking rate

Greyface (Border Leicester × Scottish Blackface) ewes, mated in October, were housed in winter and turned out after lambing in late March to one of four treatments replicated three times on perennial ryegrass (Lolium perenne)-dominated pastures. On three of the treatments the stocking rate was 12·5 ewes plus lambs ha^?1 annum^?1 (SR_12·5), whereas on the fourth it was 15·0 ewes plus lambs ha^?1 annum^?1 (SR_15·0). There were two levels of applied fertilizer nitrogen (N), 152 kg N ha^?1 annum^?1 (N₁₅₂) and 205 kg N ha^?1 annum^?1 (N₂₀₅). Three patterns of nitrogen application were used: predominantly in spring (E), predominantly in autumn (L), and mainly in spring and autumn (E + L). Grazed sward surface height was controlled within the range 3·5–5·5 cm during spring and summer, and supplementary feed was offered when sward height was below 3·5 cm. The effects on animal performance, yield of silage and requirement for supplementary feed were measured over 3 years (1986–88). Management of the sward height within the specified limits resulted in similar levels of individual animal performance for all treatments, but treatment SR_15·0N₂₀₅E + L produced a significantly (P < 0·05) greater output of lamb per hectare. There was no evidence to suggest that, by increasing the amount of N fertilizer applied in the autumn, ewe reproductive performance was increased. Neither the treatment with the lower level of N applied in the spring (SR_12·5N₁₅₂L) nor the high stocking rate treatment (SR_15·0N₂₀₅E + L) was self-sufficient in winter feed (silage) production. SR_15·0N₂₀₅E + L also required more supplementary feed in both the lactation (38·1 kg ewe^?1) and the mating (9·1 kg ewe^?1) periods. SR_12·5N₁₅₂L required the second greatest amount of supplementary feed during lactation (36·1 kg ewe^?1), whereas SR_12·5N₂₀₅E + L required the least supplementary feed over both periods (27·8 kg and 4·8 kg ewe^?1). Taking all treatments together, there were significant (P < 0·05) differences between years in lamb weaning weight (kg), lamb output (kg ha^?1), yield of silage (kg ewe^?1) and supplementary feed required (kg ewe^?1), arising mainly from a lower level of herbage production in one year. Treatment SR_12·5N₁₅₂E generated the highest gross margin per hectare.     

Canopy height and N affect herbage accumulation and the relative contribution of leaf categories to photosynthesis of grazed brachiariagrass pastures

The effects of three canopy heights (10, 25 and 40 cm) and two N rates (50 and 250 kg ha^?1 year^?1) on herbage accumulation (HA), participation of leaf categories in the leaf area index (LAI) and photosynthesis of grazed “Mulato II” brachiariagrass (Brachiaria brizantha × B. decumbens × B. ruziziensis) were investigated during two summer growing seasons in Piracicaba, São Paulo, Brazil. The HA and LAI increased linearly from 8560 to 13600 kg DM ha^?1 year^?1 and 2.3 to 5.5 m² m^?2, respectively, as canopy height increased. Mulato II brachiariagrass maintained at 10 cm canopy height showed greater proportion of young leaves, but was less productive than taller canopies. Taller canopies had greater proportion of mature leaves in the LAI as well as greater photosynthesis. The greater N rate contributed to increased LAI, leaf and canopy photosynthesis rates and HA but did not affect the LAI composition. The total leaf area is more important to HA than the proportion of young leaves in the LAI, as taller canopies were more productive, suggesting that maintaining Mulato II brachiariagrass at 25 or 40 cm is advantageous for this grass.     

The effects of stocking rate and nitrogen fertilizer on a perennial ryegrass-white clover sward

The effects of stocking rate and N fertilizer on a mixed sward of perennial ryegrass ( Lolium perenne) and white clover ( Trifolium repens ) set-stocked with sheep were examined. Sward production and composition, and sheep production were studied.
Increasing the stocking rate over the range 25–55 yearling sheep ha⁻¹ reduced herbage accumulation by about 40%, whether or not N fertilizer was applied. Increasing the stocking rate increased the density of ryegrass tillers, but reduced the density of clover stolons and the clover content of the swards. Applications of N fertilizer (200 kg N ha⁻¹ a⁻¹) increased herbage accumulation by about 20% but substantially reduced the clover content.
Liveweight gain per animal and per unit area were greater at the lower stocking rates where the clover content and nutritive value of the diet were greatest. Wool growth per unit area was greater at the higher stocking rates. Applications of 1M fertilizer increased liveweight gain at stocking rates above 25 sheep ha⁻¹, but had no effect on wool production at any stocking rate.
The results demonstrate that a stable and productive grass-clover association was maintained under conditions of set-stocking at around 23 yearling sheep ha⁻¹and that at this stocking rate, which appears to be about the biological optimum, there was no advantage in using N fertilizer.     

The effect of stocking rate and calving date on grass production,utilization and nutritive value of the sward during the grazing season

A 2‐year whole‐farm system study compared the accumulation, utilization and nutritive value of grass in spring‐calving grass‐based systems differing in stocking rate (SR) and calving date (CD). Six treatments (systems) were compared over two complete grazing seasons. Stocking rates used in the study were low (2·5 cows ha^?1), medium (2·9 cows ha^?1) and high (3·3 cows ha^?1), respectively, and mean CDs were 12 February (early) and 25 February (late). Each system had its own farmlet of eighteen paddocks and one herd that remained on the same farmlet area for the duration of the study. Stocking rate had a small effect on total herbage accumulation (11 860 kg DM ha^?1 year^?1), but had no effect on total herbage utilization (11 700 kg DM ha^?1 year^?1). Milk and milk solids (MS; fat + protein) production per ha increased by 2580 and 196 kg ha^?1 as SR increased from 2·5 to 3·3 cows ha^?1. Milk production per ha and net herbage accumulation and utilization were unaffected by CD. Winter feed production was reduced as SR increased. Increased SR, associated with increased grazing severity, resulted in swards of increased leaf content and nutritive value. The results indicate that, although associated with increased milk production per ha, grazed grass utilization and improved sward nutritive value, the potential benefits of increased SR on Irish dairy farms can only be realized if the average level of herbage production and utilization is increased.     

Sward height, structure and leaf extension rate of Lolium perenne pastures when grazed by overwintering barnacle geese

Abstract The grazing of agricultural pastures during winter and spring by geese is considered an important agricultural problem in parts of the U.K. This study describes the sward structure, leaf extension and senescence rates of Lolium perenne‐dominated pastures that are frequently grazed by barnacle geese (Branta leucopsis) during winter in South‐west Scotland, as well as the conducting of a field experiment that simulated grazing to investigate the effects of defoliation. Gross leaf extension and senescence rates were strongly related to temperature, daylength and average tiller size, resulting in positive values of net leaf extension rate per tiller for most of the winter. Total tiller length declined from November to January but increased from January to April. Sward height, however, declined consistently from October to April, suggesting that swards were becoming trampled by repeated visits by flocks of geese over this time. The structure of individual tillers was found to vary slightly over the winter, with tillers becoming more dominated by younger leaves towards the end of the winter. Experimental defoliation of tillers suggested that absolute leaf extension rates did not respond in an under‐ or over‐compensatory manner, even when tillers were nearly completely defoliated. The results suggested that sward structure and leaf extension rates are not unduly affected by repeated grazing by overwintering geese and that short‐term depletion and trampling are the main impacts.