Weed interference in maize, cowpea and maize/cowpea intercrop in a subhumid tropical environment. II. Early growth and nutrient content of crops and weeds

Early growth and nutrient content of crops and weeds from weed-free and weedy no-tillage maize (Zea mays L, cv. TZB), cowpea (Vigna unguiculata (L) Walp. cv. VITA-5) and maize/cowpea intercrop at populations of 40000, 50000 and 30000 + 40000 plants ha^?1 grown on a loamysand Oxic Ustropept in a subhumid tropical location were monitored in the early and late 1979 cropping seasons. In the first 6 weeks of growth in the early season, cropping pattern had no effect on weed growth; weeds did not suppress crop growth significantly until 5–6 weeks after sowing and total crop dry weights were not affected by cropping pattern. Three weeks after sowing, weeds from weedy crop plots had taken up two to four times as much nutrient (N, P, K, Ca + Mg) as was taken up by corresponding weed-free crops. In the late season, weed dry weight 6 weeks after sowing was depressed in the intercrop compared to monocultures and dry-matter production of the intercrop was higher than those of monocultures. The resource use index (RUI), defined as the amount of an environmental resource used by a weed-free crop divided by the combined amount of the same resource used by the corresponding weedy crop and the associated weeds, increased with age of crop and was higher for the intercrop than the monocultures only in the late season.     

Critical period of weed competition in three vegetable crops in relation to management practices

The critical period of weed competition was determined in three vegetable crops: early cabbage (Brassica oleracea var. capitata L.), pickling cucumbers (Cucumis sativus L.), and field-seeded processing tomatoes (Lycopersicon esculentum L.). There were significant interactions between weed-removal treatments, year, and row width. Cabbage yields were reduced if plots were not kept weed-free for at least 3 weeks after transplanting or if weeds which emerged with the crop were allowed to remain longer than 4–5 weeks, Cucumber yields were reduced if plots were not kept weed-free for up to 4 weeks after seeding or if plots remained weed-infested longer than 3–4 weeks. Higher crop population densities (narrower row widths) in cabbage and cucumbers resulted in smaller plants, earlier competition from weeds, and therefore a shorter period that the crop could remain weed-infested without suffering reduced yields. Yields of direct-seeded tomatoes were reduced if plots were not kept weed-free for up to 9 weeks after seeding or if weeds which emerged with the crop were allowed to remain longer than 5 weeks. In each crop the timing of the critical period of competition was verified by weed removal only during this interval. There was a true critical period in direct-seeded tomatoes, but not in cabbage or cucumbers where a single weeding was sufficient to prevent yield losses.     

Comparison of crop and weed height,for potential differentiation of weed patches at harvest

Weeds and weed control are major production costs in global agriculture, with increasing challenges associated with herbicide‐based management because of concerns with chemical residue and herbicide resistance. Non‐chemical weed management may address these challenges but requires the ability to differentiate weeds from crops. Harvest is an ideal opportunity for the differentiation of weeds that grow taller than the crop, however, the ability to differentiate late‐season weeds from the crop is unknown. Weed mapping enables farmers to locate weed patches, evaluate the success of previous weed management strategies, and assist with planning for future herbicide applications. The aim of this study was to determine whether weed patches could be differentiated from the crop plants, based on height differences. Field surveys were carried out before crop harvest in 2018 and 2019, where a total of 86 and 105 weedy patches were manually assessed respectively. The results of this study demonstrated that across the 191 assessed weedy patches, in 97% of patches with Avena fatua (wild oat) plants, 86% with Raphanus raphanistrum (wild radish) plants and 92% with Sonchus oleraceus L. (sow thistles) plants it was possible to distinguish the weeds taller than the 95% of the crop plants. Future work should be dedicated to the assessment of the ability of remote sensing methods such as Light Detection and Ranging to detect and map late‐season weed species based on the results from this study on crop and weed height differences.     

THE INFLUENCE OF IRRIGATION, FERTILIZING AND MCPA ON THE COMPETITION BETWEEN SPRING CEREALS AND WEEDS

Summary. The average air-dry yield of weeds on fallow plots was about four times as large (2660 kg/ha) as the yield of weeds which developed among spring cereal crops (740 kg/ha). Application of fertilizers increased the number of weeds and their average weight on both irrigated and non-irrigated plots. Treatment with MCPA reduced the weed yield to 36% on fallow plots and to 26% on weedy cereal plots. It was more effective on fertilized plots than on unfertilized plots, but irrigation made little difference to its effectiveness. The space left by the destroyed weeds was mainly occupied by Agropyron repens . Oats were better able to compete with weeds than were wheat and barley. On unfertilized and non-irrigated plots spraying with MCPA (1-26 kg/ha) at the beginning of shooting significantly reduced the yields of weed-free barley. The yields of sprayed oats and wheat were also below the yields from the unsprayed although the reductions were not significant at the 5% level. However, on a number of fertilized and irrigated plots, both weedy and hand-weeded , on which the plants had better growing conditions, MCPA resulted in a significant increase in the grain yield of barley and oats.
Influence de l'irrigation, de la fertilisation et du MCPA sur la compétition entre céréales de printemps et mauvaises herbes     

Competition between annual weeds and vining peas grown at a range of population densities: effects on the weeds

Linear regression of dry weight of weeds against crop density, together with the use of diversity indices and principal component analysis were used to derive information about changes in the behaviour of annual weeds over the growing season and in response to a wide range of crop densities in vining peas Pisum sativum L. Using linear regression it was possible to quantify reductions in weed dry weights per unit increase in crop plant density The ‘weed losse’ factor was acceptably consistent between experiments. Indices examining richness and evenness showed that numbers of weed species declined with increasing crop density and as the season progressed, but although species evenness became less at successive sampling dates the presence of a pea crop, whatever its density, did not radically alter the composition of the weed flora. Principal component analysis demonstrated that although there was competition within the weed flora, the crop did not replace the dominant weed species on high density plots, but reduced growth of all weed species alike.     

Weed competition in spring-planted strawberries

Annual weeds germinating after planting strawberry (Fragaria chiloensis (L.) Duch. cv. Cambridge Favourite) in late March had no effect on crop growth if removed by late May. Dense weed cover thereafter severely inhibited stolon growth, virtually eliminating it if allowed to remain beyond mid-August. Shading by weeds inhibited leaf production and caused etiolation of existing leaves. New leaves appeared shortly after weed removal and few plants died unless weed cover persisted for most of the growing season. Further weed germination was allowed on some plots. Although not removed until late autumn, these weeds only had adverse effects on crop growth where initial weeding had occurred before mid-June. In one experiment, delaying weed removal until 6 July, 31 August or 2 November in the first year reduced fruit yield in the second (weed-free) year by 34%, 54% and 67% respectively. In a later experiment, competition from weeds until July or later in the first growing season gave fruit yields similar to those in the first experiment, but totally weed-free plots and those kept clean after weeding in mid-June produced less fruit than plots which remained weedy between mid-June and mid-August. It is suggested that competition from uncontrolled stolon growth in this experiment severely inhibited crown and hence truss production on plots which did not suffer weed competition. Unless left untouched until early September, weeds had less adverse effect on truss production than the stolons which they displaced. The results are interpreted in relation to improving the efficiency of weed control techniques.     

Response of onion (Allium cepa L.) plants to fertilizers, weed competition duration, and planting times in the central Jordan Valley

The competitive ability of onion with weeds and the effects of weeds on onion crop growth and yield were investigated in three field experiments carried out in the central Jordan Valley during the 1997–1998 growing season. Fertilizer application to the onion plants in the presence of weeds significantly lowered the onion yield and reduced the bulb diameter, compared to the unfertilized, weed-infested onion crop. Weed competition and onion yield reduction increased with an increase in the fertilizer rate. All weed-free treatments with or without fertilizer gave a significantly higher onion yield than any of the fertilized, weedy plots. The highest yield of onion was obtained from the weed-free plots with a moderate application of fertilizer. Weed competition reduced the growth, bulb yield, and size of onion in the plots established from bulbs, seedlings, and seeds when compared to the relevant weed-free controls. The effect increased with competition duration and was more pronounced in direct-seeded onion than in the other two types of planting material. However, onion grown from bulbs or seedlings gave better growth, bulb weight, and number and was more competitive than the direct-seeded onion. The results showed that the minimum number of days of weed competition needed for a significant reduction in onion growth was 42 days when propagated from bulbs or seedlings and 21 days after direct-seeding. Onion grown from bulbs tolerated weed competition better and produced a higher bulb yield than that obtained from direct-seeded or transplanted onion, irrespective of the planting dates. December was found to be the best planting time for onion that was grown from bulbs and seedlings, while November was the best planting time for direct-seeded onion, provided the plots were weed-free throughout the growing season.     

Critical period of weed interference in chickpea

The critical period of weed interference in one variety of chickpea was determined in field experiments carried out at two sites, Tabriz 2002 and Kermanshah 2003, Iran. Chickpea culture was either kept free of weeds for 0, 12, 24, 36, 48 and 60 days after crop emergence (DAE) or weeds were allowed to grow for 0, 12, 24, 36, 48 and 60 DAE. In these experiments, chickpea yield increased with increasing duration of weed-free period and was reduced by increasing duration of weed-infested period. Unweeded conditions for the entire growing season caused 66.4% and 48.3% seed yield reduction when compared with the treatment that was weed-free throughout the growing season, at Tabriz 2002 and Kermanshah 2003, respectively. The results indicated that chickpea must be kept weed-free between the five-leaf and full flowering stages (24–48 DAE) and from the four-leaf to beginning of flowering stages (17–49 DAE) at the two sites, respectively, in order to prevent >10% seed yield loss. At both sites, reduction in seed yield, because of the increased weed interference period, was accompanied by simultaneous reduction in plant dry weight, number of branches, pods per plant and 100-seed weight. This was supported by significant and positive correlations between these traits and chickpea seed yield. There was no significant correlation between the number of seeds per pod and seed yield. A linear regression model was used to describe the relationship between weed dry weight and seed yield loss.     

Predicting the growth and competitive effects of annual weeds in wheat

The growth and competitiveness of 12 annual weed species were studied in crops of winter wheat, in which weeds were sown to give a wide range of plant densities. Weed growth patterns were identified; early species which senesced in mid-summer were less competitive than those with a growth pattern similar to that of the crop. Most species had little effect on crop yield in 1987, and this was attributed to a high crop den sity. Crop yield-weed density relationships for all species in 1988 and for Galium aparine in 1987 were well described by a rectangular hyperbola. Species were listed in the following competitive order based on the percentage yield loss per weed m^?2: Avena fatua > Matricaria perforata > Galium aparine > Myosotis arvenis > Poa trivialis > Alopecurus myosuroides > Stellaria media > Papaver rhoeas > Lamiumpur-pureum > Veronica persica > Veronica hederi-folia > Viola arvensis. Prediction of yield loss is discussed. The assumptions inherent in using Crop Equivalents (based on relative weights of weed and crop plants), are challenged; with intense competition, weed biomass at harvest failed to replace lost crop biomass, and harvest index was reduced. It is concluded that a competi tive index, derived from yield density relation ships, and expressed as the percentage yield loss per weed m^?2, is more likely to reflect the com petitive ability of a species than an index obtained from plant weights in the growing crop.     

Competition of sorghum cultivars and densities with Japanese millet (Echinochloa esculenta)

Field studies were conducted at two locations in southern Queensland, Australia during the 2003–2004 and 2004–2005 growing seasons to determine the differential competitiveness of sorghum (Sorghum bicolor L. Moench) cultivars and crop densities against weeds and the sorghum yield loss due to weeds. Weed competition was investigated by growing sorghum in the presence or absence of a model grass weed, Japanese millet (Echinochloa esculenta). The correlation analyses showed that the early growth traits (height, shoot biomass, and daily growth rate of the shoot biomass) of sorghum adversely affected the height, biomass, and seed production of millet, as measured at maturity. “MR Goldrush” and “Bonus MR” were the most competitive cultivars, resulting in reduced weed biomass, weed density, and weed seed production. The density of sorghum also had a significant effect on the crop's ability to compete with millet. When compared to the density of 4.5 plants per m², sorghum that was planted at 7.5 plants per m² suppressed the density, biomass, and seed production of millet by 22%, 27% and 38%, respectively. Millet caused a significant yield loss in comparison with the weed‐free plots. The combined weed‐suppressive effects of the competitive cultivars, such as MR Goldrush, and high crop densities minimized the yield losses from the weeds. These results indicate that sorghum competition against grass weeds can be improved by choosing competitive cultivars and by using a high crop density of >7.5 plants per m². These non‐chemical options should be included in an integrated weed management program for better weed management, particularly where the control options are limited by the evolution of herbicide resistance.     

Improved management of Avena ludoviciana and Phalaris paradoxa with more densely sown wheat and less herbicide

Summary The effectiveness of crop competition for better weed control and reducing herbicide rates was determined for Avena ludoviciana and Phalaris paradoxa . Four experiments, previously broadcast with seeds of the two weeds in separate plots, were sown with three wheat densities, and emerged weeds were treated with four herbicide doses (0–100% of recommended rate). The measured crop and weed traits were first analysed across experiments for treatment effects. Grain yield and weed seed production data were then analysed using cubic smoothing splines to model the response surfaces. Although herbicide rate for both weeds and crop density for P. paradoxa had significant linear effects on yield, there was a significant non-linearity of the response surface. Similarly, herbicide rate and crop density had significant linear effects on weed seed production, and there was significant non-linearity of the response surface that differed for the weed species. Maximum crop yield and reduction in seed production of P. paradoxa was achieved with approximately 80 wheat plants m⁻² and weeds treated with 100% herbicide rate. For A. ludoviciana , this was 130 wheat plants m⁻² applied with 75% herbicide rate. Alternatively, these benefits were achieved by increasing crop density to 150 plants m⁻² applied with 50% herbicide rate. At high crop density, application of the 100% herbicide rate tended to reduce yield, particularly with the A. ludoviciana herbicide, and this impacted adversely on the suppression of weed seed production. Thus, more competitive wheat crops have the potential for improving weed control and reducing herbicide rates.     

The critical period of weed control in double-cropped soybean

Double-cropped soybean after winter cereals is the most common soybean production system in the eastern Mediterranean region of Turkey. Weeds are among the main obstacles to double-cropped soybean. A study was conducted in 2002 and 2003 to determine the critical period of weed control (CPWC) for double-cropped soybean. The treatments consisted of either allowing weeds to infest the crop for increasing durations after sowing, or maintaining plots weed-free for increasing durations after sowing. The Gompertz and logistic equations were fitted to relative yields representing the critical weed-free period and the critical time of weed removal, respectively. Johnsongrass (Sorghum halepense), common cocklebur (Xanthium strumarium) and field bindweed (Convolvulus arvensis) were the dominant weeds. For 5% crop loss level, the CPWC was almost all season long, whereas it was from V1 to R6-R7 growth stages for 10% yield loss level. These findings indicate that pre-sowing or pre-emergence control methods should be applied in the region to avoid greater crop losses.     

The role of models for multicriteria evaluation and multiobjective design of cropping systems for managing weeds

Weeds are both harmful for crop production and important for biodiversity, while herbicides can pollute the environment. We thus need new cropping systems optimising all cultural techniques, reconciling agricultural production, herbicide reduction and biodiversity conservation. Here, we show how to (i) develop models quantifying the effects of cropping systems on weed dynamics, (ii) integrate interactions between weeds and other organisms, (iii) predict the impact on production and biodiversity and (iv) use the model for multicriteria evaluation and multiobjective design of cropping systems. Among the existing weed dynamics models, we chose the one closest to our requirements to illustrate these different steps, that is, FlorSys which predicts multispecific weed dynamics as a function of cultural techniques and pedoclimate. We have illustrated the development of interaction submodels with the example of a crop pathogen whose propagation is increased when infecting grass weeds. To evaluate the weed flora impact, predicted weed densities were translated into indicators of harmfulness (crop yield loss, technical harvest problems, harvest pollution, field infestation, crop disease increase) and biodiversity (weed species richness and equitability, trophic resources for birds, insects and pollinators). Simulations were run over several years and with different weather scenarios (i) to optimise cultural techniques to control harmful weeds, (ii) to analyse the impact of changing agricultural practices (e.g. simplified tillage and rotations, no‐till, temporary crops) on weed density, species and trait composition and (iii) to evaluate cropping systems for their ability to reconcile agricultural production and biodiversity, thus identifying levers for designing sustainable cropping systems.     

Problems of herbicide assessment in peanuts in Jamaica

Delays of 40 or 60 days in weeding peanuts (Arachis hypogaea L.) gave lower yields and fewer pods per plant but did not affect seed number per pod. Yield, pod number per plant, seeds per pod and mean weight per seed were all less in unweeded plots. In three herbicide trials crop yields were not closely correlated with crop vigour scores made several weeks before crop maturity, or with weediness scores. Correlations between yields and weed dry weights were better, but certain high-yielding herbicide treatments gave poor weed control and in some cases had low crop vigour scores. Most yield differences reflected differences in pod number per plant, with additional smaller compensating or additive effects on the other components. Certain herbicides apparently gave high mean seed weights without diminishing seed numbers per pod. Pre-emergence alachlor was the most promising herbicide, combined with preplant incorporated vernolate if nutgrass was serious. Napropamide and metobromuron merit further testing.     

Weed interference and control in white yam (Dioscorea rotundata Poir)

The climbing growth habit of white yam (Dioscorea rotundata) coupled with its inability to shade the ground completely at any stage of its growth and development makes it very susceptible to weed interference. The critical period of weed interference in white yam was between the 8th and 16th week after planting (w.a.p.) for yam planted at the onset of rains. Keeping yam plants weedy for 16 w.a.p. or beyond significantly reduced tuber yield; keeping them weed-free for the same period resulted in a significant increase in tuber yield. A minimum of three weedings within 16 w.a.p. are necessary to minimize yield reduction caused by weeds. The lowest weed weight and highest crop yield from herbicide treatments were obtained where a mixture of fluometuron and metolachlor each at 2.0 kg a.i./ha was used for pre-emergence weed control.     

Weed control in squash and tomato fields by soil solarization in the Jordan Valley

Black (BPE) and clear polyethylene mulches (CPE), 0.08 and 0.06 mm thick, respectively, were compared for their effectiveness for soil solarization over three seasons during 1986–1989 in weedy fields. Solarization for 6 weeks reduced weed growth and enhanced crop yields. However, further mulching with BPE after solarization with either BPE or CPE gave the best results. Not all weeds were sensitive to solarization. Some weed species were completely controlled; others were reduced to varying degrees; yet other weed species seemed to be enhanced by solarization. Solarization without further mulching was no better than farmer-practice in reducing weed growth or in increasing crop yield. Weeds required further removal after the middle of the growing season. Any soil disturbance after solarization reduced the weed control effect of solarization. Crops grew best in plots after solarization with BPE if they were planted in the same mulch after it was perforated.     

WEED COMPETITION IN TRANSPLANTED SPRING CABBAGE

Summary. The presence of weeds during autumn and winter had no effect on the growth of transplanted spring cabbage provided they were removed before rapid growth of crop and weeds began in early spring. Weeds left beyond this time competed with the crop, resulting in smaller marketable heads. Increasingly severe competition affected internal head quality, reduced the numbers of plants producing heads and resulted in the death of a proportion of the crop plants. The main weed species responsible for crop loss was Stellaria media , which survived winter frosts and grew rapidly in early spring to fill all available ground space, dominating the weed flora and shading the crop foliage.
Application of propachlor at planting time failed to give sufficient control of S. media to avoid crop loss, although the onset of competition was delayed. Trifluralin gave excellent control of S. media and resistant species were kept in check by the crop.
Comparison of cropped and uncropped plots showed that the crop exerted considerable competitive pressure on the growth and development of weeds particularly where this had been reduced or delayed by herbicide treatment. The presence or absence of the crop did not, however, affect the relative proportions of the weed species, except on plots treated with trifluralin, where the crop suppressed Capsella bursa-pastoris.
Compétition des mauvaises herbes dans les choux de printemps repiqués     

Influence de la densité et de la période de concurrence de Solanum nigrum L. sur la tomate de semis direct, en relation avec le désherbage

Effects of density and period of competition by Solanum nigrum L. on direct seeded tomatoes in relation to weed control The effects of density and period of competition from Solanum nigrum L. were measured in direct seeded tomatoes given weed control treatments currently used in south-east France. S. nigrum emerging after a diquat treatment at the 2–3 leaf stage of the crop and thinned to low densities (<12.8 plants ha^?1) at the 5–6 leaf stage of the crop caused significant yield loss if left to compete with the crop until harvest. Yield reduction was smaller if the same weed densities were present only until the onset of flowering. The regression curves of yield on weed density differed as annual climatic variations affected sowing date and plant growth; a comparison between years was made using the relation ‘crop yield × weed biomass/crop biomass’. Significant interactions between weed density and period of competition were found with yield of both green and red fruit. For late sown crops with low densities of S. nigrum two weed control treatments at the 5–6 leaf stage and at the onset of flowering were sufficient to prevent yield loss.     

Weed community characteristics and crop performance: a neighbourhood approach

The presence of weeds in crop fields often causes yield reductions. However, the effects of weed diversity have not been fully examined. This study tested the hypothesis that, holding density constant, increased weed species richness would decrease the effects of competition on spring wheat target plant performance. Measurements were taken from circular neighbourhoods (16.5 cm radius) with a single spring wheat plant surrounded by combinations of Setaria viridis , Avena fatua , Kochia scoparia , Thlaspi arvense and spring wheat plants, representing all combinations of neighbour species at four density levels. Using regression models, we found that species richness had no significant direct effect on spring wheat biomass, yield, or relative growth rate and that there were no significant neighbour species interaction terms. For weedy target plants, the presence of negative interaction terms suggested that increasing species richness had negative effects on growth of individual weed species. Additional analysis suggested that increased species richness may limit competitive ability of dominant weeds. Although we found no evidence of a direct effect of weed species richness on crop performance, increased weed species richness had no negative effect on spring wheat performance. Further, species richness of the weed community appears to influence weedy plant performance, which may offer a future opportunity to influence crop performance.     

Interference of redroot pigweed (Amaranthus retroflexus) in green bean (Phaseolus vulgaris)

Weeds that emerge along with or immediately after crop plants usually can reduce the yield of those crops. Two randomized complete block design experiments were conducted during 2006 and 2007 in Tabriz, Iran to determine the critical period of redroot pigweed control in the green bean hybrid “Cantander”. The treatments were weed‐infested and weed‐free plots at 2, 4, 6, 8, 10, and 14 weeks after bean emergence (WABE). The green bean biomass was affected by the early emergence of redroot pigweed, but it was not reduced when redroot pigweed emerged at 10 weeks after crop emergence, along with crop emergence, and grew with green bean until 4 WABE. The redroot pigweed biomass decreased by 2.7 g m^?2 per day when weed emergence was delayed. Each 100 g m^?2 of weed biomass that was produced resulted in a 1.4 kg ha^?1 loss in the green bean yield. When redroot pigweed interference lasted for ≥4 weeks after green bean emergence, the green bean yield was reduced significantly. Weeds, which emerged 2 weeks after green bean and thereafter were controlled, did not decrease crop productivity significantly. The highest crop yield was obtained when the weed emerged at 14 WABE. The critical period of redroot pigweed control, considering a 10% yield loss, was between 19 and 55 days after green bean emergence. Thus, weed control practises should be begun no later than 3 WABE and should continue until at least 8 WABE in order to obtain the maximum green bean yield.