Potential of nine multipurpose tree species to reduce saline groundwater tables in the lower Amu Darya River region of Uzbekistan

This paper evaluates the potential of nine multipurpose tree species for afforestation of degraded land in the Khorezm region, Central Asia, particularly their suitability for biodrainage i.e., lowering the elevated groundwater table through the transpirative capacity of plantations. For this purpose water use (WU), water use efficiency (WUE) and tree physiological factors influencing transpiration were assessed during two consecutive years. Mean daily leaf transpiration differed significantly among the species and ranged during the seasons from 4.5–5.2 mmol m⁻² s⁻¹ for Prunus armeniaca L. to 4.5–10 mmol ⁻² s⁻¹ for Elaeagnus angustifolia L. WU differences were triggered by species physiological features such as capability of water uptake by roots. Transpiration rates and the length of fine roots correlated highly (r = 0.7). Correlations of leaf transpiration rates with leaf area were weaker (r = 0.6). No correlations were found between salt content in plants and water uptake under conditions of slight-to-moderate rootzone soil salinity. Values of WUE per root and shoot DM were similar averaging, respectively, 0.2 and 0.3 g DM g⁻¹ water for two-year-old trees, and decreased with age. In addition to WU characteristics, also salinity tolerance, growth rate and the ability to produce fodder and fuelwood must be considered during species selection. Regarding these features, the N-fixing E. angustifolia ranked the highest, combining high WU, fast growth and production of nutritious feed. Examined Populus spp. and Ulmus pumila L. ranked lower but still represented potential candidates for biodrainage purposes. Typical fruit species in the region such as P. armeniaca and Morus alba, showed low biodrainage potential.     

Performance of Zebu Steers Grazing on Western Tanzania Native Forages Supplemented with <Emphasis Type="Italic">Leucaena Leucocephala</Emphasis> Leaf Meal

Information is lacking on the potential of leguminous fodder trees such as Leucaena leucocephala as a feed resource to supplement the native forages in traditional grazing management systems in the tropics. Two studies were conducted (1) to assess traditional fodder banks’ forage nutritive potential on animal production, and (2) to investigate the effect of Leucaena leucocephala leaf meal (LLM) supplementation on growth of steers grazing standing hay basal forages in the dry season. The traditional forages had low nutritive values indicated by low mean crude protein (CP) of 23 g kg⁻¹ dry matter (DM), and high fibre contents of 717, 546 and 153 g kg⁻¹ DM for neutral detergent fibre (NDF), acid detergent fibre (ADF) and acid detergent lignin (ADL), respectively. The forages were poorly degraded in sacco and had low metabolisable energy (ME) (4.2–4.6 MJ kg⁻¹ DM). Mean washing losses A, slowly degradable DM fraction, B, potential degradability, (A+B) and mean 48 h DM degradability (DMD) of grazing land forages were 70, 471, 541 and 326 g kg⁻¹ DM, respectively. In the supplementation study, 16 growing steers (160.8±0.24 kg) were randomly allocated into four groups, in a completely randomized design. Four LLM treatment diets (T1, T2, T3 and T4), with four levels: 0, 0.4, 0.6 and 0.8 kg DM for control, low, medium and high LLM levels, respectively, were randomly allocated to the animals in the four groups for 70 days (d). LLM supplementation (p<0.05) improved mean weight gain from −0.30 to 0.26 kg steer⁻¹ d⁻¹, for T1 and T4, respectively. Steers on T4 gained (p<0.05) more weight compared to the animals in T1 and T2, though there was no (p>0.05) difference in weight gains between animals on T3 and T4 (0.14 vs. 0.26 kg steer⁻¹ d⁻¹, respectively). Standing hay basal forages alone could not sustain animal productivity during dry seasons unless corrected for protein. Higher levels of LLM supplementation prevented weight losses and improved the performance of grazing steers, a management practice thought appropriate to low income pastoralists in semiarid western Tanzania.     

Shrub yield and fodder quality variations in a non-tropical dryland environment in West Asia

Integration of shrubs into the smallholder crop-rangeland-livestock farming systems in non-tropical dry areas could reduce feed gaps, rangeland degradation and desertification, but data on yield and fodder quality of most native and exotic shrubs are scanty. The study aimed at identifying shrubs for dryland agroforestry based on agronomic and fodder quality attributes. Fodder and wood yield, percent fodder (FBR) and fodder quality of 26 non-legume shrubs were determined from 6 to 8 months regrowth in north-west Syria. Seed yield was estimated from plants that were not cut. Fodder (31–87 Mg DM ha⁻¹), wood (16-2064 Mg DM ha⁻¹) and seed (0-132 Mg DM ha⁻¹) yield, FBR (24–87%), fodder concentrations of crude protein (69–195 g kg⁻¹.), acid detergent lignin (ADL) (24–109 g kg⁻¹), acid detergent fiber (ADF) (102–267 g kg⁻¹), neutral detergent fiber (NDF) (214–526 g kg⁻¹), in vitro organic matter digestibility (IVOMD) (391–526 g kg⁻¹), and in vitro gas production after 24 h of incubation (25–39 ml 200 mg⁻¹ DM) varied (P < 0.05) among the shrubs. Atriplex halimus-halimus and A. herba-alba from Spain, A. canescens, A. ploycarpa and A. lentiformis from USA, A. halimus and A. herba-alba from Syria and A. nummularia from Australia and South Africa had greater potential for development of dryland agroforestry technologies. The promising shrubs could be integrated into the rangeland-crop-livestock systems in non-tropical dryland environs to provide fodder, fuel-wood, shade, medicine, stabilize sand-dunes, and sequester carbon; thereby contributing to mitigation of rangeland degradation and global warming; if major constraints to adoption of fodder trees such as agronomic problems, low multipurpose value, land shortage and quality seed supply could be overcome.     

Evaluation of 15 indigenous and introduced species for reforestation and agroforestry in northeastern Mexico

The coast of the Gulf of Mexico is characterized by dry regions with high variation in climatic conditions. This area is rich in drought-tolerant or subhumid species. The species that are potentially useful for reforestation, regreening, agroforestry activities and the production of timber, fodder, fuelwood and human food have been overexploited, resulting in the gradual decrease and degradation of their populations. This study was undertaken in order to suggest ways of improving the regeneration of these species. Fifteen native and exotic multipurpose tree species of low dry shrubland planted in monoculture in four randomized blocks. Measurements of various growth parameters, volume of trees, fodder potential and agroforestry uses over 15 years were evaluated. Eucalyptus camaldulensis Dehnh., E. microtheca F. Muell., Leucaena leucocephala (Lam.) de Wit. (exotic species), Acacia farnesiana (L.) Wild and Parkinsonia aculeata L. (native species) tend to have better characteristics in terms of growing annual rate, economic value and management schemes, while Prosopis glandulosa Torr. and Helietta parvifolia (Gray) Benth. (native species) did not establish well due to biotics problems which arose under plantation conditions. Acacia rigidula Benth., A. wrightii Benth. and two Pithecellobium spp. (native species) had intermediate yields of great interest since their multipurpose potential is the best of all 15 species. The forage potential of the exotic species (483–1684 kg DM//ha/year) were notably superior to native species (76–721 kg DM/ha/year). The firewood production volume varied between averages of 0.3–1.2 (native species) and 0.4–2.5 m³/ha/year (exotic species). This revised version was published online in June 2006 with corrections to the Cover Date.     

Growth and ecological impacts of traditional agroforestry tree species in Central Himalaya, India

A number of multipurpose tree species are conserved as scattered trees in settled farms on terraced slopes by the traditional farmers in Central Himalaya, India. Knowledge on growth rates and ecological impacts of these tree species is limited. Ten locally valued multipurpose tree species, viz., Albizzia lebbek, Alnus nepalensis, Boehmeria rugulosa, Celtis australis, Dalbergia sissoo, Ficus glomerata, Grewia optiva, Prunus cerasoides, Pyrus pashia and Sapium sebiferum, were established as mixed plantations at a degraded community forest land site and an abandoned agricultural land site in a village at 1200 m altitude in District Chamoli, India. At the abandoned agricultural land site, annual food crops were grown, along with planted trees, providing supplemental irrigation and organic manure following traditional farming practices. Survival, height, stem circumference, crown depth and width, number of branches, above-ground biomass and soil physico-chemical characteristics were monitored up to five years of plantation growth. Above-ground tree biomass accumulation at the abandoned agricultural land site was 3.9 t ha⁻¹ yr⁻¹ compared with 1.1 t ha⁻¹ yr⁻¹at the degraded forest land site. B. rugulosa, C. australis, F. glomerata, G. optiva, P. cerasoides and S. sebiferum showed more prominent differences in growth at the two sites compared with A. lebbek, A. nepalensis, D. sissoo and P. pashia. A. nepalensis and D. sissoo showed best growth performance at both the sites. A significant improvement in soil physico-chemical characteristics was observed after five years at both of the sites. Carbon sequestration in soil was higher than that in bole biomass. This revised version was published online in June 2006 with corrections to the Cover Date.     

Important characteristics of some browse species in an agrosilvopastoral system in West Africa

Three browse species, Afzelia africana Sm., Khaya senegalensis (Desv.) A. Juss., and Pterocarpus erinaceus Poir. were investigated as agroforestry system components in a subhumid zone of West Africa. The foliation, flowering and fruiting of ten trees per species were recorded every 15 days for 2 years. The total foliage biomass at maximum availability was determined by complete pruning of 75 trees. The chemical composition of the foliage and the proportion of trees pruned on the pasture were determined. The phenological phases of the species began in the dry season and ended at the end of the rainy season. Afzelia africana and Pterocarpus erinaceus were totally defoliated during 2–6 weeks while K. senegalensis replaced the foliage progressively and earlier. The crude protein content was significantly different (123 g, 102 g and 92 g kg⁻¹ dry matter (DM) for Afzelia africana, Pterocarpus erinaceus and K. senegalensis, respectively). The foliage biomass per tree of K. senegalensis, Pterocarpus erinaceus and Afzelia africana differed significantly (41 kg, 30 kg and 21 kg DM ha⁻¹, respectively) while Pterocarpus erinaceus had the highest available foliage biomass per ha. The trees of Afzelia africana were intensively pruned. There was a significant relationship found between foliage biomass and circumference of the crown for Afzelia africana (R ² = 82%) and Pterocarpus erinaceus (R ² = 81%). Relationships were also found between circumference of the branches and foliage biomass. In conclusion, the trees are important potential fodder and nitrogen sources for animals in the agrosilvopastoral system and the phenological differences make the fodder available during a long period of time.     

Seasonal dynamics of mineral N pools and N-mineralization in soils under homegarden trees in South Andaman,India

Agroforestry trees are now well known to play a central role in the build up of nutrients pools and their transformations similar to that of forest ecosystem, however, information on the potential of homegarden trees accumulating and releasing nitrogen (mineralization) is lacking. The present study reports seasonal variations in pool sizes of mineral N (NH₄⁺-N and NO₃⁻-N), and net N-mineralization rate in relation to rainfall and temperature under coconut (Cocos nucifera L.), clove (Eugenia caryophyllata Thunb) and nutmeg (Myristica fragrans Houtt. Nees) trees in a coconut-spice trees plantation for two annual cycles in the equatorial humid climate of South Andaman Island of India. Concentration of NH₄⁺-N was the highest during wet season (May–October) and the lowest during post-wet season (November–January) under all the tree species. On the contrary, concentration of NO₃⁻-N was the lowest in the wet season and the highest during the post-wet season. However, concentrations of the mineral N were the highest under the nutmeg and the lowest under the coconut trees. Like the pool sizes, mean annual mineralization was the highest under the nutmeg (561 mg kg⁻¹ yr⁻¹) and the lowest under the coconut trees (393 mg kg⁻¹ yr⁻¹). Rate of mineralization was the highest during the post-wet season and the lowest during the dry season (February–April) under all the tree species. High rainfall during the wet season, however, reduced the rate of nitrification under all the tree species. The mean annual mineralization was logarithmically related with rainfall amount and mean monthly temperature.     

Litter production and decomposition in the forested areas of traditional homegardens: a case study from Barak Valley,Assam, northeast India

Homegardens are one of the oldest forms of managed land use systems characterised by high diversity and complexity of their species structure which in turn contribute to efficient nutrient cycling. Litterfall and decomposition are the two major processes that replenish the soil nutrient pools and endow sustainability to these agroforests. A study was carried out in the village Dargakona, Barak Valley, northeast India to understand the pattern of litter production and litter decomposition in the traditional homegardens. Annual litter production was 6.27 Mg ha⁻¹ with a bimodal distribution pattern and the nitrogen input through litterfall accounted for 48.17 kg ha⁻¹ year⁻¹. Litter decomposition studies for ten multipurpose trees revealed Sapium baccatum and Toona ciliata to be the most labile litter species and the decay rate coefficients varied among the species with differing rates of nutrient release pattern. Such studies can provide information regarding the litter quality of indigenous tree species and help validate farmers planting and management of multiple species which allows for efficient nutrient cycling of the system.     

Fodder production responses to pruning height and fodder quality of some trees and shrubs in a forest-savanna transition zone in southwestern Nigeria

Seasonal fodder production responses of five shrubs (Centrosema arenarium, Desmodium strigillosum, Desmodium velutinum, Phyllodium pulchellum, and Tadehagi triquetrum)and five trees (Albizia gummifera, Berlinia grandiflora, Albizia niopoides, Bauhinia monandra, and Inga edulis) to pruning heights ranging from 15 to 75 cm were evaluated during the main-wet, minor-wet, and dry seasons of 1993 and 1994 in the forest-savanna transition zone of West Africa. Fodder from the main-wet season was analysed for nitrogen (N) and phosphorus (P), and dry matter degradation characteristics after 6, 12, 24, 48, 72 and 96 h of incubation in rumen-fistulated N'Dama steers. In the minor-wet season, fodder production of all species increased in response to increasing pruning height with the exception of P. pulchellum and A. gummifera. Highest fodder production was attained at a pruning height of 45 cm for the Desmodium species, 50 cm for P. pulchellum, and 75 cm for the rest of the species. Concentrations of N and P varied significantly among the species; for N the ranges were 28.7–38.8 g kg^–1 (shrubs) and 25.3–44.5 g kg^–1 (trees), while for P the ranges were 3.26–7.04 g mg^–1 (shrubs) and 3.58–6.76 g mg^–1 (trees). Dry matter degradation characteristics differed significantly among shrubs and trees; ranges for shrubs were: soluble fraction (a), 128–185; degradable fraction (b), 664–703; potential degradability (PD), 793–857, as g kg^–1; rate of degradation (c), 0.0241–0.0308 as % h^–1 while for trees ranges were: (a), 139–160; (b), 651–826; (PD), 824–970, as g kg^–1; (c), 0.0143–0.0227 as % h^–1. Based on fodder production and quality, C. arenarium, D. strigillosum, D. velutinum, B. monandra, I. edulis, and A. niopoides were the most promising species for the development of animal agroforestry technologies in the west African forest-savanna transition zone and similar environments in the tropics.This revised version was published online in November 2005 with corrections to the Cover Date.     

Structure and function of traditional agroforestry systems in the western Himalaya. I. Biomass and productivity

Species composition, biomass, and productivity patterns of three types of traditional agroforestry systems, namely, agrisilvicultural, agrihorticultural, and agrihortisilvicultural, commonly practiced in the western Himalaya, were studied. Species composition in the systems varied depending upon the size of the land holdings, and the basic requirements of the farmer. Among three systems agrihortisilvicultural was highly diverse in vegetation, with as many as 13 tree and 5 agricultural crops mixed together. This system showed the highest productivity up to 25.8 t ha⁻¹ yr⁻¹, out of which 68 percent was contributed by the trees and the remainder by the annuals. Agrisilvicultural system having predominantly annuals had the lowest productivity of 20.4 t ha⁻¹ yr⁻¹ with only 27 percent contribution by the trees. Total aboveground biomass in agrihortisilvicultural or agrihorticultural system was around 48 t ha⁻¹ and it was about 2-fold higher than agrisilvicultural system. In fodder trees, a significant percentage of annual production, up to 48 percent, was allocated in current twigs, while in horticultural trees a major portion, up to 63 percent was partioned towards fruits. Implications of the results have been discussed in context with the growing problems of fuel, fodder, and packaging material in the western Himalaya.     

Interactions between Widely Spaced Young Poplars (Populus spp.) and Introduced Pasture Mixtures

Silvopastoral systems involving poplars are widespread in rural landscapes in a number of regions of New Zealand. The effect of widely spaced trees of Populus nigra × P. maximowiczii, aged 8–11 years, on the growth and botanical composition of understorey pasture mixes was determined over 3 years at a southern North Island hill country site. Pasture mixes comprised existing pasture, and two introduced test swards comprising new grass (Agrostis capillaris, Dactylis glomerata, Lolium perenne) and legume (Lotus uliginosus, Trifolium repens) cultivars. Pasture accumulation beneath trees (6.6 t DM ha⁻¹ year⁻¹) was 23% less than open (unshaded, no trees) pasture (8.6 t DM ha⁻¹ year⁻¹) and differences in accumulation occurred between tree aspects (North and South sides of trees) when trees were foliated. In early spring, North plots produced 11–14% more herbage than South plots whereas in late summer, South plots produced up to 44% more herbage than North plots. Yield of both test swards (6.4 and 8.0 t DM ha⁻¹ year⁻¹) was not significantly different from that of the existing sward (7.4 t DM ha⁻¹ year⁻¹) but productivity varied between swards in spring and summer. One test sward included Lotus uliginosus cv. Grasslands Maku, and the sward produced 30% more DM than the other swards in mid-summer. Dactylis glomerata cv. Grasslands Wana comprised 37% of the DM of swards in which it was sown and was the most successful cultivar, followed by Grasslands Maku. Both cultivars are recommended for silvopastoral systems where pasture is lightly grazed.     

Litter dynamics of six multipurpose trees in a homegarden in Southern Kerala, India

Multipurpose trees, the integral components of homegardens, contribute significantly to the closed nutrient cycling processes and sustainability of the ecosystem. Although, the litter production and probable nutrient returns via litter in homegardens have been documented, quantification and characterization of the decomposition and bioelement release from the litter have received relatively little scientific attention. The objective of the present study is to explore the litter dynamics of six locally important multipurpose trees (Mangifera indica L., Artocarpus heterophyllus Lamk., Anacardium occidentale L., Ailanthus triphysa Dennst., Artocarpus hirsutus L. and Swietenia macrophylla L.), in an agroforestry homegarden in Southern Kerala, India. Litterfall and nutrient additions in the six species ranged from 383 to 868 g m⁻² yr⁻¹, nitrogen, 6.4 to 8.8, phosphorus, 0.17 to 0.42 and potassium, 1.1 to 2.8 g m⁻² yr⁻¹. The annual litter output in the homegarden was 425 kg with A. hirsutus, M. indica, A. heterophyllus and A. occidentale recording significantly higher litter and nutrient additions. Leaf litter decay studies revealed A. heterophyllus and A. occidentale to be the most labile litter species and S. macrophylla the most recalcitrant. The decay rate coefficients varied significantly among the species. Foliage decomposition rates related to the initial chemical composition of the litter revealed best correlation with lignin. NPK release was almost complete by the end of decay in all species inspite of the initial phases of accumulation observed for nitrogen and phosphorus. Two-way analysis of variance test revealed significant differences in the contents of the three elements as a function of species and time elapsed. Macronutrients were released in the order K>N/P. The higher rates of decay and nutrient turnover in M. indica, A. heterophyllus and A. occidentale foliage indicated the potential of using these species’ litter as nutrient inputs in agriculture while A. triphysa, A. hirsutus and S. macrophylla perform better as organic mulches taking a longer time for decay and hence nutrient release.     

Pasture production under different tree species and densities in an Atlantic silvopastoral system

We studied the effect of six tree species planted at six different densities on pasture production seven years after establishment. Annual and seasonal pasture production was studied every six months, over three years. Pasture production was lower under conifer trees (Pseudotsuga menziesii (Mirb.) Franco, Pinus pinaster Aiton, Pinus radiata D. Don) than under broadleaved trees (Betula alba L., Quercus rubra L. and Castanea sativa Mill.). Annual pasture production under Pseudotsuga menziesii and Pinus pinaster decreased progressively starting from 952 trees ha⁻¹, while decline in herbage production under Pinus radiata began to occur at 427 trees ha⁻¹. Tree density effect on pasture production was detected at 2,000 trees ha⁻¹ for all of the deciduous species studied. This effect on pasture production was more important in the first six months of the year (June sampling), while from June to December herbage production was less affected by tree density. The tree effect became more noticeable over time, with the last sampling showing the inverse relationship between tree density and herbage production most clearly. Seven years after tree establishment, pasture production was quite consistent under tree densities between 190 trees ha⁻¹ and 556 trees ha⁻¹ and declined remarkably from 556 trees ha⁻¹ to 2,500 trees ha⁻¹. The study also indicated that by the sixth growing season, annual pasture production under different tree species is inversely correlated with tree leaf area index.     

Do multipurpose companion trees affect high value timber trees in a silvopastoral plantation system?

Establishment of native timber trees on deforested land may contribute to the livelihood of farmers, to improved ecosystem services and to increased greenhouse gas uptake. Here, we present a new silvopastoral planting design to assess species performance and interspecific competition or facilitation effects among native timber and multipurpose trees in Central America. Two timber species, Tabebuia rosea and Cedrela odorata, were established in three low-density planting regimes allowing combined tree and future livestock production: (1) solitary planting, (2) companion planting with Guazuma ulmifolia, and (3) companion planting with the nitrogen-fixing Gliricidia sepium. We quantified survival, growth and reforestation potential of the two timber species subjected to the different planting regimes for the first 2 years after establishment. Nitrogen concentration as well as stable nitrogen and carbon isotope composition (δ¹⁵N, δ¹³C) of leaves of the timber saplings were determined. T. rosea showed higher survival and better growth than C. odorata under varying environmental conditions (soil, concomitant vegetation). Performance of the timber saplings was unaffected by either companion species. Planting regimes had no effect on foliar nitrogen concentration and δ¹⁵N of the two timber species, although δ¹⁵N values indicated nitrogen fixation activity in G. sepium trees. Planting regimes affected foliar δ¹³C values in T. rosea. δ¹³C values were significantly higher in solitarily growing individuals, suggesting lower exposition to water stress conditions in saplings surrounded by companion species. As we found positively correlated growth traits among timber and multipurpose trees, a combined planting may benefit farmers by providing additional goods and services.     

Nitrogen and fine root length dynamics in a tropical agroforestry system with periodically pruned Erythrina poeppigiana

The effect of pruning all branches (complete pruning) or retaining one branch (partial pruning) on the dynamics of nitrogen cycling in aboveground biomass, nitrogen supplying power of an amended Eutric Cambisol, and fine root length, was studied in an Erythrina poeppigiana (Walp.) O.F. Cook—tomato (Lycopersicon esculentum Mill.) alley cropping practice in Turrialba, Costa Rica during 1999–2000. Over the 1 year pruning cycle, in which trees were completely or partially pruned four times, respective aboveground biomass production was 4.4 Mg or 7 Mg ha⁻¹ (2-year-old trees) and 5.5 Mg or 9 Mg ha⁻¹ (8-year-old trees); N cycled in aboveground biomass was 123 kg or 187 kg ha⁻¹ (2-year-old trees) and 160 kg or 256 kg N ha⁻¹ (8-year-old trees); mean fine root length was 489 or 821 m (2-year-old-trees), 184 or 364 m per tree (8-year-old-trees). Pruning intensity did not significantly affect net N mineralisation and net nitrification rates during the tomato-cropping season. For the tomato crop, pre-plant mean net N mineralisation rate of 2.5 mg N kg⁻¹ soil day⁻¹ was significantly lower than 16.7 or 11.6 mg N kg⁻¹ soil day⁻¹ at the end of vegetative development and flowering, respectively. Mean net nitrification rates of 3.5, and 4.3 mg N kg⁻¹ soil day⁻¹, at pre-plant and end of vegetative development, respectively, were significantly higher than 0.3 mg N kg⁻¹ soil day⁻¹ at end of flowering. In humid tropical low-input agroforestry practices that depend on organic inputs from trees for crop nutrition, retention of a branch on the pruned tree stump appears to be a good alternative to removal of all branches for reducing N losses through higher N cycling in aboveground biomass, and for conserving fine root length for higher N uptake, although it might enhance competition for associated crops.     

Effect of dried Gliricidia sepium leaf supplement on feed intake, digestibility and nitrogen retention in sheep fed dried KW4 elephant grass (Pennisetum purpureum) ad libitum

A feeding experiment was conducted to study the effects of supplementation with dried Gliricidia sepium leaves on dry matter (DM) intake, digestibility and nitrogen retention in sheep fed KW4 elephant grass. Four mature rams were fed elephant grass ad libitum supplemented with four levels of gliricidia leaves (0, 4, 8 and 12 g DM kg⁻¹ lwt day⁻¹) in a 4 × 4 Latin square design. Supplementation with gliricidia leaves decreased elephant grass DM intake (g DM day⁻¹ or g DM kg⁻¹ BW^0.75 day⁻¹). However, the effect of supplementation on total DM intake of the rations was not significant (P >; 0.05). Total crude protein intake significantly (P < 0.001) increased with increasing levels of gliricidia supplementation. Total DM digestibility and body weight changes were significantly (P < 0.05) improved by gliricidia supplementation; with the highest digestibility coefficient (60.5%) and body weight gain (89.3 g/day) obtained at 8 g DM kg⁻¹ lwt day⁻¹. Gliricidia supplementation significantly (P < 0.001) improved nitrogen intake, absorbed nitrogen and retained nitrogen but with no significant difference at 8 and 12 g DM kg⁻¹ lwt day⁻¹ level of supplementation. The highest efficiency of N retention by sheep (44.9%) was obtained at 8 g DM kg⁻¹ lwt day⁻¹ level of gliricidia supplementation. The results indicated that supplemention of KW4 elephant grass with small quantities of gliricidia leaves up to 8 g DM kg⁻¹ lwt day⁻¹ enhance utilisation efficiency of the total ration. Further increases in the level of gliricidia supplementation, under the conditions of this experiment, reduced the intake of elephant grass leading to substitution effects of the basal. This revised version was published online in August 2006 with corrections to the Cover Date.     

Dry matter production,morphology and nutritive value of <Emphasis Type="Italic">Dactylis glomerata</Emphasis> growing under different light regimes

Plant growth, morphology and nutritive value under shade can differ between temperate grasses. Therefore, the aim of this study was to quantify the dry matter (DM) production, sward morphology, crude protein (CP%), organic matter digestibility (OMD) and macro-nutrient concentrations (P, K, Mg, Ca and S) in a grazed cocksfoot (Dactylis glomerata L.) pasture under 10-year-old Pinus radiata D. Don forest. Four levels of light intensity were compared: full sunlight (100% photosynthetic photon flux density-PPFD), open + wooden slats (∼43% PPFD), trees (∼58% PPFD) and tree + slats (∼24% PPFD). The mean total DM production was 8.2 t DM ha⁻¹ yr⁻¹ in the open and 3.8 t DM ha⁻¹ yr⁻¹ in the trees + slats treatment. The changes in cocksfoot leaf area index (LAI) were related to variations in morphological aspects of the sward such as canopy height and tiller population. CP% increased as PPFD declined with mean values of 18.6% in open and 22.5% in the trees + slats treatment. In contrast, the intensity of fluctuating shade had little effect on OMD with a mean value of 79 ± 3.2%. The mean annual macro-nutrient concentrations in leaves increased as the PPFD level declined mainly between the open and the trees + slats treatments. It therefore appears that heavily shaded dominant temperate pastures in silvopastoral systems limit animal production per hectare through lower DM production rates and per animal through reduced pre-grazing pasture mass of lower bulk density from the etiolated pasture.     

Effects of mother tree ages,different rooting mediums,light conditions and auxin treatments on rooting behaviour of <Emphasis Type="Italic">Dalbergia sissoo</Emphasis> branch cuttings

Dalbergia sissoo Roxb. is one of the promising multipurpose tree species of South Asia. Most of the plantations of D. sissoo from seeds are facing severe threats due to the die-back disease, which ultimately causes death of this potential tree-species within a few months. Vegetative propagation could avoid the die-back disease. Thirty mother trees of different age-groups of D. sissoo were selected for evaluating the rooting behaviour of branch cuttings from D. sissoo as influenced by auxins (IAA or IBA at 100, 200, 500 mg·L⁻¹), ages of mother trees (10, 4 and 2 years old) and different environment conditions, i.e., different mediums (soil and sand) or light conditions (in shade and open condition). The results show that application of IAA and IBA induced more numbers of cuttings (collected from 10-year-old mother trees) to root compared to control. Branch cuttings of D. sissoo collected from 10-year-old mother trees and planted in soil bed in open conditions had 100.0% of cuttings to root in IAA (100 mg·L⁻¹) and IBA (200 mg·L⁻¹) treatments. Both rooting medium (Soil and sand) influenced significantly (p<0.05) on rooting response of branch cuttings. Soil medium was found to achieve maximum no. of branch cuttings to root, compared to sand medium.     

Management of pigeon pea in short fallows for crop-livestock production systems in the Guinea savanna zone of northern Ghana

Crop and livestock production in the Guinea savanna zone of northern Ghana has been declining over the past years as a result of increasing pressure on land. To sustain soil productivity, pigeon pea(Cajanus cajan), a leguminous perennial crop was evaluated for its potential as a short duration fallow crop for fodder and grain, and maize (Zea mays)production. It involved comparing a natural fallow (i.e., control) and four improved fallows of pigeon pea pruned annually at 30 cm, 60 cm and 90 cm from the ground, and unpruned pigeon pea over a two-year period. After this time, the land was cleared manually and planted to maize. The highest mean annual biomass of pigeon pea over the two-year period of 6.1 t ha⁻¹ dry matter (DM) was obtained by pruning at 60 cm. The highest leaf litter production and pigeon pea seed yield was obtained from the no pruning treatment. The mean maize grain yield from the improved fallow (3.02 t ha⁻¹) in the first year after clearing was significantly (P < 0.05) greater than that of the natural fallow (1.54 t ha⁻¹). Considering the biomass of pigeon pea from pruning, pigeon pea seed yield and maize grain yield after the pigeon pea, pruning pigeon pea at 60 cm is the most promising regime for crop-livestock production systems. This revised version was published online in June 2006 with corrections to the Cover Date.     

Biomass production, foliar and root characteristics and nutrient accumulation in young silver birch (Betula pendula Roth.) stand growing on abandoned agricultural land

The above-ground biomass and production, below-ground biomass, nutrient (NPK) accumulation, fine roots and foliar characteristics of a 8-year-old silver birch (Betula pendula) natural stand, growing on abandoned agricultural land in Estonia, were investigated. Total above-ground biomass and current annual production after eight growing seasons was 31.2 and 11.9 t DM ha⁻¹, respectively. The production of stems accounted for 62.4% and below-ground biomass accounted for 19.2% of the total biomass of the stand. Carbon sequestration in tree biomass reaches roughly 17.5 t C ha⁻¹ during the first 8 years. The biomass of the fine roots (d < 2 mm) was 1.7 ± 0.2 t DM ha⁻¹ and 76.2% of it was located in the 20 cm topsoil layer. The leaf area index (LAI) of the birch stand was estimated as 3.7 m² m⁻² and specific leaf area (SLA) 15.0 ± 0.1 m² kg⁻¹. The impact of the crown layer on SLA was significant as the leaves are markedly thicker in the upper part of the crown compared with the lower part. The short-root specific area (SRA) in the 30 cm topsoil was 182.9 ± 9.5 m² kg⁻¹, specific root length (SRL), root tissue density (RTD) and the number of short-root tips (>95% ectomycorrhizal) per dry mass unit of short roots were 145.3 ± 8.6 m g⁻¹, 58.6 ± 3.0 kg m⁻³ and 103.7 ± 5.5 tips mg⁻¹, respectively. In August the amount of nitrogen, phosphorus and potassium, accumulated in above ground biomass, was 192.6, 25.0 and 56.6 kg ha⁻¹, respectively. The annual flux of N and P retranslocation from the leaves to the other tree parts was 57.2 and 3.7 kg ha⁻¹ yr⁻¹ (55 and 27%), respectively, of which 29.1 kg ha⁻¹ N and 2.8 kg ha⁻¹ P were accumulated in the above-ground part of the stand.