Effect of spatial heterogeneity on natural regeneration of Manchurian ash

lntr0ducti0nSpatialheterogeneityisanimportantcharacteristicofecoIogicatsystem,itisreceivingm0reandmoreattentionbecauseofitsdeterminingroIeinecoIogicaIpatternandprocess(Robertsonetal1993,CadweIletal.1994,W8ngetal1997,1998,Lieta/1998,W8ng1999,Davieetal.1998).Spatialstructurecre-atedbymanyfactorsprofOundIyinfIuencesthedy-namics,composition,andbiodiversity0fcommunity(Tilman1994).DimensionanaIysisinspaceisindis-pensabletoexplaintheage-oIdquestionsaboutthemechanismsofplantcoexistenceinseeming…     

STUDY ON THE NATURAL REGENERATION EFFECT OF MANCHURIAN ASH BY OPENING UP EFFECT BELT IN DAHURIAN LARCH PLANTATION

NaturalregenerationofmanchurianashusuallytakesplacewheretherearesuitablesoilmoistureandseedbankofsecondarymanchurianashforestintheEasternMountainousAreaofNortheastChina.Sometimes,naturalregenerationofmanchurianashcanbefoundinpureconiferousplantation.Inordertofindoutthepossibilityofintroducingnaturalregenerationofmanchurianashinpureconiferousplantation,trialsandresear-cheshavebeencarriedoutindahurianlarchp1antationbyopeningupeffectbeltinthestands.lntheend,charactersofcommunitystructureandmicr…     

水曲柳单木生长模型的研究

根据黑龙江省、市、县576块水曲柳固定样地的解析木数据,在水曲柳单木直径生长模型中引入林木大小、竞争和立地条件因素,采用回归的方法建立水曲柳单木直径生长模型。研究表明：水曲柳单木生长模型与该林木自身的大小（lnD）、林分平均胸径（Dg）、对象木直径与林分中最大林木直径之比（DDM）以及土壤有较大的关系。林木的直径越大其生长量越大;竞争因子DDM说明水曲柳单木直径越大;其定期生长量越小;立地因子土壤表明,土层越浅其生长的越快。模型的检验精度为94%,说明模型具有良好的拟合效果,可用于实际的生产中。     

水曲柳地板毛坯的快速干燥

本文用连续升温基准在火管式加热木材干燥窑中对水曲柳地板坯进行了干燥试验。试验结果表明：用火管式加热木材干燥几连续升温基准干桫曲柳地板毛坯时，干燥速度快，干燥质量了，干燥基准易操作。     

Soil seed banks in four 22-year-old plantations in South China: Implications for restoration

To better understand the potentials of the soil seed banks in facilitating succession towards a more natural forest of native tree species, we quantified the size and composition of the soil seed banks in established plantations in South China. The seed banks were from four typical 22-year-old plantations, i.e., legume, mixed-conifer, mixed-native, and Eucalyptus overstory species. Species diversity in the seed banks was low, and the vegetation species differed from those found in the seed bank in each plantation. A total of 1211 seedlings belonging to eight species emerged in a seedling germination assay, among which Cyrtococcum patens was most abundant. All species detected were shrubs and herbs, and no viable indigenous tree seeds were found in soil samples. Size and species composition of the seed banks might be related to the overstory species compositions of the established plantations. The seed bank density in soils was highest in the mixed-conifer plantation followed by Eucalyptus, mixed-native, and legume plantations. Species richness among the seed banks of plantations was ranked as follows: Eucalyptus > mixed-conifer > mixed-native = legume. The results indicated that the soil seed banks of the current plantations are ineffective in regenerating the former communities after human disturbances. Particularly, the absence of indigenous tree species seeds in the seed banks would limit regeneration and probably contribute to arrested succession at the pioneer community stage. It would appear from these data that the soil seed banks under the current plantations should not be considered as a useful tool leading the succession to more natural stages. Introduction of target indigenous species by artificial seeding or seedling planting should be considered to accelerate forest regeneration.     

GROWTH EFFECT OF DAHURIAN LARCH-MANCHURIAN ASH MIXED FORESTS

STUDYAREAANDareTH0DThestUdyareawasatthecompatheentllofXinkenIllWorkingUnitinchoershanExPenmentalForestFann,NortheastF0restryUniversity.Dunngthespnngofl967,Dahu-nanlarchwasplantedintheleftlandofsecon-daryf0rest.Beforeorafterplanhng,mal1yMancl1tlrial1ash,e1l11withwhitebarkandsolneotherbroad-leavedtLreesinvadedandgrewwell.NowManchunanashandDahu-nanlarchn1ixedforestshavebeenf0n11ed,inwhichManchurianashisdolninant.lnl984,2Pen1lanentplotsweresetupwiththeslzeof0.O6hnl',facil1gsoutheast,…     

Observed dynamics of ponderosa pine (<Emphasis Type="Italic">Pinus ponderosa</Emphasis> var. <Emphasis Type="Italic">ponderosa</Emphasis> Dougl. ex Laws.) seedling recruitment in the Cascade Range,USA

An observational study of early seedling establishment (first 1–2 summers after emergence) was conducted in four ponderosa pine (Pinus ponderosa var. ponderosa Dougl. ex Laws.) stands east of the Cascade Range crest in central Oregon, USA. Newly emerged ponderosa pine seedlings were identified at the start of summer and were monitored through their second summer; a subsequent cohort of seedlings was identified and monitored for one summer. About 3% of the viable seed that was produced resulted in new seedlings. Mortality was substantial immediately following emergence. Most seedlings emerged without shading from understory vegetation, but few survived. Most seedlings alive after one year had emerged beneath live cover; by the end of two summers only shaded seedlings were alive. 63–85% of new seedlings initiated in clusters attributed to rodent caching, and seedlings originating in clusters remained a substantial proportion of the cohort (60%) after two seasons. Results suggest that understory vegetation in these types of stands facilitates the recruitment of ponderosa pine seedlings, most likely by buffering them against environmental stressors during the seedling establishment phase. They also reveal that seed caching rodents are highly active in seed redistribution and can exert a lasting influence on seedling recruitment.     

花楸种子变温处理及造林技术

对花楸种子进行沙藏变温处理能够打破种子深度休眠,促进发芽。另外采用和落叶松等针叶树混交,成活率高,生长快。     

楸树叶面积与叶形指数的回归分析

本实验采用数码相机拍照,利用CAD．图像处理方法得到楸树叶片的面积,并同时测量叶长、叶宽计算出长宽比、长乘宽,通过逐步回归的方法对与叶面积有关的指标进行筛选。用最后得到的最能反映叶面积的指标长乘宽,建立不同叶形的叶片长乘宽与叶面积的一元线形回归预测模型方程,分别是倒心形Y=0．652x＋16．041,决定系数R2=0．7221,水滴形Y=0．5058x＋112．47,决定系数R2=0．4994,三裂Y=0．3211x＋155．98,决定系数R2=0．6006,分析认为可将叶长乘宽作为重要的叶形指数,进行楸树生长等相关研究。     

种子不同保存方法及播种时间对水紫树发芽和幼苗生长的影响

以田间调查数据为基本资料,对极耐湿的水紫树种子进行沙藏和冷藏保存,分别于3月17日、4月1日播种,对其播种苗的发芽、幼苗生长进行统计分析。结果表明：冷藏保存的种子发芽慢,持续时间长,而沙藏保存的种子发芽相对较快,持续时间短,苗齐较冷藏处理早21 d。沙藏保存种子的幼苗高、地径的全年平均累计生长量极显著大于冷藏处理。3月17日播种的幼苗高、地径的全年平均累计生长量极显著大于4月1日播种幼苗的生长量。     

城市近郊林土壤种子库研究

对沈阳、抚顺两城市交接的近郊林区不同林分类型土壤种子库进行研究.结果表明:不同林分类型土壤种子库均具有较好的储量和活力,其储量大小顺序为:乔木林地＞灌木林地＞宜林荒地,土壤种子库物种多样性指数大小顺序为:灌木林地＞乔木林地＞宜林荒地,种子种类以乡土树种为主;数量化分析表明坡位对乔木林地及灌木林地中土壤种子库储量和活力影响较大,而植被覆盖度对宜林荒地中土壤种子库储量和活力影响较大.     

Effects of logging, liana tangles and pasture on seed fate of dry forest tree species in Central Brazil

We assessed seed fate of six tree species in a seasonally deciduous forest of Central Brazil. Seed germination, predation, removal and death caused by pathogens or dessication were followed in an undisturbed forest site, a logged forest site, and an active pasture. In both forest types we sowed seeds under patches of liana tangles (Low Forest habitat) and under patches of mature forest (High Forest habitat). Seeds were monitored weekly for up to 8 months. Overall, small-seeded species (Astronium fraxinifolium and Tabebuia impetiginosa) had high germination, although germination was lower in the pasture due to insect predation. The hard-coated seeds of Erythrina sp. had higher germination rates in the pasture. The large-seeded Cavanillesia arborea and Swartzia multijuga had high seed removal (ca. 90%) in the pasture and under High Forest, but only 23–34% under Low Forest in the logged site. Lower removal of large seeds in Low Forest suggests that this habitat conceals the seeds and constrains the movement of large mammals. Both species were favored in Low Forest, suggesting that caution should be used when managing lianas. When protected from predation, seeds with a soft coat and high water content, such as the seeds of Eugenia dysenterica and S. multijuga, died from desiccation in the pasture. Restoration efforts must take into consideration high mortality caused by desiccation or insect predation in abandoned pastures of dry forests.     

七叶树种子级别与种子活力及苗木品质的相关性研究

以七叶树种子为实验材料，并分级别绘制了种子的吸水曲线、测定了种子活力指数；同时，把种子按不同级别温室播种，分别测定其幼苗的苗高、地径、水势及电导率并作相关性分析。结果表明，种子级别与种子活力有相关性，中粒种子活力最强，小粒种子活力最弱；种子级别与苗木品质也有极显著的相关性，大粒种子萌发苗品质最好，小粒种子萌发苗品质最差。     

Differentiation of seed germination and early seedling growth in ten provenances ofEucalyptus microtheca

An investigation of seed germination and early seedling growth ofEucalyptus microtheca was based on seed collection from 10 widely separated provenances in Australia. Genetic variation of seed germination and early seedling growth was observed among a series of provenances whose natural habitats range from different climatic condition. In the ten provenances, both the model of seed relative germination percentage and the model of seed total germination percentage fitted Logistic regression [y=a/(1+exp(-ex+b))]. In comparison with provenances from four high temperature (mean annual maximum temperature >30.0 °C; mean annual minimum temperature >17.0 °C) areas, six low temperature (mean annual maximum temperature <30.0 °C; mean annual minimum temperature <17.0 °C) areas showed the fast germination rate and the high total germination percentage. For each provenance we have 45 seedlings equally divided into three watering levels (100%, 50%, and 25% of field capacity), and studies on relationship between early seedling growth and climatic factors of the natural habitat of provenance. In control treatment, height growth of the seedling has been associated with intrinsically the driest quarter precipitation in the seed collection areas of provenance. In all the treatments, length growth of the biggest leaf of the seedling was related to mean annual maximum temperature and mean annual minimum temperature in origin of provenance. In contrast, basal diameter growth of the seedling was related to mean annual minimum temperature of the seed collection areas in water stress treatment. From an ecological viewpoint, the fast germination rate and the high total germination percentage of the seed and rapid early growth of the seedling appear to be favourable adaptations to the climatic conditions prevailing in the natural habitat of provenance. Responsible Editor: Chai Ruihai     

冷箭竹种子特性及自然更新

冷箭竹(Bashania fangina)为四川西部特产,分布于海拔2100～3600米之针阔叶混交林和亚高山针叶林下,是邛崃山系大熊猫的主食竹种。1983年,该竹种发生大面积的开花、结实而枯死,开花面积占竹子分布总面积的65～95％。据史书记载,冷箭竹的开花周期为50～60年。竹种萌发成林约需10年以上时间,从而导致以此为食的大熊猫发生周期性的食物危机。因此,研究冷箭竹的种子特性及幼苗更新规律,采取人为措施加速幼苗的更新复壮,这不仅是研究该物种生态生物学特性的需要,也是保护大熊猫,促进其种群繁衍的主要途径。本文描述了冷箭竹种子的净度、含水量、千粒重、生活力、发芽率、休眠期和生命期限等特性,并介绍了该竹和在自然条件下林地上的种子数量、种子发芽持续时间、幼苗更新的期数量的动态变化、幼苗成活率和幼苗的生长变化。通过这些研究,初步掌握了冷箭竹种子生命活动的一般规律和幼苗生长的基本特征,为提高高山地区冷箭竹的自然更新能力提供了科学依据。     

Bird communities and seedling recruitment in restoring seasonally dry forest using the framework species method in Northern Thailand

This study examined the effects of framework trees, planted in 1998, and bird community on the natural recruitment of tree seedling species in a forest restoration experiment designed to test the framework species method of forest restoration established by Chiang Mai University’s Forest Restoration Research Unit (FORRU-CMU). Tree seedlings establishing beneath five framework tree species: Erythrina subumbrans (Hassk.) Merr., Hovenia dulcis Thunb., Melia toosendan Sieb. & Zucc., Prunus cerasoides D.Don and Spondias axillaries Roxb., were surveyed. Five trees of each species were selected in the 8-year-old trial plots. Birds visiting each tree were observed to determine possible seed dispersal activities. Thirty-six tree seedling species were found beneath the selected trees, of which 11 were wind-dispersed and 25 were animal-dispersed. The population density of animal-dispersed tree seedlings was higher than the wind-dispersed seedlings beneath all selected framework trees. The sample plots beneath P. cerasoides supported the highest population density of tree seedlings. Forty-nine bird species were recorded visiting the framework trees between July 2006 and June 2007. Non-frugivorous birds were recorded more frequently than the frugivorous birds. The effects of birds on seedling recruitment were different among each of the selected framework tree. Bigger trees, which attracted high number of birds by providing food resources, roosting and nesting sites may increase the seed deposition more than smaller trees with fewer attractants.     

凉风坳亚热带次生常绿阔叶林土壤种子库和种子雨的特征研究

选择四川省沐川县凉风坳亚热带次生常绿阔叶林为研究对象,在2004年5月、2004年12月和2005年5月对其林下土壤种子库、种子雨以及种子天然萌发状况等方面进行系统调查及分析研究。结果显示,种子下落前后土壤种子库储量,变动为643.2～889.2粒.m-2,物种和种子密度多年生草本物种均占优势。天然林土壤种子库Shannon-Wiener指数均在2004年5月最大,2004年12月次之,2005年5月最小;Simpson指数却和Shannon-Wiener指数H＇的变化趋势相反;Pielou均匀度指数在2004年12月有所降低,此后又升高。种子均呈明显垂直分布,其中0～2 cm土层种子数量最多,其次是枯枝落叶层,其数量随着土层加深而降低。土壤种子库年输入活力种子142.3粒.m-2,年输出活力种子268.9粒.m-2,动物取食是造成种子损失的主要原因。对照人工林土壤种子库物种和密度均低于天然林地;种子库与地上植被物种相似性系数,人工林较高,而天然次生常绿阔叶林较低。     

Assessing the potential for ash canopy tree replacement via current regeneration following emerald ash borer-caused mortality on southeastern Michigan landscapes

The emerald ash borer (EAB) has killed millions of ash trees in Michigan and at least fourteen other states since its first detection near its introduction point in metropolitan Detroit in 2002. Despite overstory ash mortality near 100% in many areas, ash seedling and saplings remain unaffected and provide the potential for ash re-establishment into the canopy of deciduous forests of the region. We examined the potential for ash re-establishment in areas of heavy mortality by measuring the status and change of ash regeneration at 45 sites across southeastern Lower Michigan in 2007 and 2009. Ash regeneration was found to be abundant in all forest types, particularly in the smallest height classes, though it was more abundant in ash species of upland forests compared to those of lowland forests. New seedlings 1-2 years old were less common than other regeneration size classes, and declined substantially between 2007 and 2009, suggesting a depletion of the ash seed bank in these forests. Regeneration density was not explained well by the presence of overstory ash prior to EAB introduction, suggesting that competitive interaction with other tree species in higher canopy strata is an important driver of ash regeneration density. Regeneration is sufficient to replenish overstory ash to pre-EAB levels in upland forests, though ecological changes caused by overstory ash mortality in lowland areas may affect local hydrology in a way that reduces opportunities for tree regeneration. Despite abundant ash regeneration, it remains unclear whether ash will recover to the overstory of forests in southeastern Michigan because of uncertainties in the future dynamics of EAB in the region.     

Seed dispersal,dormancy, seedbank and regeneration of Amur corktree

Amur corktree (Phellodendron amurense Rupr.) is one of the important accompanying species of Korean pine (Pinus Koraiensis Sieb et Zuce) broadleaved forest in northeast China. This paper reports on the seed dispersal, dormancy, seed bank and regeneration of amur corktree. Seeds of amur corktree arc dispersed mainly by the aid of birds and gravity. The aril of the seeds may increase seed longevity. In natural secondary forests, the store of the seed bank may reach 750, 000 seeds per heetare. Regeneration was correlated wirh disturbance, and was best under intensive disturbance.     

Seedling emergence, growth, and allocation of Oriental bittersweet: effects of seed input, seed bank, and forest floor litter

The establishment of invasive plant populations is controlled by seed input, survival in the soil seed bank, and effects of soil surface disturbance on emergence, growth, and survival. We studied the invasive vine Celastrus orbiculatus Thunb. (Oriental bittersweet) to determine if seedlings in forest understory germinate from the seed bank or from seed rain. We also conducted a greenhouse experiment to investigate the role of leaf litter mass and physical texture on seedling survival, growth, and allocation. In the understory of an invaded mixed hardwood forest, we measured seed input, seedling emergence with seed rain, and seedling emergence without seed rain. Mean seed rain was 168 seeds m⁻²: mean seedling emergence was 107 m⁻², and there was a strong correlation between seed rain and seedling emergence. The ratio of seedlings to seed input (0.61) was close to the seed viability (0.66) leaving very few seeds to enter the seed bank. Seed bank germination under field conditions was low (1 seedling m⁻²). Soil cores were incubated in a greenhouse to determine seed bank viability, and germination from these soil cores did not occur. To determine how litter affects seedling establishment and growth, we measured seedling emergence and biomass allocation in a greenhouse experiment. Seeds were placed below intact and fragmented deciduous leaf litter in amounts ranging from zero to the equivalent of 16 Mg ha⁻¹. Seedling emergence was not affected by fragmented litter, but decreased to <20% as intact litter increased to 16 Mg ha⁻¹. Increasing litter resulted in greater allocation to hypocotyl and less to cotyledon and radicle, and this effect was greater in intact litter. C. orbiculatus seedlings achieve emergence through forest floor litter through plasticity in allocation to hypocotyl growth. The low survival of C. orbiculatus in the seed bank suggests that eradication of seedling advance regeneration and adult plants prior to seed rain may be an effective control strategy. However, the intact forest floor litter of an undisturbed forest will not prevent seedling establishment.