Productivity,grazing utilization,forage quality and primary production controls of species‐rich alpine grasslands with Nardus stricta in northern Spain

Species‐rich alpine grasslands with Nardus stricta are important communities for both animal production and environmental conservation in Europe. We selected two contrasting types of Nardus grasslands (mesic and wet) within a rangeland of northern Spain and measured annual above‐ground net primary productivity (ANPP), botanical components, forage utilization and their respective seasonal patterns, during a 5‐year period. We analysed their chemical properties and recorded soil moisture and temperature in order to construct models able to explain grassland productivity. Mean annual ANPP of mesic Nardus grassland was about half (216 g DM m^?2 year^?1; ±29·8 s.e.) that of the wet grassland (406 g DM m^?2 year^?1; ±54·3 s.e.), with significant intra‐ and interannual variability. Mesic grassland, with a more important contribution of forbs and legumes over graminoids in its botanical composition, was the preferred forage source of grazing livestock and showed better chemical properties in spring and early summer. In summer and autumn, wet grassland had a higher utilization owing to its ability to maintain high biomass production. This was partially explained by soil moisture, a limiting factor of mesic grassland productivity. Our results provide new and relevant information on key aspects of species‐rich alpine Nardus grasslands, potentially useful for the definition of management options for these habitats of priority conservation.     

Spatial and structural uniformity of lowland agricultural grassland in England: a context for low biodiversity

The contention that agricultural grassland within the UK is spatially and structurally uniform across wide differences in livestock farming intensity is examined. Total nitrogen (N) input was used as a surrogate for management intensity. Farms with average annual N inputs of >200 kg ha^?1 were categorized as highly intensive, those with N inputs of 50–200 kg N ha^?1 as moderately intensive and those with inputs of <50 kg N ha^?1 were categorized as extensive. Four farms within each management category were selected in two discrete regions: one in south‐west England, typifying a landscape dominated by agricultural grassland, the other in south‐east England with a more mixed‐farming landscape. Specific N input and management data, and sward botanical composition and structure, were obtained on four fields per farm. The south‐west region had a higher proportion of farms with dairy cattle than in the south‐east England region. The average N input and stocking rates were higher within the intensive‐management category farms in south‐west England than in south‐east England. Old permanent grasslands contained variable amounts of sown species, such as Lolium perenne, and were dominated by a few (generally <5) non‐sown grass species. Grassland that received >75 kg N ha ^?1 year^?1 contained <7 plant species and <3 forb species m^?2. Grassland communities with >12 plant species and >5 forb species m^?2 were only found in study fields that received <50 kg N ha^?1 year^?1. Grasslands with >10 forb species m^?2 were found only in sites receiving <15 kg N ha^?1. Of importance at a landscape scale was ubiquity of species‐poor and similar grassland plant communities across the management intensity range within and between livestock farms. Grassland in the extensive management category was similar in structure in terms of sward height and/or sward density to more intensively managed grasslands. This study revealed, albeit in two lowland sample regions in the UK, the ubiquity of grass‐dominated, species‐poor and structurally uniform grasslands, irrespective of apparently broad differences in farming intensity. The consequences of this spatial uniformity on grassland biodiversity are likely to be profound.     

The effect of stocking rate and calving date on grass production,utilization and nutritive value of the sward during the grazing season

A 2‐year whole‐farm system study compared the accumulation, utilization and nutritive value of grass in spring‐calving grass‐based systems differing in stocking rate (SR) and calving date (CD). Six treatments (systems) were compared over two complete grazing seasons. Stocking rates used in the study were low (2·5 cows ha^?1), medium (2·9 cows ha^?1) and high (3·3 cows ha^?1), respectively, and mean CDs were 12 February (early) and 25 February (late). Each system had its own farmlet of eighteen paddocks and one herd that remained on the same farmlet area for the duration of the study. Stocking rate had a small effect on total herbage accumulation (11 860 kg DM ha^?1 year^?1), but had no effect on total herbage utilization (11 700 kg DM ha^?1 year^?1). Milk and milk solids (MS; fat + protein) production per ha increased by 2580 and 196 kg ha^?1 as SR increased from 2·5 to 3·3 cows ha^?1. Milk production per ha and net herbage accumulation and utilization were unaffected by CD. Winter feed production was reduced as SR increased. Increased SR, associated with increased grazing severity, resulted in swards of increased leaf content and nutritive value. The results indicate that, although associated with increased milk production per ha, grazed grass utilization and improved sward nutritive value, the potential benefits of increased SR on Irish dairy farms can only be realized if the average level of herbage production and utilization is increased.     

Effects of functional groups and species richness on biomass constituents relevant for combustion: results from a grassland diversity experiment

Grassland biomass has been identified as a potential energy source. The combustion of mature and fibrous biomass, as occurs in extensive grasslands managed with low cutting frequencies, is one possible conversion technique. This study tested the relationship between plant diversity and biomass constituents relevant for combustion, as they determine energy content, energy yield and emission and corrosion risks. The biomass from a biodiversity experiment, with a species richness (SR) gradient of 1–60 species from Central European mesophilic grasslands divided into four functional groups (grasses, legumes, small and tall herbs), was harvested twice a year (in 2008 and 2009). The higher heating value (HHV) was estimated from carbon, hydrogen and oxygen contents to give insight into the energy potential of the species mixtures. The potential risk of emission and corrosion was assessed by analysing ash content, potassium, calcium, magnesium, nitrogen, sulphur and chloride content. HHV was independent of SR, and the overall mean was 18·13 MJ kg^?1 DM. Biomass and gross energy (GE) yield were positively affected by SR. The presence of legumes in a mixture resulted in increased HHV, biomass yield and GE, irrespective of the level of SR. Annual GE varied between 59 (average of monocultures) and 152 GJ ha^?1 year^?1 (mixture of sixty species). The concentration of ash‐forming elements was generally high, suggesting a pre‐treatment of the biomass prior to combustion. Emission‐ and corrosion‐related constituents were clearly affected by the different functional groups, and sulphur and nitrogen both declined with SR. The results of this study show that high SR in experimental grassland communities is beneficial for the energy output and that legumes play a key role for the energy potential. However, identifying a functional group as being solely beneficial or disadvantageous for fuel quality was difficult.     

Biomass production and nutrient removal by tropical grasses subsurface drip‐irrigated with dairy effluent

Effluent lagoons on dairy farms can overflow and potentially pollute adjacent land and associated water bodies. An alternative solution to effluent disposal is needed by dairy operators in island environments. An attractive win‐win alternative is to recycle nutrients from this resource through effluent irrigation for forage grass production that minimizes environmental pollution. This study assessed biomass production and nutrient removal by, and high application rates to, tropical grasses that were subsurface drip‐irrigated with dairy effluent. Four grass species – Banagrass (Pennisetum purpureum K. Schumach.), California grass (Brachiaria mutica (Forssk.) Stapf.), Stargrass (Cynodon nlemfuensis Vanderyst) and Suerte grass (Paspalum atratum Swallen) – were subsurface (20–25 cm) drip‐irrigated with effluent at two rates based on potential evapotranspiration (ET_p) at the site (Waianae, Hawaii) ?2·0 ET_p (16 mm d^?1 in winter; 23 mm d^?1 in summer) and 0·5 ET_p (5 mm d^?1 in winter; 6 mm d^?1 in summer). Treatments were arranged in an augmented completely randomized design. Brachiaria mutica and P. purpureum had the highest dry‐matter yield (43–57 t ha^?1 year^?1) and nutrient uptake especially with the 2·0 ET_p irrigation rate (1083–1405 kg ha^?1 year^?1 N, 154–164 kg ha^?1 year^?1 P, 1992–2141 kg ha^?1 year^?1 K). Average removal of nutrients by the grasses was 25–94% of the applied nitrogen, 11–82% of phosphorus and 2–13% of the potassium. Average values of crude protein (90–160 g kg^?1), neutral detergent fibre (570–620 g kg^?1) and acid detergent fibre (320–360 g kg^?1) were at levels acceptable for feeding to lactating cattle. Results suggest that P. purpureum and B. mutica irrigated with effluent effectively recycled nutrients in the milk production system.     

Grazing and fertilization influence plant species richness via direct and indirect pathways in an alpine meadow of the eastern Tibetan Plateau

Grazing and fertilization influence plant species richness through multiple pathways, but the exact pathways and their relative contributions are unclear. Here, we report on a 3‐year field experiment on an alpine meadow in which we manipulated grazing and fertilization simultaneously to assess their direct and indirect effects on plant species richness. Results indicated that both grazing and fertilization significantly influenced plant species richness via above‐ and belowground pathways. Specifically, the direct effect of grazing (?0·30, effect size) on species richness was negative, mainly due to selective feeding. This was comparable, in magnitude, to a positive indirect effect of grazing (0·36) on species richness, due to decreased aboveground biomass and increased belowground biomass. The direct effect of fertilization (?0·34) on species richness was negative, which was attributed to the changes of soil pH and available nitrogen; this effect was double the indirect effect (?0·16), which largely stemmed from the increased aboveground biomass rather than changes in belowground biomass. No significant interaction of grazing and fertilization on species richness was observed, but the effect of fertilization on species richness was masked by the effect of grazing in the combined treatment. It was concluded that moderate nitrogen addition (<51·0 kg N ha^?1 year^?1) could increase local pasture productivity while also sustaining species richness stability.     

The efficiency of nitrogen in cattle manures when applied to a double‐annual forage cropping system

The use of cattle manure (CM) for fertilization presents challenges for optimizing nitrogen (N) use. Our work aimed to assess N efficiencies, in a 6‐year experiment with three biennial rotations of four crops: oat–sorghum (first year) and ryegrass–maize (second year) in a rainfed humid Mediterranean area of Spain. Fertilization treatments included the following: control (no N), 250 kg mineral N ha^?1 year^?1 (250MN), three CM rates (supplying 170, 250 and 500 kg N ha^?1 year^?1) and four treatments where the two lowest CM rates were complemented with either 80 or 160 kg mineral N ha^?1 year^?1. Treatments were distributed randomly in each of three blocks. Maximum dry‐matter yield (~44–49 t ha^?1 rotation^?1) was achieved in the third rotation, and only the control and the 170CM yielded significantly less. Within the limitations of the EU Nitrate Directive, the N steady state supply of 170CM always requires a complement of mineral N (80 kg N ha^?1) to maximize N agronomic efficiency. The maximum N‐fertilizer replacement value (250CM vs. 250MN) was 0·67, without significant differences between the two treatments in other N‐related efficiency indexes, which indicates that plants took advantage of residual‐N effects. Nitrogen losses by leaching in the 250CM treatment were around 5–7% of the N applied. This reinforces the sustainability of manure recycling in long cropping seasons.     

Herbage dry‐matter production and forage quality of three legumes and four non‐leguminous forbs grown in single‐species stands

Agronomic data on most broad‐leaved species of grasslands are scarce. The aim of this study was to obtain novel information on herbage DM yield and forage quality for several forb species, and on species differences and seasonal patterns across harvests and in successive years. Four non‐leguminous forbs [salad burnet (Sanguisorba minor), caraway (Carum carvi), chicory (Cichorium intybus) and ribwort plantain (Plantago lanceolata)] and three leguminous forbs [yellow sweet clover (Melilotus officinalis), lucerne (Medicago sativa) and birdsfoot trefoil (Lotus corniculatus)] and a perennial ryegrass–white clover mixture were investigated in a small‐plot cutting trial in Denmark during 2009 and 2010. Plots were harvested four times per year. On average, annual herbage yield was highest for lucerne (15·4 t DM) and grass–white clover (12·5 t DM ha^?1), and lowest for salad burnet (4·6 t DM ha^?1) and yellow sweet clover (3·9 t DM ha^?1). Ribwort plantain and lucerne had the highest concentrations of acid detergent fibre (339 and 321 g kg^?1 DM respectively) and lignin (78 and 67 g kg^?1 DM respectively); contents in other species were similar to grass–white clover (275 and 49 g kg^?1 DM respectively). No common feature was found within the functional groups of non‐leguminous forbs and leguminous forbs, other than higher crude protein contents (198–206 g kg^?1 DM) in the legumes. DM yield and fibre content were lowest in October. Digestibility declined with higher temperature and increasing fibre content. Results are discussed in terms of the potential of forbs to contribute to forage resources in farming practice.     

Biomass yield and nutrient content of a tropical mulberry forage bank: effects of season,harvest frequency and fertilization rate

Effects of season (SEAS), organic fertilization with laying hen manure (N) and harvest frequency (FREQ) on forage yield and composition were evaluated in a high‐density mulberry forage bank, established under low‐input tropical farming conditions. The experiment, replicated over two consecutive years, was arranged in 18 treatments resulting from a 3 × 3×2 factorial design which combined three FREQ (60, 90 and 120 d), 3 N rates (100, 300 and 500 kg N ha^?1 year^?1) and the two seasons (RAIN and DRY). Forage yield, leaf‐to‐stem ratio, the edible fraction (EDIB, %), nutrient content and the maturity indexes were monitored. Either forage yield or qualities by plant fraction were strongly affected by SEAS, FREQ, N rate and by their interactions. Effects of FREQ and N were modulated by SEAS. Increasing FREQ during RAIN negatively affected EDIB yield and quality after 90 d, while increasing ligneous fraction. During DRY, longer FREQ allowed higher total forage and EDIB yields. Leaves were more abundant in RAIN, and its yield was increased with fertilization but only during this season. Maturity indexes were higher in DRY than in RAIN. Under the conditions of this experiment, harvesting at 60 and 90 d in RAIN and DRY, respectively, with an N rate of 300 kg N ha^?1 year^?1 seems the best agronomic choice for an optimal compromise between forage yield and nutritive value.     

Foliar N/P ratio and nutrient limitation to vegetation growth on Keerqin sandy grassland of North‐east China

To examine whether the critical leaf N/P ratios (of 14, 16) are valid to test nutrient limitation in the context of semi‐arid sandy grasslands, an experiment was conducted on a Keerqin sandy grassland in North‐east China to investigate the responses of plant biomass and nutrient concentrations to fertilization. Plant biomass production and leaf nutrient concentrations were measured after five consecutive years of fertilization with N (20 g N m^?2 year^?1) and/or P (10 g P₂O₅ m^?2 year^?1). Nitrogen fertilization increased the shoot biomass by twofold and consequently the shoot/root ratio, whereas P fertilization had little effect on either shoot biomass or shoot/root ratio. Leaf N/P ratio varied among species with an average of 5·6 in the control, while the mean leaf N/P ratio (7·5) under the N fertilization treatment remained below the threshold of 14. Our results suggest that the critical N/P ratio (14, 16) is not applicable as a test for nutrient limitations in the context of semi‐arid, sandy grassland.     

‘MaxClover’ grazing experiment: I. Annual yields,botanical composition and growth rates of six dryland pastures over nine years

Six dryland pastures were established at Lincoln University, Canterbury, New Zealand, in February 2002. Production and persistence of cocksfoot pastures established with subterranean, balansa, white or Caucasian clovers, and a perennial ryegrass‐white clover control and a lucerne monoculture were monitored for nine years. Total annual dry‐matter (10.0–18·5 t DM ha^?1) and sown legume yields from the lucerne monoculture exceeded those from the grass‐based pastures in all but one year. The lowest lucerne yield (10 t ha^?1 yr^?1) occurred in Year 4, when spring snow caused ungrazed lucerne to lodge and senesce. Cocksfoot with subterranean clover was the most productive grass‐based pasture. Yields were 8·7–13·0 t DM ha^?1 annually. Subterranean clover yields were 2·4–3·7 t ha^?1 in six of the nine years which represented 26–32% of total annual production. In all cocksfoot‐based pastures, the contribution of sown pasture components decreased at a rate equivalent to 3·3 ± 0·05% per year (R² = 0·83) and sown components accounted for 65% of total yield in Year 9. In contrast, sown components represented only 13% of total yield in the ryegrass‐white clover pastures in Year 9, and their contribution declined at 10·1 ± 0·9% per year (R² = 0·94). By Year 9, 79% of the 6.6 t ha^?1 produced from the ryegrass‐white clover pasture was from unsown species and 7% was dead material. For maximum production and persistence, dryland farmers on 450–780 mm yr^?1 rainfall should grow lucerne or cocksfoot‐subterranean clover pastures in preference to ryegrass and white clover. Inclusion of white clover as a secondary legume component to sub clover would offer opportunities to respond to unpredictable summer rainfall after sub clover has set seed.     

Performance and environmental effects of forage production on sandy soils. I. Impact of defoliation system and nitrogen input on performance and N balance of grassland

Grassland and its management is central to the productivity of and nitrogen (N) losses from dairy farms in north‐west Europe. Botanical composition, production and N surplus of grassland were assessed during five consecutive years. The experiment consisted of all combinations of five defoliation systems: cutting‐only (CO), rotational grazing (GO), grazing + one (MSI) or two silage cuts (MSII) and simulated grazing (SG). Four mineral N fertilization rates (0–300 kg N ha^?1 year^?1) and two slurry levels (0 and 20 m³ slurry ha^?1 year^?1) were applied. Fertilizer N was more efficient in producing net energy (NEL) in grazing‐dominated, low white clover systems (GO and MSI systems: 70 and 88 MJ NEL kg^?1 N) than in white clover‐rich systems (MSII, CO and SG systems: ≤60 MJ NEL kg^?1 N). While sward productivity in system MSI was similar to that in system GO, system MSII benefited from increased N₂ fixation at low N rates. There were small differences in NEL concentrations of the herbage between defoliation systems. Crude protein concentration of the herbage increased with increasing N supply from fertilizer, excreta and N₂ fixation. N surpluses (?63 to +369 kg N ha^?1 year^?1) increased with increasing grazing intensity and increasing N fertilization rate. The average response in N surplus applied was 0·81, 0·59, 0·40, 0·33 and 0·24 kg N ha^?1 in systems GO, MSI, MSII, CO and SG respectively.     

Production and nutritional value of sorghum and black oat forages under nitrogen fertilization

The effect of nitrogen (N) fertilization on the dry‐matter (DM) yield and nutritional value of sorghum (Sorghum sp., cv. Jumbo) and black oat (Avena strigosa cv., IPR 61) was investigated in the context of forage and livestock production in southern Brazil. Sorghum was cultivated with 0, 37·5, 75, 150, 225, 300 and 375 kg N ha^?1 during the summer crop seasons of 2010/11 and 2011/12. Black oat received 0, 40, 80, 120, 160, 200 and 240 kg N ha^?1 in the winter of 2011. According to the adjusted polynomial regression, sorghum DM yield increased in response to N up to 288 (12·9 t ha^?1) and 264 kg ha^?1 (5·6 t ha^?1) in 2010/11 and 2011/12 respectively. Crude protein (CP) content of sorghum was highest at 349 and 328 kg N ha^?1, but in vitro dry‐matter digestibility (IVDMD) was highest at 212–207 kg N ha^?1 in 2010/11 and 2011/12 respectively. Sorghum neutral detergent fibre (NDF) and acid detergent fibre (ADF) were not affected by N fertilization. In black oat, the maximum DM yield (6·0 t ha^?1) was obtained with 187 kg N ha^?1; the IVDMD, NDF and ADF were not affected by N fertilization, but the CP content increased up to 220 kg N ha^?1. It is concluded that these forage species can improve the year‐to‐year amount and quality of forage produced but high rates of N fertilizer are required to achieve high yields. Fertilizer N rates of 210–280 kg N ha^?1 in sorghum and 180 kg N ha^?1 in black oat in the crop rotation provide the greatest responses in DM yield consistent with good nutritional quality for livestock production.     

An economically based evaluation index for perennial and short‐term ryegrasses in New Zealand dairy farm systems

Grazed pastures based on ryegrass species provide most of the feed for dairy cattle in New Zealand. There are many cultivars of perennial (Lolium perenne), annual and Italian (L. multiflorum), and hybrid (L. boucheanum) ryegrasses available for dairy farmers to use in pasture renewal. This study describes an index which ranks ryegrass cultivars relative to a genetic base according to the estimated economic value (EV) of seasonal dry matter (DM) traits. A farm system model was used to derive EVs (Grazed pastures based on ryegrass species provide most of the feed for dairy cattle in New Zealand. There are many cultivars of perennial (Lolium perenne), annual and Italian (L. multiflorum), and hybrid (L. boucheanum) ryegrasses available for dairy farmers to use in pasture renewal. This study describes an index which ranks ryegrass cultivars relative to a genetic base according to the estimated economic value (EV) of seasonal dry matter (DM) traits. A farm system model was used to derive EVs ($ ha^?1 calculated as change in operating profit divided by unit change of the trait) for additional DM produced in different seasons of the year in four regions. The EV of early spring DM was consistently high across all regions, whereas EV for late spring DM was moderate to low. Genotype × environment analysis revealed significant reranking of DM yield among ryegrass cultivars across regions. Hence, separate performance values (PVs) were calculated for two mega‐environments and then combined with the corresponding season and region EV to calculate the overall EV for twenty‐three perennial ryegrass and fifteen short‐term ryegrass cultivars. The difference in operating profit between the highest ranked and lowest ranked perennial ryegrass cultivar ranged from $556 ha^?1 to $863 ha^?1 year^?1 depending on region. For short‐term ryegrasses used for winter feed, the corresponding range was $394 to $478 ha^?1 year^?1. Using PV for DM yield, it was estimated that plant improvement in perennial ryegrass has added $12–$18 ha^?1 year^?1 (depending on region) operating profit on dairy farms since the mid‐1960s.     

Herbage and nitrogen yields,fixation and transfer by white clover to companion grasses in grazed swards under different rates of nitrogen fertilization

In grass–legume swards, biologically fixed nitrogen (N) from the legume can support the N requirements of the grass, but legume N fixation is suppressed by additional fertilizer N application. This study sought to identify a fertilizer N application rate that maximizes herbage and N yields, N fixation and apparent N transfer from white clover to companion grasses under intensive grazing at a site with high soil‐N status. During a 3‐year period (2011–2013), swards of perennial ryegrass and of perennial ryegrass–white clover, receiving up to 240 kg N ha^?1 year^?1, were compared using isotope dilution and N‐difference methods. The presence of white clover increased herbage and N yields by 12–44% and 26–72%, respectively. Applications of N fertilizer reduced sward white clover content, but the effect was less at below 120 kg N ha^?1. The proportion of N derived from the atmospheric N fixation was 25–70%. Nitrogen fixation ranged from 25 to 142 kg N ha^?1 measured using the isotope dilution method in 2012 and from 52 to 291 kg N ha^?1 using the N‐difference method across all years. Fertilizer N application reduced the percentage and yield of fixed N. Transfer of N from white clover to grass was not confirmed, but there was an increased N content in grass and soil‐N levels. Under intensive grazing, the maximum applied N rate that optimized herbage and N yields with minimal effect on white clover content and fixation rates was 60–120 kg N ha^?1.     

Effects of nitrogen fertilization and cutting intensity on the agronomic performance of warm‐season grasses

Responses of grasses to N fertilization are affected by cutting intensity although little is known regarding the interactions of these factors in warm‐season grasses. Pre‐cutting canopy height, herbage accumulation and changes in the nutritive value of warm‐season grasses in response to four different management strategies were assessed from October 2011 to September 2014. Treatments included two cutting intensities (70 vs. 50% depletion of canopy height set by 95% light interception), two N fertilization levels (zero vs. 300 kg N ha^?1 year^?1) and six perennial C₄ grass species (Axonopus catharinensis; Cynodon spp. hybrid Tifton 85; Hemarthria altissima cv. Flórida; Megathyrsus maximus cv. Aruana; Paspalum notatum cv. Pensacola; and Urochloa brizantha cv. Marandu) grown in monoculture in a factorial experimental design. Canopy height varied among grass species, cutting intensity and N treatments, mainly among seasons, indicating that more than one management target (i.e. canopy height) existed throughout the plant growth cycle for each species. The largest herbage accumulation occurred in the N fertilization treatments for most species, regardless of cutting intensity. Nitrogen fertilization and 50% depletion of canopy height increased the leaf proportion and decreased the neutral detergent fibre content. Overall, N fertilization had a stronger positive impact than cutting intensity on the acid detergent fibre content, dry‐matter digestibility and crude protein content, but the magnitudes of the responses were species‐specific.     

Changes in the area of permanent grassland and its implications for the provision of bioenergy: Slovakia as a case study

Throughout Europe, grasslands are managed primarily for agricultural production but also provide a range of ecosystem services, the magnitude of which is influenced by their area, management and the abiotic properties of the ecosystem. The grassland area in Europe has been affected by significant changes in recent decades, including abandonment and conversion to arable land. This study presents an assessment of changes in the permanent grassland area with reference to Slovakia and provides an assessment of the bioenergy potential of permanent grassland surplus to agricultural needs. In 2015, of the total permanent grassland area (868,000 ha) 42% had ceased to be used as feed for milk or meat production and had been abandoned. Results from field experiments show that low‐level improvement of permanent grasslands, e.g., by oversowing commonly sown agricultural grassland species and low‐dose fertilization (up to 30 kg N ha^?1 year^?1) could provide herbage biomass of 1.7 million tonnes of herbage dry matter per year. This would be sufficient to generate an energy output of approximately 31.3–31.9 PJ/year (4.5% of the current gross inland energy consumption of Slovakia). Low‐intensity use of the currently surplus permanent grassland is also considered to have potential beneficial outcomes in terms of delivery of ecosystem services. Results are also discussed in the context of countries beyond the case‐study area.     

Radiation‐use efficiency for forage kale crops grown under different nitrogen application rates

Crop growth is related to radiation‐use efficiency (RUE), which is influenced by the nitrogen (N) status of the crop, expressed at canopy level as specific leaf N (SLN) or at plant level as N nutrition index (NNI). To determine the mechanisms through which N affects dry‐matter (DM) production of forage kale, results from two experiments (N treatment range 0–500 kg ha^?1) were analysed for fractional radiation interception (RI), accumulated radiation (R_acc), RUE, N uptake, critical N concentration (N_c), NNI and SLN. The measured variables (DM, RI and SLN) and the calculated variables (NNI, R_acc and RUE) increased with N supply. RUE increased from 0·74 and 0·89 g MJ^?1 IPAR for the control treatments to 1·50 and 1·95 g MJ^?1 IPAR under adequate N and water in both experiments. This represented an increase in RUE of 52–146% for the range of N treatments used in both experiments, whilst R_acc increased by 9–17%, compared with the control treatments. Subsequently, the total DM yield of kale increased from 6·7 and 8 t DM ha^?1 for the control treatments to ≥ 19 t DM ha^?1 when ≥150 kg N ha^?1 was applied. The DM yields for the 500 kg N ha^?1 treatments were 25·5 and 27·6 t DM ha^?1 for the two experiments. RUE increased linearly with SLN, at an average rate of 0·38 g DM MJ^?1 IPAR per each additional 1 g N m^?2 leaf until a maximum RUE of 1·90 g MJ^?1 IPAR was reached in both experiments. There were no changes in RUE with SLN of > 2·6 g m^?2 and NNI >1, implying luxury N uptake. RUE was the most dominant driver of forage kale DM yield increases in response to SLN and NNI.     

Evaluation of forage legumes for introduction into natural pastures of semi‐arid rangelands of Kenya

To address the potential of legumes to contribute to improved quality and quantity of natural pastures in the semi‐arid rangelands of Kenya, five legume species were introduced and evaluated in a small‐plot field experiment over three growing seasons. The investigated species were glycine (Neonotonia wightii), siratro (Macroptilium atropurpureum), dolichos (Lablab purpureus cv. Rongai), velvet bean (Mucuna pruriens) and shrubby stylo (Stylosanthes scabra cv. seca). Treatments included two cutting heights (ground level and 15 cm) and two cutting intervals (at 2 and 4 months). The mean dry matter (DM) yields of glycine and siratro were highest when the legumes were harvested at ground level at 2‐month intervals (10·31 and 7·81 t ha^?1 year^?1 respectively). Mean DM yield of stylo was highest when the legume was harvested at 15 cm after 4 months (3·52 t ha^?1 year^?1). These three legumes also produced high organic matter through litter fall, which contributed to soil fertility. Evidence from a supporting pot experiment showed effective nodulation and potential for N fixation. These legumes also possessed deep tap roots and withstood heavy defoliation. These three legumes were selected for further integration with grasses in natural pastures. The DM yields of dolichos and velvet bean when harvested at 15 cm after 2 or 4 months were low (2·48 and 1·91 t ha^?1 year^?1), and these species were considered inappropriate for further investigation.     

Phosphorus application and liming effects on forage production,floristic composition and soil chemical properties in the Campos biome,southern Brazil

Studies of responses of native and introduced grassland species to lime and phosphorus (P) applications could contribute to improved understanding of the potential production of South American natural grasslands. To determine the effect of applying lime and different P sources on forage production, diversity and floristic composition, and on soil chemical properties, a small‐plot experiment was conducted over 12 years in natural grassland oversown with Lolium multiflorum and Trifolium vesiculosum in the Campos of southern Brazil. In treatments with soluble phosphate application, dry‐matter (DM) yield in November 2008, after 164 d of winter and early spring growth, increased from 2·3 to 3·2 t ha^?1. Differences in DM yield in March 2009, after 111 d of growth during late spring and early summer, were not significant. The DM yield in April 2010, after 419 d of growth, increased from 7·7 to 9·2 t ha^?1 in the treatments with P, regardless of the P source. Increased forage yield during the slow growth period in winter was only possible with the introduction of winter‐growing species (L. multiflorum and T. vesiculosum) and soluble phosphate application. Assessment of annual forage yield showed that the effectiveness of Gafsa rock phosphate was equivalent to that of soluble phosphates in the long term. Soluble phosphates and liming modified the botanical composition of natural Campos grassland in the long term, but floristic diversity was not altered.