Correction to: Silvicultural prescriptions for mixed-species forest stands. A European review and perspective

European Journal of Forest Research -     

Modeling diameter distributions of mixed-species forest stands

This study compares three diameter distribution models to fit mixed-species forest stands using four example plots with two or three species components in Daxing'an Mountain, PR China. The methods include (1) a finite mixture model (FMM) to fit two or three species components simultaneously, (2) a single Weibull function to fit the whole plot only, and (3) a single Weibull function to fit each species component separately and the summation of the individual species produced the whole plot. Our results indicated that Method 2 is only suitable to regular and unimodal diameter distributions with a balanced reversed J-shape. Method 3 may be able to fit each species component well if its frequency distribution is known and available in the data. However, Method 3 ignores the interspecies relationships within a given plot. Thus, the summation of the species components may not produce a good fit for the whole plot. In contrast, Method 1 (FMM) fits the species component distributions simultaneously with the constraint that the individual components add up to the whole plot, without requiring the observed frequencies for each species across the diameter classes. The FMM models are more flexible to describe highly skewed and irregular diameter distributions for the whole plot, as well as provide the acceptable estimation for each species component and the mixing proportions. Thus, the FMM models can be a useful tool for effectively managing mixed-species forest stands.     

Replacing coniferous monocultures with mixed-species production stands: An assessment of the potential benefits for forest biodiversity in northern Europe

Conifer dominated plantations in central and northern Europe are associated with relatively low ecological values, and in some cases, may be vulnerable to disturbances caused by anthropogenic climate change. This has prompted the consideration of alternative tree species compositions for use in production forestry in this region. Here we evaluate the likely biodiversity costs and benefits of supplanting Norway spruce (Picea abies) monocultures with polycultures of spruce and birch (Betula spp.) in southern Sweden. This polyculture alternative has previously been evaluated in terms of economic, recreational, and silvicultural benefits. By also assessing the ecological implications we fill a gap in our understanding of the range of socio-ecological benefits that can be achieved from a single polyculture alternative. We project likely broad scale changes to species richness and abundance within production stands for five taxonomic groups including ground vegetation, tree-living bryophytes, lichens, saproxylic beetles, and birds. Our research leads us to three key findings. First, the replacement of spruce monocultures with spruce–birch polycultures in the managed forest landscapes of southern Sweden can be expected to result in an increase in biological diversity for most but not all taxa assessed, but it is unlikely to improve conditions for many red-listed forest species. Second, modification of other aspects of forest management (i.e. rotation length, dead wood and green tree retention, thinning regimes) is likely to contribute to further biodiversity gains using spruce–birch polycultures than spruce monocultures. Third, the paucity of empirical research which directly compares the biodiversity of different types of managed production stands, limits the extent to which policy relevant conclusions can be extracted from the scientific literature. We discuss the wider implications of our findings, which indicate that some climate change adaptation strategies, such as risk-spreading, can be readily integrated with the economic, environmental and social goals of multi-use forestry.     

A comparison of annual transpiration and productivity in monoculture and mixed-species Douglas-fir and red alder stands

Although much is known about drivers of productivity in Douglas-fir and red alder stands, less is known about how productivity may relate to stand transpiration and water use efficiency. We took advantage of a 15-year-old experiment involving Douglas-fir (Pseudotsuga menziesii) and red alder (Alnus rubra) in the western Cascade Range of western Oregon to test the following hypotheses: (a) more productive stands transpire more water, (b) the relationship between productivity and transpiration differs between species, and (c) the relationship between productivity and transpiration differs between sites varying in soil moisture and fertility. Furthermore, the experimental design included alder, a facultative nitrogen-fixing species, which could also affect fertility. Fixed area plots (20 × 20 m) were planted as monocultures of each species or in mixtures at a common density (1100 trees ha⁻¹) in a randomized-block design. Transpiration of Douglas-fir and red alder was measured using heat dissipation sensors installed in eight trees per plot and scaled to the plot level based on sapwood basal area for each species. Although up to 53% of the variability in tree transpiration was explained by basal area, irrespective of species or site conditions, the two stands with the highest biomass and sapwood basal area did not transpire the most. Instead of more productive stands transpiring more water, the greatest variability in both productivity and transpiration was determined by site conditions and to a lesser degree, species composition. For example, 70% of the variation in tree biomass increment (TBI) was determined by leaf area index, which was much higher at the site with higher fertility and soil moisture (p < 0.05). Despite marked phenological and physiological differences, Douglas-fir and red alder performed similarly. Only 19% of annual water use of Douglas-fir occurred between October and March when alder was leafless. Also, there was no evidence of a fertilization effect of the nitrogen-fixing red alder on the Douglas-fir: the nitrogen concentration and N-isotopic ratio of Douglas-fir needles did not differ whether trees were grown in monoculture or in mixtures with red alder. We conclude that lower soil fertility and contrasting microclimate at one site relative to the other suppressed NPP while maintaining higher transpiration, thus reducing water use efficiency.     

The nutrient status of Norway spruce in pure and in mixed-species stands

Atmospheric deposition of N and S appears to have caused nutrient imbalance in Norway spruce stands in southern Sweden. This calls for a change of forest management to procedures that promote nutrient balance. Studies have shown lower soil acidity in Norway spruce/deciduous mixed stands than in spruce monocultures, but the tree nutrient status in such mixtures has not been much investigated so far.

The nutrient status of Norway spruce foliage and top mineral soil chemistry in monocultures and in stands mixed with beech, birch, or oak was investigated through paired comparisons on 30 sites in southern Sweden (27 sites) and eastern Denmark (three sites). In total, 45 mixed stands and 34 pure stands were included in the study.

Spruce needles from mixed stands had higher concentrations and ratios to N of K, P, and Zn than needles from pure spruce stands. Among the mixed stands, the K status appeared to be positively correlated with the percentage of deciduous tree basal area. Soil samples from mixed stands had a higher Mg concentration, base saturation, and BC/Al ratio than soil samples from pure stands. The spruce needle nutrient status was comparable in pure stands on fertile sites and in mixed stands on poor sites. We did not detect any differences in spruce tree growth between pure and mixed stands.

This paper discusses possible reasons for a positive effect on the tree nutrient status in mixed-species stands and the possibility of using mixed-species stands as a forest management procedure to avoid nutrient imbalance.     

Determining the optimal selective harvest strategy for mixed-species stands with a transition matrix growth model

An optimization model was developed to determine the optimal harvesting strategy needed for uneven-aged mixed-species stands in the Changbai Mountain region of northeast of China. The model takes into account four variables including residual basal area (RBA), the diameter of the largest tree, harvest cycle and a constant representing the ratio of the number of trees in a given diameter class to those in the next larger diameter class (‘q’). According to model simulations, under the objective of maximizing net revenue, the optimal harvesting strategy is defined when the residual basal area equals to 19 m² ha⁻¹, the diameter of the largest tree equals to 44 cm, q 1.3 and the harvest cycle equals to 20 years. If the objective is to maximize the total volume yield, the optimal harvesting strategy is defined when RBA equals to 13 m² ha⁻¹, the diameter of the largest tree equals to 36 cm and the constant ‘q’ equals to 1.9 and the harvest cycle equals to 15 years.     

The effects of understory vegetation on P availability in Pinus radiata forest stands: A review

In many second-rotation Pinus radiata forest planta-tions, there has been a steady trend towards wider tree spacing and an increased rate of application of P fertiliser. Under these regimes, the potential for understory growth is expected to in-crease through increased light and greater nutrient resources. Therefore, understory vegetation could become a more signifi-cant component of P cycling in P. radiata forests than under closely-spaced stands. Studies have shown that growth rates and survival of trees is reduced in the presence of understory vegeta-tion due to the competition of understory vegetation with trees. Other studies have suggested that understory vegetation might have beneficial effects on nutrient cycling and conservation within forest stands. This review discusses the significance of understory vegetation in radiata pine forest stands, especially their role in enhancing or reducing P availability to forest trees.     

Point process models for mixed sessile forest stands

• Background   

Growth modelling of complex stands calls for the use of spatially explicit single-tree models. Such models require spatially explicit tree locations as the initial state to run simulations. Given the cost of such data, virtual forest stands, where tree locations are simulated, are generally used as the initial state.     

Silvicultural strategies for increased timber harvesting in a Central European mountain landscape

The demand for wood as construction material, renewable source for energy and feedstock for chemicals is expected to increase. However, timber increments are currently only partly harvested in many European mountain regions, which may lead to supply shortages for local timber industries, decreases in forest resistance to disturbances and functioning as protection from gravitational hazards. Using an inventory-based forest simulator, we evaluated scenarios to increase wood mobilization in the 7105-km² Swiss canton of Grisons for the period 2007–2106. Scenarios varied with respect to landscape-scale harvesting amounts and silvicultural strategies (low vs. high stand-scale treatment intensity) and accounted for regulations and incentives for protection forest management. With 50 and 100% increases of harvests, the current average growing stock of 319 m³ ha^?1 was simulated to be reduced by 12 and 33%, respectively, until 2106 in protection forests of Northern Grisons, where management is prioritized due to subsidies. Outside protection forests and in Southern Grisons, growing stock was simulated to continually increase, which led to divergent developments in forest structure in- and outside protection forests and in the Northern and Southern Grisons. The effect of silvicultural strategies on simulated forest structure was small compared to the effect of future harvesting levels. We discuss opportunities and threats of decreasing management activities outside protection forests and advocate for incentives to promote natural regeneration also outside protection forests to safeguard long-term forest stability.     

Growth and yield models for uneven-sized forest stands in Finland

Uneven-sized forestry is gradually gaining popularity and acceptability also in the Nordic countries. This is because of the willingness of the public and some forest owners to avoid clear-fellings and pursue more near-nature forest structures. It has also been realized that the profitability of uneven-sized forestry may be competitive with even-aged forestry. In Finland, management of uneven-sized stands is hampered by the lack of information about the dynamics of such stands, and about the yield and profitability of uneven-sized forestry. This study developed models which allow managers to simulate the growth and yield of uneven-sized stands in Finland, making it possible to predict the yield and analyze the sustainability of different management options. The model set consists of individual-tree diameter increment, height and survival models, and a model for ingrowth. The modeling data consisted of two long-term field experiments of uneven-sized forest management, a set of temporary sample plots measured earlier for growth modeling purposes, and the sample plots of the third National Forest Inventory of Finland. The application area of the models covers all growing sites, all main tree species, and the whole surface area of Finland. According to the models, the sustainable harvest of a fertile (Oxalis-Myrtillus site) uneven-sized Norway spruce forest varies between 5.5 and 7 m³ ha⁻¹ a⁻¹ in Central Finland, depending on the length of the cutting cycle, stand density, and shape of the diameter distribution. It is profitable to harvest large diameter classes more heavily than small ones. Due to the large amount of data the models for diameter increment are highly significant and reliably show the growth level of trees in uneven-sized stands. The weakest models are the ingrowth models, which are based on a clearly smaller data set than the other models.     

国家网络森林医院中建立林木健康卡构想

为了适应林改形势,强化公共服务,完善服务功能,建议在国家网络森林医院中,针对具有林权证的用户建立林木健康卡,记录林地的基本信息、林木防治活动以及用户在网院的咨询与专家的解答等信息,实现网院"服务社会、服务林改、服务现代林业"的宗旨。本文系统地探讨了建立林木健康卡的可行性、设计原则、实现方法与使用方法。     

A flexible modeling of irregular diameter structure for the volume estimation of forest stands

Simple distribution models (SDMs) have deficiencies in portraying irregular forest diameter structure. This paper introduces mixture distribution models (MDMs) to improve the estimation accuracy of stand volume of Siberian larch (Larix sibirica Ledeb.) forests. Stand volume was estimated by combining the suitable diameter mixture model and diameter–height model. Appropriate mixture models are derived by integrating multiple SDMs of Burr III and XII, Johnson S_B, Weibull or lognormal probability density functions (pdf) that satisfied the criteria of goodness of fit tests. Results showed that the average bias of volume estimation for all of the study plots using SDM and MDM approaches are underestimated by 6.93 and 2.42 m³, respectively. Each of the estimates is equivalent to an estimation error of 25.59 ± 18.18 and 7.08 ± 2.97 %. This suggests that the MDM approach is a more flexible and suitable modeling technique for forest volume estimation, in particular for forests that have been frequently disturbed by natural events.     

Yield—Density diagram for natural deciduous broadleaved forest stands

A yield—density diagram for natural deciduous broad-leaved forests in Hokkaido was devised. It consists of Y—N curves, equivalent diameter curves, equivalent mean diameter curves, a B-point curve and a full density curve. The Y—N curve is derived from the relation between cumulative summation (Y) of individual tree volume from the maximum sized tree to one of a certain given volume, and number (N) of these trees, and is expressed by the reciprocal equation of two parameters, A and B. The equivalent diameter curve expresses the number of trees whose diameters are larger than a certain size. The equivalent mean diameter curves express the number and volume of trees of selected mean diameters. The B-point curve is a hypothetical trajectory of the B-point of the C—D curved ruler (C—D rule) which is fitted to each Y—N relation, throughout the course of stand development. The full density curve indicates the upper limit of the number and volume of the standing trees. Utilizing this diagram, we can comprehend numbers, volumes and tree diameters in a stand and among stands.A method is devised of predicting the number and volume of each diameter class of a stand several years after thinning. Thinned trees are assumed to be selected equally from each diameter class so that the Y—N curves are fitted equally to the Y and N data before and after thinning. Volume growth of individual trees is assumed to be proportional to their volumes after thinning. The current volume growth of the stand after crown closure is assumed to be constant irrespective of the stand age. Tree mortality after thinning is assumed to be negligible. With these assumptions, a trajectory of the Y—N curve through thinning is determined, and a yield after n years from thinning is predicted. Comparing the Y—N curve of a thinned stand with that of an unthinned stand after n years from thinning, we can comprehend the thinning effect.     

提高林分单位面积蓄积量的对策

林分单位面积蓄积量是林地重要的生态和价值指标,分析了遂昌县影响单位面积蓄积量是营林生产投入不足、造林作业方式的改变、择伐的普遍性、香茹原料的生产等原因,据此提出了拓宽营林生产投资渠道、推行工程造林质量管理、因地制宜炼山造林、联户经营限制择伐、加强林区索道管理等措施。     

Clustering forest harvest stands on spatial networks for optimised harvest scheduling

Context

Clustering forest harvest stands reduces the movements of the harvesters, forwarders and staff. Moreover, it simplifies the subsequent log transport, when compared with dispersed stands.

Aims

Harvesting activities are generally based on silvicultural motivated planning data. The development of an analytical method to cluster harvest stands with respect to the spatial network of roads should improve the harvesting effort.

Material and method

The clustering of harvest stands was developed for Aracruz (Brazil) in 2004, where it is used there successfully since. The hierarchical method ??single linkage cluster analysis?? is applied. As a distance function, the Euclidian distance was substituted by the shortest path on the spatial network.

Result

The clustering method is based on the minimum spanning tree, which is the spatial equivalent to the dendrogram of an ordinary cluster analysis. Applying the Delaunay triangulation to fill the distance matrix reduces the distance calculation time from O(n ²) to O(n). The method is illustrated by a planning district of the Aracruz enterprise.

Conclusion

Harvesting units are properly clustered spatially by the discussed method. Topographic obstacles are automatically avoided and the need to relocate machinery is reduced as well as the total driving distance.     

紫金山风景林林相改造异龄复层混交模式研究

林相改造异龄复层混交林模式研究表明,在郁闭度0.5~0.75的疏林中"林下植阔、林中插绿"能形成较稳定的地带性异龄复层混交林群落,提高风景林生态系统的质量和生态功能。