The influence of heterogeneous habitats on the germination ofAcanthopanax senticosus seeds and the dynamics of its seed pool

Differences in germination ofA. Senticosus, an important shrub under the Korean Pine broad-leavd forest and its secondary forest in heterogeneous habitats and the dynamics of its seed pool have been studied by observations in location. The results showed that transformation rates from seeds to seedlings were in the following order: 16.8% in Larch plantations, 4.1% in Mongolian oak forests, 2.7% in Birch forests, 1.8% in Korean Pine plantations, and 0.5% in hard wood forests. The life-span of seeds was determined to be four years in the simulating seed pool under Larch plantations. The percentage of the seedling output was, 14.5% in the 2nd year, 10.1% in the 3rd year, and 1.8% in the 4th year. Other types of the seed output were those: decayed 33.1%, senescent, 22.3%, predated by mice 14.1%, which was variable due to the variation in the number of mouse, and only 1.17% were eaten by insects and ofher soil animals. The project was supported by National Science Foud of China     

Evaluation of regeneration potential of Pinus koraiensis in mixed pine-hardwood forests in the Xiao Xing’an Mountains, China

Large scale harvest of Korean pine(Pinus koraiensis) seeds as a food product in the mixed Korean pine-hardwood forest of northeastern China poses a serious threat to the sustainability and restoration of this endangered regional ecosystem.Seed collection over past decades greatly reduced the seed bank and subsequent seedling and sapling recruitment,and impacting a wide array of granivorous animals that rely on the pine seeds.We surveyed Korean pine seeds,including solid seeds(SS),insect consumed seeds(ICS) and other(animal) consumed(OCS) kernels,of the seed bank(forest floor and the top 10 cm of mineral soil),the seedlings and saplings from 1 m 2 sample plots in five forest types in Liangshui Nature Reserve(LNR) of the southern Xiao Xing’an Mountains in northeastern China to provide accurate information for assessing the Korean pine regeneration potential.The average number of pine seeds in the seed bank were 11.2 seeds/m 2,9.1 seeds/m 2,4.6 seeds/m 2,1.1 seeds/m 2,and 0.2 seeds/m 2 in Korean pine-basswood forest,mixed Korean pine-hardwood forest,mixed conifer-hardwood forest,white birch forests,and oak forests,respectively.In the first three forest types,percentages of SS(potentially viable seeds) were 11.2%,3.5% and 27.8%,respectively.The percentages of ICS(not viable seeds) were consistent at around 35%.The higher but variable percentages of OCS(not viable seeds) indicated high seed predation in these forests.Compared with other studies,we recorded higher percentages of seed damage,probably due to our survey approach and the increased depth of seed bank sampled in our study.Depletion of pine seeds in the seed bank greatly reduced seedling and sapling recruitment.Densities of pine seedlings varied from about 180 trees/ha in the mixed Korean pine-hardwood forest to about 5,400 trees/ha in the mixed conifer-hardwood forests and showed a high degree of spatial variation.Saplings were rare in the mixed Korean pine-hardwood forest,but ranged in the thousands in other forests.Large scale pine seed harvest has seriously threatened the sustainability of the mixed Korean pine-hardwood forest ecosystem.Scaling down the seed harvest or supplemental planting of pine saplings are urgently needed to maintain the health of the existing Korean pine forests and to restore this endangered ecosystem.     

凉风坳亚热带次生常绿阔叶林土壤种子库和种子雨的特征研究

选择四川省沐川县凉风坳亚热带次生常绿阔叶林为研究对象,在2004年5月、2004年12月和2005年5月对其林下土壤种子库、种子雨以及种子天然萌发状况等方面进行系统调查及分析研究。结果显示,种子下落前后土壤种子库储量,变动为643.2～889.2粒.m-2,物种和种子密度多年生草本物种均占优势。天然林土壤种子库Shannon-Wiener指数均在2004年5月最大,2004年12月次之,2005年5月最小;Simpson指数却和Shannon-Wiener指数H＇的变化趋势相反;Pielou均匀度指数在2004年12月有所降低,此后又升高。种子均呈明显垂直分布,其中0～2 cm土层种子数量最多,其次是枯枝落叶层,其数量随着土层加深而降低。土壤种子库年输入活力种子142.3粒.m-2,年输出活力种子268.9粒.m-2,动物取食是造成种子损失的主要原因。对照人工林土壤种子库物种和密度均低于天然林地;种子库与地上植被物种相似性系数,人工林较高,而天然次生常绿阔叶林较低。     

Scots pine (Pinus sylvestris L.) plantations contribute to the regeneration of holly (Ilex aquifolium L.) in mediterranean central Spain

The aim of this study is to evaluate the recruitment possibilities of holly (Ilex aquifolium L.) populations under Pinus sylvestris plantations in the mountains of Central Spain. Population recruitment of holly under P. sylvestris L. plantations was studied at two localities. We sampled holly seed rain, seed bank, and seedling emergence and survival (of both emerged and planted seedlings) in both localities. Seed germination was analysed by randomly sowing Ilex seeds and their subsequent recollection and analysis over the following 3 years (only in one locality). Seed rain, seed bank and emerged seedling densities did not differ between localities. Seed bank density was between 10 and 20 seeds m⁻², considerably higher than the observed seed rain (0.4–1.2 seeds m⁻²), and also higher than the emerged seedling density (0–4 seedlings m⁻²). But Ilex is not very efficient in germinating under pine forests (29.1% germination after 3 years). This may indicate a seed accumulation process of at least 5 years. Seedling survival for the first year differed among the studied sites (50–60 vs. 17%). Nevertheless these survival rates are higher than those found for the same seedling populations in different forest habitats (not Scots pine forests) from near locations. Differences in summer water stress between the study sites are proposed as the main cause of the observed differences in seedling survival. It is concluded that P. sylvestris forests may provide one of the best regeneration habitats for holly in the Mediterranean area, where this species is close to the Southern limit of its distribution.     

诱导改造对大兴安岭低质林土壤理化性质的影响

2009年对大兴安岭地区的阔叶混交次生林及白桦萌生低质林进行不同带宽皆伐,将每条皆伐带分为3段,分别种植西伯利亚红松、落叶松和樟子松,2011年8月测定林分的土壤理化指标。结果表明:不同方式改造后土壤密度均不同程度地降低,而土壤毛管持水量、毛管孔隙度均不同程度地升高;与对照样地相比,阔叶混交次生林不同诱导改造后土壤全氮含量均升高(落叶松林升高98.26%,西伯利亚红松林升高72.70%,樟子松林升高44.91%),除落叶松林土壤全磷含量3.35g·kg-1保持不变外,西伯利亚红松林和樟子松林全磷含量均降低;与对照样地相比,白桦萌生低质林不同诱导改造后土壤全磷、全氮含量均有所降低,全磷含量降低程度表现为西伯利亚红松林降低38.90%、落叶松林降低33.15%、樟子松林降低11.78%,全氮含量降低程度表现为西伯利亚红松林降低36.02%、落叶松林降低36.75%、樟子松林降低37.68%;综合分析所有改造林,土壤有机质含量与全氮、全钾含量显著正相关(P﹤0.05);不同带宽西伯利亚红松林的土壤pH值、有机质含量、全氮含量及全钾含量变异系数最大,说明带宽对西伯利亚红松诱导林土壤理化性质有显著影响。     

油松、辽东栎混交林中生化他感作用的研究

研究了油松（ＰｉｎｕｓｔａｂｕｌａｅｆｏｒｍｉｓＣａｒｒ．）、辽东栎（ＱｕｅｒｃｕｓｌｉａｏｔｕｎｇｅｎｓｉｓＫｏｉｄｚ．）混交林中的生化他感作用，结果表明：油松、辽东栎纯林及松栎混交林下的枯落叶、半分解枯落叶及表层土壤水浸液对油松及其它植物的种子发芽和幼苗胚根、幼茎生长有显著的影响，一般在高浓度下表现为强烈的抑制作用，低浓度下作用不显著或起促进作用。其中辽东栎枯落叶水浸液的抑制作用最为强烈，混交林中枯落叶的抑制作用有所下降。说明在某些条件（干旱、混交比例和混交方式不当）下，两树种混交可引起林地生化他感物质富积，从而导致油松生长不良，林地生产力下降。解释了实际林地的现象，为营造松栎混交林提供了依据。     

Seed Biology,Stand Characteristics,and Regeneration of Fraxinus rhynchophylla Hance

Fruit size and seed moisture content were measured and seed production and dispersal were monitored to understand the seed biology of the Korean ash (Fraxinus rhynchophylla Hance) in two Korean ash forests. A combination of four ground treatments and four crown closure levels were created in a natural forest to understand the site conditions needed for seedling emergence in natural stands and to determine practical regeneration methods for direct seeding. The seed size of Korean ash increased until late May and its moisture content decreased rapidly in early November. Prolific seed bearing occurred every 3 yr. The distance of seed dispersal by wind was about 30 m, but more than 90% of the seeds dispersed within 10 m from the seed trees. Twenty-five seed trees per hectare provided for successful natural regeneration. The seedling emergence in the natural stands was best with scarification treatment and 25～50% of crown closure.     

关于发展人工林与建立人工林业问题探讨

全世界森林面积为 34 5 4× 1 0 8hm2 ,其中 3%是人工林。人工林是重要的森林生态系统 ,在木材生产、环境保护和景观建设中具有重要功能和作用。人工林在生产功能方面的补偿作用 ,使天然林可以更好地发挥生态环境价值、森林娱乐和生物多样性保护功能。人工林可持续经营的前提是遗传材料的正确选择与营林实践艺术的结合以及保证这种结合的林业政策。愈来愈多地使用外来树种营建人工林是全球趋势。人工林对于环境和生物多样性的影响 ,可以通过合理的规划和经营措施得到解决。林木育种与遗传改良和经营制度的技术创新是不断提高人工林生态系统生产力的动力。人工林的可持续经营是一个十分复杂的问题 ,中国人工林实现可持续经营面临技术和体制两方面的挑战 ,但是 ,根本出路在于林业经济体制的改革 ,象农业那样 ,在市场经济环境中实行人工林产业化经营 ,建立人工林业 (Forestplantationindustry)。     

二滩库区锥连栎林土壤种子库和幼苗格局初步研究

本文对二滩库区群落优势树种锥连栎林土壤种子库和幼苗分布格局进行了调查研究，结果表明：(1)锥连栎群落土壤种子库不存在霉烂种子，虫蛀种子的数目较少，仅为2.08个／m^2；而败育种子和种子壳数目相对较多，平均密度分别达到42．78个／m^2和42.96个／m^2。(2)锥连栎土壤种子库组成中完好种子的密度为0．28个／m^2；雨季已萌发的种子密度为0．92个／m^2。(3)土壤种子库各组成的垂直分布较为明显，随土层厚度加深而减少，但霉烂种子例外。(4)经测算，锥连栎种子散布后的主要命运是动物捕食和转移，种子的保存率和萌发率分别仅占到单个树木种子产量的0、59％和1．95％。(5)在锥连栎树木分布的上坡有较多的萌生苗，而下坡实生苗的数量较多。(6)距树干距离不同，萌生苗和实生苗的分布模式有差异。     

西安市近自然林经营模式探讨

介绍了近自然林业经营理论及其在我国的研究与实践,分析了西安市林业建设中应用近自然森林经营理论的必要性。对西安市周边风景片林近自然林营造模式,秦岭北麓浅山区灌木林地近自然化改造模式和秦岭北麓油松、落叶松高密度人工纯林的近自然化改造模式进行了探讨。       

Tree Recruitment from On-site Versus Off-site Propagule Sources During Tropical Forest Succession

The management of successional forests has become an increasingly important tool for tropical forest restoration on disturbed lands. Successful management depends on understanding factors governing tree recruitment during succession. Recruits can be on-site propagules surviving disturbance (e.g. saplings) or seeds arriving after disturbance. Among disturbed sites, the importance of seed arrival from off-site should increase as availability of on-site propagules declines. This study examines how seed arrival from off-site seemed to influence natural forest regrowth on logged plantations in Uganda. Specifically, the effect of seed arrival from mammals (determined directly) and birds (determined inferentially) was estimated in two plantations with high and three plantations with low quantities of saplings surviving logging. In both plantation types, first-year recruitment seemed to originate from on-site (seed bank and sprouts from damaged trees) and off-site sources (seed-dispersing birds). Recruitment declined 2–3 years after logging in all plantations despite high densities of seed-dispersing birds. Recruitment was generally greater in plantations with low initial quantities of on-site propagules, and by 4–6 years forest structure was similar between plantation types. Recruitment was dominated by bird-dispersed species, rather than mammal- or wind-dispersed species. Taken together, these results illustrate that disturbed areas with low initial on-site propagule availability have much potential for forest regrowth.     

Long-lasting effects of unplanned logging on the seed rain of mixed conifer-hardwood forests in southern South America

Understanding the effects of disturbance on seed rain is critical to predict changes in forest species composition and diversity.Logging effects on seed rain in a mixed conifer-hardwood forest complex in southern Brazil were evaluated.One year of seed rain data were collected from a large-scale observational experiment in logged and protected forests and quantity and average seed size weighted by species abundance(CWM) were compared between old-growth and logged stands 55 years after logging activities.Using these data,variations in frequency of functional groups of species in the seed rain were examined to see if they could be attributed to logging.Results show that the number of seeds per trap was highly right-skewed,ranging from 13 to 12,788 seeds per trap in one year.Seed rain was affected by logging history,with seed traps in old-growth plots receiving significantly less seeds than traps in logged plots.All species included mean seed size weighted by species abundance were significantly smaller in logged than in old-growth forests.This difference persisted after the exclusion of Arawcaria a large-seeded pioneer which was intensively logged,although the difference of seed size between the two forest classes was greatly reduced.Species abundance in the seed rain differed significantly from the established tree community,between logged and oldgrowth stands.The composition of the seed rain was much more variable than the composition of the established tree community and its points more scattered over the ordination space than the points corresponding to the protected forests.The number of collected seeds across different functional groups of species significantly differed between logged and old-growth plots.The seed rain of logged forests reflects their arrested succes sion as indicated by reduced abundance of functional groups such as pioneers,large seeded pioneers and A raucaria,as well as reduced functional diversity.S eed rain differences between logged and old-growth stands reflect the intensive logging of A.angustifolia.     

Seed dispersal,dormancy, seedbank and regeneration of Amur corktree

Amur corktree (Phellodendron amurense Rupr.) is one of the important accompanying species of Korean pine (Pinus Koraiensis Sieb et Zuce) broadleaved forest in northeast China. This paper reports on the seed dispersal, dormancy, seed bank and regeneration of amur corktree. Seeds of amur corktree arc dispersed mainly by the aid of birds and gravity. The aril of the seeds may increase seed longevity. In natural secondary forests, the store of the seed bank may reach 750, 000 seeds per heetare. Regeneration was correlated wirh disturbance, and was best under intensive disturbance.     

Direct seeding of late-successional trees to restore tropical montane forest

Natural regeneration of large-seeded, late-successional trees in fragmented tropical landscapes can be strongly limited by a lack of seed dispersal resulting in the need for more intensive restoration approaches, such as enrichment planting, to include these species in future forests. Direct seeding may be an alternative low-cost approach to planting nursery-raised tree seedlings, but there is minimal information on its efficacy or when in the successional process this technique will be most successful. We tested directly seeding five native tree species into habitats representing passive and active restoration approaches: (1) recently abandoned pasture; (2) naturally establishing, young secondary forests; and (3) young, mixed-species (fast-growing N-fixers and commercially valuable species) tree plantations established to facilitate montane forest recovery in southern Costa Rica. We monitored germination, survival, growth, and above- and below-ground biomass over a 2-year period. Germination in pastures, secondary forests, and tree plantations was similar (∼43%). Seedling survival after one and two years was significantly higher under tree plantations (91% year 1, 75% year 2) compared to secondary forests (76, 44%) or pastures (74, 41%). Moreover, seedlings had greater total biomass and lower root:shoot ratios in the plantations, suggesting higher nutrient availability in that treatment. Costs for direct seeding were 10- to 30-fold less per 100 seedlings after 2-year compared to nursery-raised seedlings planted at the same sites; however, there are important trade-offs to the two restoration approaches. Planting nursery-raised seedling is a more effective but higher cost approach for rapidly establishing canopy cover and restoring large areas whereas direct seeding is a more efficient way to enrich an existing system. We particularly recommend using direct seeding as a complimentary measure to the more intensive restoration approach of planting fast-growing and N-fixing trees.     

呼伦贝尔沙地樟子松天然林自然落种特征研究

研究了呼伦贝尔沙地樟子松天然林自然落种及其种子萌发特征,结果表明:丰年期落种器共接到99粒种子,且集中分布在母树的正东、东北侧,欠年期则接到落种42粒且集中分布在正东、东南侧。丰年期不同方位、不同落种距离接到的种子平均发芽率为54.5%,欠年期为28.6%。樟子松林缘落种的传播距离范围可以达到25 m,并且在林缘外5 m范围内种子数量最多,林缘外5～20 m范围平均发芽率为60.9%。樟子松单株树木下土壤种子库中种子数量在4 m处最高,并且随着距树干距离的增加种子数量也随之降低,种子总数由2 m处样方的16粒降低到10 m处样方的7粒,降低百分率达129%。     

Effects of seed quality and seed location on the removal of acorns and beechnuts

We examine whether different guilds of foragers remove seeds differentially according to seed quality (seed size and insect infestation) and seed location (habitat and microhabitat) in a mixed oak-beech forest. Video recordings indicated that the wood mouse (Apodemus sylvaticus) was first to encounter seeds. Foragers preferred acorns to beechnuts, large to small size and sound to infested. Nevertheless, infested seeds were removed by rodents even when sound seeds were present. Seeds that were not preferred by scatter-hoarding rodents remained longer on the ground and were more vulnerable to predation and desiccation (4% moisture loss per day). However, seeds that were removed by scatter-hoarders were moved away from their mother trees (96%) and cached individually (32%), increasing their moisture content (3% per day). Buried seeds, simulating scatter-hoarding behavior, experienced only a 17% removal after 4 months. Seed removal differences among habitats were not due to habitat attributes but to the spatial distribution of rodent-preferred microhabitats. Thus, a significant lower seed removal was observed under the tree canopy with no shrubs. However, seed removal in forest gaps with deadwood cover was not significantly different from the preferred microhabitat (under shrub cover). In pure beech forests, seed removal by rodents only occurred under Ilex aquifolium (the only perennial cover) and under woody debris. This study concludes that seed quality and seed location determine the contribution of different removers (predators vs. dispersers), their seed selection and their removal speed, leading to different seed fates which will eventually affect tree regeneration.     

三峡地区柏木林种子雨和种子库的研究

柏木种子雨在8月底开始降落,10月达到顶峰,翌年5月结束,历经9个月左右。整个过程呈单峰曲线,10~11月的种子雨量占全年的67.5%。种子雨时间变化格局差异极显著,而水平空间分布格局没有显著差异。种子雨中柏木种子的年均密度为5 982.2粒.m-2。63.4%的柏木种子直径大于2.5 mm,构成了柏木种子雨的主体。由于这一级别柏木种子的萌发能力较强,因此三峡地区柏木林的更新能力强。柏木林中非柏木种子的种子雨每年主要发生在1月和5月,形成2个高峰,年均种子雨密度为1 842.8粒.m-2。土壤种子库中柏木种子的平均密度为608.8粒.m-2。柏木种子在土壤种子库中垂直分布格局差异极显著。0~2 cm土壤层所分布的柏木种子占种子库中柏木种子的80.49%。其它土壤层次所含有的种子比例很低,且萌发能力较低,因此,柏木林的更新主要依赖于表层土壤种子库的种子。柏木种子在种子库中的水平分布格局虽然随坡位的不同而有所变化,但是差异不显著,表明整个柏木林的更新能力在水平格局上无显著差异。     

英国西加云杉和落叶松的无性繁殖

本文概述了英国西加云杉和落叶松无性繁殖的历史、研究背景、生产方法、扦插苗的造林表现和大规模应用的发展前景。目前已优化无性繁殖系统的扦插苗生产成本为常规的１／３，且扦插苗不增加造林成本。     

Tropical reptiles in pine forests: Assemblage responses to plantations and plantation management by burning

Worldwide, the land area devoted to timber plantations is expanding rapidly, especially in the tropics, where reptile diversity is high. The impacts of plantation forestry and its management on native species are poorly known, but are important, because plantation management goals often include protecting biodiversity. We examined the impact of pine (Pinus caribaea) plantations, and their management by fire, on the abundance and richness of reptiles, a significant proportion of the native biodiversity in tropical northern Australia, by (i) comparing abundance and diversity of reptiles among pine plantations (on land cleared specifically for plantation establishment), and two adjacent native forest types, eucalypt and Melaleuca woodlands, and (ii) comparing reptile abundance and richness in pine forest burnt one year prior to the study to remove understorey vegetation with pine forest burnt two years prior to the study. We also examined the influence of fire on reptile assemblages in native vegetation, by comparing eucalypt woodland burnt two years prior to the study and unburnt for eight years. To quantify mechanisms driving differences in reptile richness and abundance among forest types and management regimes, we measured forest structure, the temperatures used by reptiles (operative temperature) and solar radiation, at replicate sites in all forest types and management regimes. Compared to native forests, pine forests had taller trees, lower shrub cover in the understorey, more and deeper exotic litter (other than pine), and were cooler and shadier. Reptile assemblages in pine forests were as rich as those in native forests, but pine assemblages were composed mainly of species that typically use closed-canopy rainforest and prefer cooler, shadier habitats. Burning did not appear to influence the assemblage structure of reptiles in native forest, but burning under pine was associated with increased skink abundance and species richness. Burned pine was not warmer or sunnier than unburned pine, a common driver of reptile abundance, so the shift in lizard use after burning may have been driven by structural differences in understorey vegetation, especially amounts of non-native litter, which were reduced by burning. Thus, burning for management under pine increased the abundance and richness of lizard assemblages using pine. Pine plantations do not support the snake diversity common to sclerophyllous native forests, but pine may have the potential to complement rainforest lizard diversity if appropriately managed.     

水源涵养林结构与功能耦合关系模型研究——以北京山区水源涵养林为例

采用野外调查和数学模拟方法,选择树种组成、年龄、郁闭度、起源、层次、生物量以及土壤厚度等7个林分结构因子,建立了北京山区水源林结构与功能耦合关系模型。利用结构与功能耦合关系模型,比较了北京密云水库上游8种水源林森林类型,结果表明:1)保持水土功能大小顺序是阔叶混交林>桦树林>蒙古栎林>山杨林>油松林>刺槐林>侧柏林>落叶松林;2)涵养水源功能大小顺序是山杨林>蒙古栎林>桦树林>阔叶混交林>落叶松林>侧柏林>油松林>刺槐林;3)改善水质功能大小顺序是桦树林>蒙古栎林>山杨林>阔叶混交林>侧柏林>刺槐林>油松林>落叶松林。