Frequency distributions of sward height under sheep grazing

Cattle and sheep can create and maintain a mixture of relatively tall and short patches in grass swards through selective grazing. In swards that are grazed by cattle this heterogeneous structure can result in the frequencies of height measurements having a skewed distribution that has variously been better described by the double‐normal distribution the gamma distribution and the Weibull distribution than by the more common normal distribution. The fit of these statistical distributions, and the adequacy of the potentially useful log‐normal distribution, to sward height frequencies were tested in sown temperate swards grazed by sheep and compared within a single sward. It was concluded that the single‐normal and Weibull distributions were inadequate and that overall the log‐normal and gamma distributions had the best fit to the measurements.     

Application of double normal frequency distributions fitted to measurements of sward height

Herbage heights were measured with a rising plate meter in mixed perennial ryegrass/white clover ( Lolium perenne/Trifolium repens ) swards maintained under five different systems of management. Double normal distributions fitted to the height frequencies were used to interpret changes over the grazing season in the mean heights of the shorter 'frequently' and the taller 'infrequently' grazed components of the sward, the variability of height within the two components, and the proportions of the two components in the sward. An attempt was also made to demonstrate that the overlap of these components, implicit in fitting double normal distributions, is a true reflection of the height distributions of frequently and infrequently grazed areas of the sward, and that the fitting of such a distribution provides a useful estimate of the proportion of a sward which is infrequently grazed.     

Defoliation and productivity of a phalaris-subterranean clover sward, and the influence of grazing experience on sheep intake

Swards of Phalaris aquatica-Trifolium subterraneum were subjected to four defoliation treatments—zero, low (11 sheep ha⁻¹) and high (22 sheep ha⁻¹) stocking rates, and weekly cutting. At high stocking rate the annual grass Hordeum leporinum dominated while clover was dominant at low and zero stocking rates. Weekly cutting suppressed species other than clover and so failed to simulate grazing.
There were similarities in net herbage production between zero and lightly grazed swards and between heavily grazed and repeatedly cut swards. Net herbage production decreased in the order undisturbed sward < lightly grazed sward < heavily grazed sward < repeatedly cut sward.
When sheep grazed swards where herbage mass was low their daily consumption of herbage, and therefore liveweight change, depended on their recent grazing experience. Sheep accustomed to swards where herbage mass was low ate more because they grazed for much longer each day than unaccustomed sheep, although they selected a diet of similar digestibility.     

Persistence and colonization of gaps in sown swards of grass and clover under different sward managements

The effects of extensive sward management and patch size on the persistence and colonization of gaps in sown swards was examined by creating gaps of five different sizes (2·3, 7, 10, 14 and 19 cm in diameter) in four different sward treatments: a fertilized sward grazed to 4 cm, i.e. relatively intensive management, and three extensively managed unfertilized swards, which were not grazed or grazed to 4 cm or 8 cm. The swards were originally sown with ryegrass (Lolium perenne L.) and white clover (Trifolium repens L.), but had developed differences in species composition as a result of the management treatments imposed 2 years before and during the experiment. Light quality measurements, i.e. red-far red (R/FR) ratio, were used to determine when the light environment in the gaps no longer differed from that in control, uncut patches and this was used as an estimate of gap persistence. Persistence of gaps depended on both sward management and gap size. Gaps disappeared most rapidly in the ungrazed sward and fertilized 4-cm sward, and most slowly in the unfertilized 8-cm sward. Small gaps persisted for up to 2 weeks in all but 8-cm swards, whereas larger gaps were estimated to persist for up to 20–25 weeks in unfertilized, grazed swards. There was no evidence that the number of grass or dicotyledonous species increased in the gaps compared with the control areas. There were significant positive linear relationships between the vegetation that developed in gaps and that in the control, uncut patches, reflecting the different species composition of the established sward of the grazed (grass-dominant) and ungrazed (Ranunculus repens-dominant) treatments. For total grass dry matter and tiller numbers, as well as L. perenne tiller numbers, there was a small, but significant, effect of both patch size and sward management on the slopes of the regressions between the controls and gaps. The results are discussed in relation to the potential for species composition of sown swards to change as a result of gap creation.     

THE EFFECT OF FERTILIZER NITROGEN AND WHITE CLOVER ON HERBAGE PRODUCTION

The effect of a range of N rates on herbage production from grass/white-clover and grass swards was investigated in two long-term grazed experiments. The mixed sward yielded more than the grass swards over the range of fertilizer rates tested which were 0–120 lb N/ac (134.5 kg/ha), but the response of the mixed sward to fertilizer N was less than that of tbe grass sward. Tbis was due to the direct and indirect effect of clover, which was considerable with no fertilizer N but decreased witb increasing rates of application. However, clover was not completely suppressed by N at eitber site and contributed to berhage yields at all N rates. Rate and time of N application and the amount of clover in tbe sward affected tbe seasonal distribution of DM yields. The mid-season decline was less marked at bigb- tban at intermediate-levels of N supply.     

Intake and behaviour responses by sheep to changes in sward characteristics under rotational grazing

A 10-week grazing experiment was conducted on a perennial ryegrass sward with lactating ewes and their twin lambs. Three paddocks were rotationally grazed with rest periods of from 4 to 5 weeks. Sward surface heights at the start of each grazing were 145, 259 and 250mni for treatments RG1, RG2 and RG3. A further four paddocks were maintained by continuous variable stocking (CS) at sward surface heights (SSHs) of about 30, 60, 90 and 120mm. Sward and animal measurements were made on the two different grazing managements as the RG swards were grazed down, giving measurements at similar sward heights for treatments RG and CS.
There was less green leaf and the total herbage mass present under RG was less than on CS swards at the same sward heights, demonstrating the differences in structure between rotationally and continuously grazed swards.
Regression analysis of animal factors on sward factors showed that grazing behaviour was more highly correlated with green leaf mass than SSH or any of the other sward measurements. On the RG swards, maximum intake per animal was reached at about 1500 kg green leaf mass ha⁻¹. A SSH of 60mm allowed the CS ewes to achieve the highest intake rate, but at this height the ewes on treatments RG2 and RG3 were restricted to approximately half this rate. The results suggest that green leaf mass or leaf area index, rather than sward surface height, could be used as a rational basis to relate intake of herbage to sward state for swards changing rapidly in leaf to stem ratio.     

Sward growth under cutting and continuous stocking managements: sward canopy structure, tiller density and leaf turnover

The change in structure of continuously grazed versus infrequently cut swards of perennial ryegrass ( Lolium perenne L), cv. S23, was investigated during their first full harvest year. Measurements were made from early May until late September. The intensity of stocking by sheep in the grazed sward was adjusted in an attempt to maintain a high level of radiation interception and the cut sward was harvested at approximately monthly intervals.
The herbage mass, lamina area index and radiation interception of the cut sward varied in a cyclic pattern between harvests but in the grazed sward these parameters showed considerably less variation, although they all increased early in the season and then declined later. The proportion of dead material above ground increased throughout the season in both sward types but was more marked in the grazed sward.
There were major differences between the grazed and cut swards in the number of tillers per unit ground area; the difference became more marked throughout the season and by September the tiller densities in the grazed and cut swards were 3·20⁴ m^-2 and 6·20³ m^-2 respectively. Divergence in tiller density was associated with differences in specific stem weight and leaf area per tiller.
Rates of appearance and death of leaves on tillers in the grazed sward were determined. During May, leaf appearance exceeded leaf death but this was reversed in June. During the rest of the season as a new leaf appeared on a tiller so the oldest leaf died.     

The effect of summer management of perennial ryegrass-dominant swards on plant and animal responses in the autumn when grazed by sheep. 1. Tissue turnover and sward structure

An experiment was conducted lo compare the effects of the grazing by ewes and weaned lambs on aftermath and previously continuously grazed perennial ryegrass-dominant swards, at two sward heights (4 and 8 cm) in (he autumn, on changes in structure and growth of the swards. The experiment had a factorial design, was replicated twice and was conducted from mid-August to early November with measurements being made on three occasions in the autumn. Aftermath swards had lower tiller population densities and lower herbage masses than those that had been previously continuously grazed, the differences being greatest in August. On an area basis net growth rates of herbage on aftermaths were lower than those on previously continuously grazed swards in August and September but not in October. Growth rates of herbage were higher on the taller sward, but the senescence rates were similar at the two sward heights. It was concluded that autumn swards may be managed at taller sward heights than summer swards without increasing senescence of the sward and a consequent reduction in efficiency of utilization. The effects of previous sward management on tissue turnover in the autumn were not long lasting.     

Effects of grazing by sheep or goats on sward composition in ryegrass/white clover pasture and on subsequent performance of weaned lambs

An experiment was conducted to assess the effects of grazing a perennial ryegrass (Lolium perenne) / white clover (Trifolium repens) sward by sheep or goats on sward composition and structure and on subsequent diet selection, herbage intake and liveweight gain by weaned lambs. From mid-May to late July (phase 1), ewes with twin lambs or yearling Scottish Cashmere goats grazed continuously swards maintained at 4- or 8-cm sward surface height. From mid-August to the end of September (phase 2), weaned lambs continuously grazed the same swards maintained at 4 cm (treatment 4–4) or at 8 cm (treatment 8–8) or which had been allowed to increase from 4 cm to 8 cm (treatment 4–8). By the end of phase 1, swards grazed by goats had higher proportions of white clover in the whole sward (0.377 vs. 0.181; s.e.d 0.0382; P < 0.001) than those grazed by sheep, irrespective of sward height treatment. This resulted in phase 2 in a higher proportion of white clover selected ( P <0.001), higher herbage intakes ( P < 0.001) and higher liveweight gains ( P < 0.001) by weaned lambs grazing swards previously grazed by goats compared with those previously grazed by sheep. There were higher proportions of clover present in the swards from treatment 4–8 at the beginning of phase 2 compared with the other sward height treatments and consequently weaned lambs had, on this treatment, a higher proportion of clover in their diet ( P <0.001), higher herbage intakes ( P <0001) and higher liveweight gains ( P <0.001). It is concluded that goats can be integrated into sequential grazing systems with sheep on grass/clover swards and this can result in an increase in the proportion of clover in swards and increased sheep performance.     

Defoliation of ryegrass-dominated swards by sheep

Patterns of defoliation in perennial ryegrass-dominated swards were examined in two experiments. In Experiment I measurements were made on four swards after a single overnight grazing. In Experiment 2 observations were made repeatedly on marked tillers in two continuously stocked swards which were grazed down over a 24-d period. In both experiments the chance of defoliation was related to lamina height both within and between age classes. Lamina angle was measured in Experiment 2 and within age classes the chance of defoliation was also independent of lamina angle. In Experiment 1 tillers most frequently had both of the two youngest laminae grazed; in Experiment 2 tillers with only lamina 1 grazed occurred most frequently, but defoliation of the two youngest laminae was again common.
The different frequencies of grazing of the different classes of laminae is interpreted as being a function of their frequency of occurrence within the grazed layer at the top of the sward. The pseudostem apparently restricted the depth of the grazed layer in the very short swards of Experiment 2.
Grazing behaviour in relation to sward structure and the patterns of defoliation and their implications for herbage production are discussed.     

Influence of structure and composition of ryegrass and prairie grass-white clover swards on the grazed horizon and diet harvested by sheep

Diet selection from ryegass-and prairie grass-white clover swards, vertically stratified into three horizons (A > 6 cm, B 3–6 cm, C > 3 cm), was studied using oesophageally fistulated sheep during summer and autumn. Animals grazed for 3-day periods. Apparent herbage intake was calculated from total herbage disappearance. The composition of each horizon and of the diet selected was measured daily.
Herbage mass (DM ha^-1) and sward height (cm) prior to grazing were not significantly different between swards in each season, and were 2·0 and 20 in summer and 1·6 and 10 in autumn. In summer, 36% and 5% of the green grass leaf (GGL) for prairie grass and ryegrass, respectively, was distributed in horizons A and B. In autumn 39% and 29% of GGL occurred above 3 cm for prairie grass and ryegrass, respectively. GGL distribution determined which sward horizons were grazed. Sheep grazed horizon C (0–3 cm) of summer ryegrass pasture, and the surface canopy (>3 cm) of all other swards.
In summer, apparent intake achieved by sheep grazing prairie grass swards was 87% higher than that achieved on ryegrass swards. In autumn a greater GGL distribution above 3 cm with prairie     

Effects of grazing by sheep or cattle on sward structure and subsequent performance of weaned lambs

The study was designed to test the hypothesis that grazing management in early season could alter sward structure to facilitate greater animal performance during critical periods. The effects of grazing a mixed perennial ryegrass/white clover sward at different sward surface heights, by cattle or sheep, in early season on sward composition and structure, and on the performance of weaned lambs when they subsequently grazed these swards in late season were determined. In two consecutive years, from mid‐May until mid‐July, replicate plots (three plots per treatment) were grazed by either suckler cows and calves or ewes and lambs at 4 or 8 cm sward surface heights (Phase 1). From mid‐August (Year 1) or early August (Year 2), weaned lambs continuously grazed, for a period of 36 d (Year 1) or 43 d (Year 2) (Phase 2), the same swards maintained at 4 cm (treatment 4–4), 8 cm (treatment 8–8) or swards which had been allowed to increase from 4 to 8 cm (treatment 4–8). Grazing by both cattle and sheep at a sward surface height of 4 cm compared with 8 cm in Phase 1 resulted in a higher (P < 0·001) number of vegetative grass tillers per m² in Phase 2, although the effect was more pronounced after grazing by sheep. Sheep grazing at 8 cm in Phase 1 produced a higher number of reproductive tillers per m² and a greater mass of reproductive stem (P < 0·001) than the other treatment combinations. The mass of white clover lamina was higher under cattle grazing (P < 0·05), especially on the 8‐cm treatment, and white clover accounted for a greater proportion of the herbage mass. These effects had mainly disappeared by the end of Phase 2. On the 4–4 and 8–8 sward height treatments the liveweight gain of the weaned lambs was higher (P < 0·05) on the swards previously grazed by cattle than those grazed by sheep. The proportion of white clover in the diet and the herbage intake also tended to be higher when the weaned lambs followed cattle. However, there was no difference in liveweight gain, proportion of white clover in the diet or herbage intake between swards previously grazed by cattle or sheep on the 4–8 sward height treatment. It is concluded that grazing grass/white clover swards by cattle compared with sheep for the first half of the grazing season resulted in less reproductive grass stem and a slightly higher white clover content in the sward, but these effects are transient and disappear from the sward by the end of the grazing season. They can also be eliminated by a short period of rest from grazing in mid‐season. Nevertheless these changes in sward structure can increase the performance of weaned lambs when they graze these swards in late season.     

THE FREQUENCY OF DEFOLIATION OF INDIVIDUAL TILLERS IN A SET-STOCKED SWARD

A method designed to study the defoliation of individual tillers in a set-stocked sward is described. This was used on S23 ryegrass swards carrying 19 (medium-stocked) and 30 (heavily-stocked) sheep/ac.
The tillers in the heavily-stocked sward were defoliated, on average, every 7–8 days, and those in the medium-stocked sward every 11–14 days. The total green leaf length (GLL) of tillers grazed by the sheep was generally greater than that of tillers not grazed. On average 27 and 40% of the GLL of tillers in the medium- and heavily-stocked swards respectively was removed at each grazing. Older leaves on the grazed tillers were defoliated much less frequently than were younger leaves.
It is suggested that a rational analysis of the relationship between the grazing animal and the grazed sward cannot be made without more work of this nature.     

The influence of sward canopy structure on foraging decisions by grazing cattle. I. Patch selection

Patch selection by grazing dairy cows in response to simultaneous variation in combinations of sward structural characteristics was examined in three experiments in which four mature dairy cows were offered a choice of patches (typically 0.9 m × 0.9 m) of perennial ryegrass (Lolium perenne) presented in a linear arrangement. Treatments involved combinations of variations in sward height, stubble height and/or depth of regrowth, prepared by preliminary cutting treatments. They were arranged in balanced sets of four to nine treatments, which were arranged in linear sequences of eighteen to twenty‐seven patches. Within experiments, sequences were balanced across replicate sets of patches, which were grazed separately by individual cows. The number of bites removed and the residence time for each patch were highly correlated in all three experiments, and the results are reported using number of bites per patch as an estimator of foraging behaviour. In the first experiment, with vegetative swards, cows preferentially selected the tallest swards. When swards comprising reproductive stem were offered in Experiment 2, cows selectively grazed short‐stubble swards rather than tall‐stubble swards, although both offered a similar depth of regrowth. Cows did not exhibit preference for swards comprising the greatest quantity of leaf mass, indicating that the spatial distribution of plant components assumed greater importance. In the third experiment, the number of bites removed increased with increasing depth of regrowth, and was negatively correlated with sward height. The three patch‐appraisal cues investigated were broadly ranked in order of importance as (i) depth of regrowth, (ii) sward maturity and (iii) sward height. There was no evidence, at least at a short temporal scale, that patch behaviour was influenced by conditions in adjacent patches, suggesting that the cows assessed grazing opportunities on a patch‐by‐patch basis.     

Effect of sward surface height on the performance of ewes and Iambs continuously grazed on grass/clover and nitrogen-fertilized grass swards

In three successive years, sward height was maintained at 3, 5, 7 or 9 cm on grass swards receiving a total of 300 kg N ha^?1 in six equal monthly dressings from April, and on grass/clover swards receiving 50 kg N ha^?1 as a single dressing in early spring. From turnout in April until weaning in July, 64 ewes and their lambs (mean litter size 1·5) were continuously grazed at the four sward surface heights on the two sward types. White clover content of grass/clover swards remained low throughout the experiment ranging from 0·2 to 7·4% of the herbage mass. During the first two years, lamb gains averaged over sward types were 204, 260, 285 and 308 g d^?1 up to weaning, while in the third year gains were 238, 296, 296 and 260 g d^?1 on 3, 5, 7 and 9 cm swards respectively. Ewes lost live weight on 3 cm swards but apart from this sward height had little effect on performance. During the autumn, weaned lambs gained — 27, 87, 147 and 167 g d^?1 on 3, 5, 7 and 9 cm swards respectively. Sward type had only a small effect on the performance of lambs up to weaning but in the autumn, mean gains of weaned lambs were lower on grass/N swards (73 g d^?1) than on grass/clover swards (115 g d^?1). Relative to 3 cm swards, carrying capacities of 5, 7 and 9 cm swards were 0·76, 0·57 and 0·52 respectively from turnout to weaning and 0·66, 0·52 and 0·44 respectively during autumn. Grass/clover swards carried 0·67 of the ewes carried by grass/N swards from turnout to weaning and 0·51 of the live weight carried by grass/N swards during autumn. The reaction of the two sward types to sward height did not appear to differ but in the third year there was evidence of a reduction in white clover content when swards were grazed at 9 cm. The data suggest that lamb growth rates will increase as sward height increases up to 9 cm and the evidence for this was stronger with weaned lambs in autumn than with suckling lambs in spring.     

The diet ingested by sheep grazing swards differing in white clover and perennial ryegrass content

A series of twenty-four swards containing different proportions of white clover (0·20-0·25) and perennial ryegrass were created by using different seed mixtures, herbicide applications and previous cutting Frequencies. These swards were used to study the diet of oesophageally-fistulated wether sheep which grazed the various swards for a 30-min period after 1, 2 and 3 weeks of regrowth.
The proportion of white clover in the diet was generally greater than that in the sward. Fifty-seven percent of the variation in the proportion of white clover in the diet could be attributed to the proportion of white clover in the sward. White clover and perennial ryegrass leaf and stem were grazed to the same height and the proportion of white clover in the grazed horizon of the sward explained 83% of the variation in the proportion of white clover in the diet. The proportion of white clover in the diet was greater than the proportion in the grazed horizon of the sward in week 3 of regrowth, but not in weeks 1 and 2, and greater when the proportion of white clover in the grazed horizon was lower than 0·20. Both these observations were interpreted as indicating selection for white clover by the sheep within the grazed horizon.
There was a positive and linear relationship between the depth of the grazed horizon and sward height which, together with the relationship between the proportion of white clover in the grazed horizon and in the diet, would allow the prediction of the proportion of white clover of the diet from the height and the white clover content of the grazed horizon of the sward.     

Lactation performance of spring-calving dairy cows grazing mixed perennial ryegrass/white clover swards of differing composition and height

An experiment was designed to examine the changes in clover content of three mixed perennial ryegrass/white cover swards of differing initial clover contents subjected to different grazing height management regimes and their effect on lactation performance of 48 Friesian dairy cows and heifers. Two paddocks were established for each treatment and grazed on alternate days. Treatments T17 and Tl3 consisted of swards with initial clover contents of 0·17 and 0·13 of the dry matter (DM) mass, respectively, grazed to maintain compressed sward heights of 6 cm throughout the season. A third treatment, SI5, consisted of a sward with an initial clover content of 0·15 grazed to maintain a compressed sward height of 4·5 cm for the first 78 days of the grazing season (period 1). Throughout period 1, half the animals on each treatment each received 4 kg of a concentrate supplement daily, while the others remained unsupplemented. From days 79 to 90, the cattle on treatment S15 grazed a similar sward, while the compressed sward height of the S15 paddocks was allowed to increase to 6 cm before re introduction of the animals. The three swards were then grazed for a further 47 days (period 2) before the animals were housed and milk yield recorded for a further 63 days (period 3). While sward T17 showed little change in clover content over the first 29 days of grazing, remaining at just below 0·18 of DM mass, swards T13 and S15 showed a marked decline in clover content to 0·05 and 0·07 of DM mass respectively. However, by the end of period 1 the clover content of all three swards had increased markedly (0·25, 0·15 and 0·15 of DM mass respectively). By the end of period 2, clover proportions were slightly higher than initial values (0·19. 0·15 and 0·15 of DM mass for treatments T17, T13 and S15, respectively). Owing to the relatively small differences in clover content of swards TI7 and T13, there were no significant effects of these two treatments on milk yield or composition in any period. Supplementation had no effect on milk composition and had little effect on milk yield, except when sward height was maintained at 4·5 cm. There was no carryover effect of supplementation on milk yield or composition in periods 2 or 3.     

An ultrasonic rangefinder for measuring the undisturbed surface height of continuously grazed grass swards

An ultrasonic rangefinder (sonic sward stick) was developed for rapidly measuring and recording the undisturbed surface height of swards. HFRO and ultrasonic sward stick measurements of undisturbed sward height over continuously grazed pasture were closely related. The measurements obtained with both instruments were related to the sward mass; the distribution of that mass between leaf, stem and dead material had little effect.     

Differences in the vertical distribution of plant material within swards continuously stocked with cattle

A mosaic of short, frequently grazed areas and tall, infrequently grazed areas is a characteristic of swards continuously stocked with cattle. Inclined point quadrat measurements were used to compare sward structure on the frequently and infrequently grazed areas in an experiment on mixed grass/clover swards with beef cattle, in which the swards were maintained at overall mean rising-plate target heights of 3·0, 5·5 or 7·0 cm. The results demonstrate large within treatment differences in the vertical distribution of plant parts between the frequently and infrequently grazed areas, and smaller differences within these areas between treatments. The consequences of these observations and the need for greater effort in recording the detailed structure within these areas in grazing experiments is noted.     

Herbage losses from tiller pulling in a continuously grazed perennial ryegrass sward

Tiller pulling was studied in a perennial ryegrass sward that was continuously grazed by cattle. The treatments included severe (sward height after grazing 25 mm), medium (50 mm) or lenient (75 mm) grazing from turn-out in April to 1 June, followed by grazing to 50 mm in the remainder of the season. Tiller pulling was confined to the midsummer-autumn period of the grazing season. The losses were most severe in swards that had been leniently grazed to a mean height of 75 mm during the spring and least severe in swards grazed to a height of 25 mm. The lenient grazing treatment allowed both the true stem development and aerial tillering whereas in the more tightly grazed swards true stem development was significantly less and aerial tillering was virtually absent. The pulled organic matter in the lenient treatment was equivalent to about 5·3% of the total harvested yield. In the severely grazed swards, pulling losses were equivalent to about 1·7% of the total harvested yield. A high rate of turnover of the pulled herbage was found in all the treatments with between 69 and 78% of the freshly pulled herbage disappearing within a week of being pulled. Tiller pulling was found to have no effect on either the subsequent autumn-winter tiller density or yields of cuts taken in the following year.