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1.
Fire severity as a key factor in post-fire regeneration of Pinus pinaster (Ait.) in Central Portugal
Context
Mediterranean pine woodlands are strongly affected by wildfires; however, there are knowledge gaps in the role of fire severity on Pinus pinaster Ait. regeneration.Objectives
The principal questions were: (a) does post-fire regeneration of Maritime Pine differ where canopies were consumed to a low vs. high degree and (b) which factors, besides fire severity, could explain these differences.Methods
Pine recruitment was monitored from 2 to 36?months after a summer wildfire in 18 25?m2 plots, equally divided over two fire severity classes based on crown consumption. Besides the degree of crown consumption, three quantitative fire severity indices as well as post-fire site conditions, seed input and understory vegetation recovery were measured.Results
Pine seedling densities were consistently higher in the plots with low than high crown consumption but due to marked spatial variability, they were only significantly different at two out of the eight sampling occasions. This variability could be explained by a quantitative index based on the diameter of twigs, as well as by seed input and post-fire site conditions (ash and litter cover).Conclusion
Fire severity was found to strongly affect the pine recruitment following wildfire, using both a qualitative visual severity index and a quantitative, more labour-intensive one. 相似文献2.
Teresa Valor Míriam Piqué Bernat C. López José Ramón González-Olabarria 《Annals of Forest Science》2013,70(5):503-513
Context
After wildfire, surviving trees are of major ecological importance as they can help in the post-fire regeneration process. Although these trees may be damaged, they may also benefit from reduced fuel hazard and competition. However, little is known about the long-term growth response of surviving trees.Aims
This study aims to explain short- to long-term variations in the postfire growth of surviving black pines in an area burnt in 1994, focusing on levels of fire severity and tree sizes.Methods
Relative basal area increments were used to detect time-course variations in postfire radial tree growth depending on fire severity. Linear mixed-effects models were used to describe the factors affecting postfire ring growth.Results
In the short term, fire caused stronger reduction in growth in small trees with increasing bole char height. However, as time since fire increased, a positive effect of fire on growth due to reduced competition counteracted the short-term fire impacts. Indeed, small surviving trees demonstrated a surge in growth 15 years after the fire.Conclusion
It was concluded that reduced competition might offset the short-term negative effects of fire in surviving black pines. 相似文献3.
Using density management diagrams to assess crown fire potential in Pinus pinaster Ait. stands 总被引:1,自引:0,他引:1
Ibán Gómez-Vázquez Paulo M. Fernandes Manuel Arias-Rodil Marcos Barrio-Anta Fernando Castedo-Dorado 《Annals of Forest Science》2014,71(4):473-484
Context
Density management diagrams (DMDs) are useful for designing, displaying and evaluating alternative density management regimes for a given stand-level management objective. The inclusion of variables related to crown fire potential within DMDs has not previously been considered.Aims
The aim of this study was to include isolines of variables related to crown fire initiation and spread in DMDs to enable identification of stand structures associated with different types of wildfire.Methods
Biometric and fuel data from maritime pine (Pinus pinaster Ait.) stands in NW Spain were used to construct DMDs. Different surface and crown fire behaviour models were used together to estimate crown fire potential.Results
The crown fire potential varied greatly throughout development of the maritime pine stands. Low stands were more prone to crowning. The type of crown fire was mainly determined by stand density.Conclusion
The DMDs developed can be used to identify relationships between stand structure and crown fire potential, thus enabling the design of thinning schedules aimed at reducing the likelihood of crowning. 相似文献4.
Models of the longitudinal distribution of ring area as a function of tree and stand attributes for four major Canadian conifers 总被引:1,自引:0,他引:1
Context
It is widely accepted that ring area increment generally increases from the tree apex to the crown base and is more-or-less constant below the crown base (Pressler’s law), but few quantitative models of this distribution have been developed.Aims
The aim of this study was to develop a model of ring area increment using easily obtained crown features and other tree or stand characteristics in order to further the understanding and prediction of tree growth, form, and wood quality.Methods
The models were fit to stem analysis observations from white spruce, black spruce, balsam fir, and lodgepole pine.Results
In the final model, which includes tree crown and stand variables, ring area increment within the crown region was slightly curvilinear, the slope of ring area increment below the crown was non-zero, and the effect of butt swell was appreciable up-to a relative height of 0.10.Conclusions
The high accuracy of the mixed effects model suggests that the three-component model form is appropriate for describing ring area profiles, whereas some tree-to-tree variation remains unexplained. The tree and stand variables used in these models can be easily measured in the field or obtained from remote sensing techniques. 相似文献5.
Context
In the Mediterranean area, different post-fire management strategies are used for coppices of resprouting species to promote a more regular forest structure, enhance plant growth, and reduce fire risk. However, the effects of these management treatments on forest-associated fauna are unknown, which in turn could be limiting their beneficial effects.Aims
The aim of this work was to determine whether forest management of a recently burned area dominated by a vigorous resprouting tree species (Arbutus unedo L.) affects ant communities.Methods
Ant communities, sampled using pitfall traps, were examined from unmanaged and selective thinning coppices of A. unedo. Ants are here used as bioindicators of ecosystem health and surrogates for other animal groups.Results
Very limited effects of these post-fire management strategies on the structure and composition of ant communities were found. The lack of effects could be due to the reported small changes in physical conditions among treatments; or either, the most sensitive ant species to these post-fire management treatments might be the same ones affected by fire and, consequently, the ant species that would potentially be affected most were no longer in the study area.Conclusion
The lack of any significant effects caused by these post-fire management practices on the associated fauna of A. unedo coppices points out the suitability of these treatments in these circumstances. 相似文献6.
J. Julio Camarero Rubén D. Manzanedo Raúl Sanchez-Salguero Rafael M. Navarro-Cerrillo 《Annals of Forest Science》2013,70(8):769-780
Context
Tree populations at the rear edge of species distribution are sensitive to climate stress and drought. However, growth responses of these tree populations to those stressors may vary along climatic gradients.Aims
To analyze growth responses to climate and drought using dendrochronology in rear-edge Pinus nigra populations located along an aridity gradient.Methods
Tree-ring width chronologies were built for the twentieth century and related to monthly climatic variables, a drought index (Standardized Precipitation–Evapotranspiration Index), and two atmospheric circulation patterns (North Atlantic and Western Mediterranean Oscillations).Results
Growth was enhanced by wet and cold previous autumns and warm late winters before tree-ring formation. The influence of the previous year conditions on growth increased during the past century. Growth was significantly related to North Atlantic and Western Mediterranean Oscillations in two out of five sites. The strongest responses of growth to the drought index were observed in the most xeric sites.Conclusion
Dry conditions before tree-ring formation constrain growth in rear-edge P. nigra populations. The comparisons of climate-growth responses along aridity gradients allow characterizing the sensitivity of relict stands to climate warming. 相似文献7.
Key message
Pinus radiata trees showed significantly reduced basal area increments and increased latewood/earlywood ratios, when their stem was charred by surface fires even if no needle damage occurred. An interaction of fire damage and precipitation on growth was observed.Context
Heat from forest fires is able to penetrate beyond the bark layer and damage or completely kill a tree’s cambium. Short-term growth reductions following surface fires have been reported for some species. However, most studies have in common that they describe a compound effect of stem and foliage damage.Aims
This study investigated the impact of surface fires on the radial growth of Pinus radiata, where only the stem of trees was charred, while no needle damage was recorded.Methods
Tree ring measurements were performed on cores obtained at breast height. Analysis of variance and tests, based on annual basal area increment values were calculated to quantify pre- and post-fire growth differences of tree ring width and latewood/earlywood ratios.Results
The analysis revealed significant growth reductions following a surface fire on P. radiata in the year on which the fire occurred as well as in the following year. As a consequence of the fire, basal area increment and latewood/earlywood ratios were significantly reduced. An interaction of fire damage and precipitation on growth was observed.Conclusion
The obtained results show how fires without crown damage can affect growth and tree ring structure of P. radiata trees and indicate that stem char could be associated with a significant decrease in ring width and latewood/earlywood ratio.8.
9.
Context
Excessive recruitment in post-fire regeneration of Pinus halepensis can require pre-commercial thinning. The 1994 Moratalla fire (Spain) and the thinning applied there since 2004 provided good conditions for testing pre-commercial thinning effectiveness.Aims
To analyse pre-commercial thinning effects on tree size, reproductive potential, stem biomass and annual growth in 15-year-old saplings.Methods
Twenty nine circular plots (5 m radius) were established based on (1) years since thinning and (2) aspect. Non-thinned plots were included as controls. Sapling variables were measured and pine cones were counted and classified according to maturity. Three saplings per plot were felled, five sample disks were extracted from each and tree rings were analysed with Windendro® software.Results
Sapling size, annual growth and stem biomass were higher in thinned plots than in control plots 2–4 years after thinning. Annual growth declined five years after thinning. Cone counts were higher 2–5 years after thinning, depending on the cone type. Aspect influenced some variables.Conclusion
Thinning accelerated stand maturity but positive effects were indiscernible until 2–5 years later. Serotinous cones, which are fundamental to regeneration after fire, increased in number after 5 years. The effects of aspect were inconclusive due to interactions with years since thinning. 相似文献10.
Variations of wood basic density with tree age and social classes in the axial direction within Pinus massoniana stems in Southern China 总被引:1,自引:0,他引:1
Xiangwen Deng Liyun Zhang Pifeng Lei Wenhua Xiang Wende Yan 《Annals of Forest Science》2014,71(4):505-516
Context
Reliable estimates of wood density (WD) within individual trees could maximize the value of Pinus massoniana for specific end-use.Aim
We examined and quantified the axial patterns of WD in trees with different social status in the stands.Methods
Wood disks were sampled at the bottom, breast height, and middle of each 1-m sections from 108 stems, harvesting from three social classes in subtropical forests. A mixed-effects model was designed to quantify axial variation.Results
The WD at different height was significantly different from the whole-stem WD (WWD) except the relative height of 0.1. An overall decrease of 133.8 kg m?3 in WD was found from stem base to top. WD was significantly influenced by relative heights, tree age, and social class. WD of each relative height in mature trees was significantly higher than that of younger trees. Tree social class can affect WD development in the axial direction at age classes 2 and 3. Combining the fixed plus random effects, the final model explained 91 % of the observed variation in WD.Conclusion
The WD development patterns in the axial direction vary considerably among tree age, diameter at breast height, and social class. To distinguish and supply timber for specific end-uses, we should use the axial variation in disk WD (DWD) instead of WWD directly. The accurate predictions of WD provided by the model could be used to optimally classify logs into different product classes and maximize economic benefits. We can use DWD at the relative height of 0.1 instead of WWD of a single tree. 相似文献11.
Daniele Castagneri Assunta Esposito Giovanni Bovio Stefano Mazzoleni Umberto Seneca Ambra Elena Catalanotti Davide Ascoli 《Annals of Forest Science》2013,70(8):863-871
? Context
Prescribed burning is increasingly recommended to control encroaching shrublands in the Mediterranean area.? Aims
The aims of this paper are to analyze the fuel structural characteristics of Spartium junceum and how they influence fire behaviour during prescribed burning.? Methods
Two winter–spring prescribed burns were conducted in 2009 and 2011. Fuel load and structure of S. junceum shrubs were assessed using the Cube Method, and shrub 3-D models were built using the FIRE PARADOX FUEL MANAGER software. Allometric equations to estimate S. junceum fuel load were developed. During burning, thermocouples measured temperature variations, which were then analyzed relative to fuel characteristics.? Results
Fuel load components and distribution were strictly related to shrub height; in tall shrubs, most of the fine fuel was more than 1.5 m aboveground. Due to fuel vertical discontinuity, not all shrubs were burned in the fires, but wind increased fire sustainment and fuel consumption. Maximum temperatures (over 800 °C) and residence times were positively related to fuel load.? Conclusion
S. junceum tall shrublands represent high hazard formations due to their elevated fuel load, mostly in fine fuel fractions. Vertical discontinuity among fuel strata limits fire propagation in mild weather conditions. Winter–spring prescribed burning cannot eliminate S. junceum shrublands, but do create shrub cover discontinuity. As S. junceum has fire-adapted morphological traits, a single burn is insufficient to control it. 相似文献12.
Sergio de-Miguel Timo Pukkala Nabil Assaf Zuheir Shater 《Annals of Forest Science》2014,71(1):101-112
? Context
Biomass prediction is important when dealing for instance with carbon sequestration, wildfire modeling, or bioenergy supply. Although allometric models based on destructive sampling provide accurate estimates, alternative species-specific equations often yield considerably different biomass predictions. An important source of intra-specific variability remains unexplained.? Aims
The aims of the study were to inspect and assess intra-specific differences in aboveground biomass of Pinus brutia Ten. and to fill the gap in knowledge on biomass prediction for this species.? Methods
Two hundred one trees between 2.3 and 55.8 cm in diameter at breast height were sampled throughout the eastern- and southernmost natural distribution area of P. brutia, in Middle East, where it forms different stand structures. Allometric equations were fitted separately for two countries. The differences in biomass prediction at tree, stand, and forest level were analyzed. The effect of stand structure and past forest management was discussed.? Results
Between-country differences in total aboveground biomass were not large. However, differences in biomass stock were large when tree components were analyzed separately. Trees had higher stem biomass and lower crown biomass in dense even-aged stands than in more uneven-aged and sparse stands.? Conclusion
Biomass and carbon predictions could be improved by taking into account stand structure in biomass models. 相似文献13.
Delphine Jullien Robert Widmann Caroline Loup Bernard Thibaut 《Annals of Forest Science》2013,70(2):133-142
Aims
In European Beech (Fagus sylvatica L.) large growth stresses lead to severe log end splitting that devaluate beech timber. Our study aimed at detecting relationships between growth stress and some morphology parameters in trees.Methods
Growth stress indicators were recorded for 440 mature trees in nine stands from five European countries, together with morphology parameters.Results
Most trees displayed an uneven distribution of growth stress around the trunk. Moreover, growth stress intensity varied largely between individual trees. Geometry of the trunk was a poor predictor of growth stress intensity. Crown asymmetry resulted in a larger stress dissymmetry within trees. Trunk inclination was not correlated to max tension stress, contrary to what is usually found in younger trees. In the case of small inclination, growth stress was close to expected from biomechanics of restoring verticality. Trees exhibiting a larger inclination probably evolved a different mechanical solution: a rather large crown, lower tree slenderness and a sufficient asymmetry in growth stress as to prevent a higher inclination due to growth.Conclusion
A large slenderness is the best accurate predictor of a large growth stress, although variations in the ratio height/diameter at breast height explained only 10 % of the variability of growth stress. A large crown surface was the best predictor of a low level of growth stress. A large spacing between trees seems a good solution to lower the risk of growth stress in mature beech. 相似文献14.
Felipe Crecente-Campo José Javier Corral-Rivas Benedicto Vargas-Larreta Christian Wehenkel 《Annals of Forest Science》2014,71(1):51-70
? Context
Tree height prediction is an important issue in forest management since tree heights are usually measured only in a sample of trees. Although numerous model approaches have been used for this purpose, no agreement on which one is more appropriate has been achieved.? Aims
To analyse the random effects of basic and generalised height–diameter (h–d) models fitted to multi-species uneven-aged forest stands, and to establish their ability to explain differences between ecoregions, plots and species.? Methods
Height and diameter measurements for 29,084 trees from 187 sample plots located in the state of Durango (Mexico) were used. Basic and generalised h–d models were fitted in a mixed-models framework. The variability between ecoregions, plots and species was considered in the random effects definition. Model calibration for different height sampling designs and sampling sizes was also analysed.? Results
Random components performed well in explaining the differences in the h–d relationship between the different plots and species; however, no significant variance for the random effects was found for the different ecoregions. A calibrated basic h–d model produced similar results to a fixed-effects generalised h–d model when a sufficiently large number of trees was used in the calibration process.? Conclusion
From a practical point of view, if no calibration is carried out, different models should be used for the different species, so that at least the variation among species is captured. 相似文献15.
Elizabeth Kearsley Pieter CJ Moonen Koen Hufkens Sebastian Doetterl Janvier Lisingo Faustin Boyemba Bosela Pascal Boeckx Hans Beeckman Hans Verbeeck 《Annals of Forest Science》2017,74(1):7
Key message
Tree heights in the central Congo Basin are overestimated using best-available height-diameter models. These errors are propagated into the estimation of aboveground biomass and canopy height, causing significant bias when used for calibration of remote sensing products in this region.Context
Tree height-diameter models are important components of estimating aboveground biomass (AGB) and calibrating remote sensing products in tropical forests.Aims
For a data-poor area of the central Congo Basin, we quantified height-diameter model performance of local, regional and pan-tropical models for their use in estimating AGB and canopy height.Methods
At three old-growth forest sites, we assessed the bias introduced in height estimation by regional and pan-tropical height-diameter models. We developed an optimal local model with site-level randomizations accounted for by using a mixed-effects modeling approach. We quantified the error propagation of modeled heights for estimating AGB and canopy height.Results
Regional and pan-tropical height-diameter models produced a significant overestimation in tree height, propagating into significant overestimations of AGB and Lorey’s height. The pan-tropical model accounting for climatic drivers performed better than the regional models. We present a local height-diameter model which produced nonsignificant errors for AGB and canopy height estimations at our study area.Conclusion
The application of general models at our study area introduced bias in tree height estimations and the derived stand-level variables. Improved delimitation of regions in tropical Africa with similar forest structure is needed to produce models fit for calibrating remote sensing products.16.
Marianne Peiffer Nathalie Bréda Vincent Badeau André Granier 《Annals of Forest Science》2014,71(7):821-829
Context
In the context of a probable increase in intensity and frequency of extreme summer drought events, a better understanding of the key processes involved in water relations is needed to improve the theoretical foundations of predictive process-based models.Aims
This paper aims to analyse how temperate deciduous trees cope with water shortage.Methods
The exceptional summer drought of 2003 in Europe provided an opportunity to monitor stomatal conductance and twig water potential in European beech (Fagus sylvatica L.) at predawn and midday and to analyse variations with respect to leaf height within the canopy. By comparing our field measurements of twig water potential to values found in the literature, we confirmed the strong impact of soil water shortage on crown water relations.Results
This paper shows that (1) the vertical gradient of stomatal conductance within the crown disappeared under extreme soil water depletion; (2) at maximum drought intensity, predawn twig water potential (ψ pd) reached ?2.3 MPa at a height of 14 m in the crown and ?2.0 MPa at a height of 10 m. The significant differences in ψ pd between the two measurement heights in the canopy may be due to night transpiration; (3) there was a close relationship between predawn twig water potential and relative extractable soil water; (4) as drought conditions intensified, there was a close relationship between canopy radiation interception and predawn water potential, as estimated daily from relative extractable soil water. 相似文献17.
18.
Variation in biomass expansion factors for China’s forests in relation to forest type, climate, and stand development 总被引:1,自引:0,他引:1
Yunjian Luo Xiaoke Wang Xiaoquan Zhang Yin Ren Hendrik Poorter 《Annals of Forest Science》2013,70(6):589-599
? Context
Biomass expansion factors (BEFs, defined as the ratios of tree component biomass (branch, leaf, aboveground section, root, and whole) to stem biomass) are important parameters for quantifying forest biomass and carbon stock. However, little information is available about possible causes of the variability in BEFs at large scales.? Aims
We examined whether and how BEFs vary with forest types, climate (mean annual temperature, MAT; mean annual precipitation, MAP), and stand development (stand age and size) at the national scale for China.? Method
Using our compiled biomass dataset, we calculated values for BEFs and explored their relationships to forest types, climate, and stand development.? Results
BEFs varied greatly across forest types and functional groups. They were significantly related to climate and stand development (especially tree height). However, the relationships between BEFs and MAT and MAP were generally different in deciduous forests and evergreen forests, and BEF–climate relationships were weaker in deciduous forests than in evergreen forests and pine forests.? Conclusion
To reduce uncertainties induced by BEFs in estimates of forest biomass and carbon stock, values for BEFs should be applied for a specified forest, and BEF functions with influencing factors (e.g., tree height and climate) should be developed as predictor variables for the specified forest. 相似文献19.
Jorge Olivar Stella Bogino Cyrille Rathgeber Vivien Bonnesoeur Felipe Bravo 《Annals of Forest Science》2014,71(3):395-404
? Context
Modification of stand density by thinning may buffer the response of tree growth and vigor to changes in climate by enhancing soil water availability.? Aims
We tested the impact of thinning intensity on cambial growth of Aleppo pine (Pinus halepensis L.) under semi-arid, Mediterranean conditions.? Methods
A multiple thinning experiment was established on an Aleppo pine plantation in Spain. We analysed the stem growth dynamics of two different crown classes under four different thinning intensities (15 %, 30 %, and 45 % removal of the basal area) for 2 years, based on biweekly band dendrometer recordings. Local relative extractable soil water was derived from the use of a water balance model Biljou© (available at https://appgeodb.nancy.inra.fr/biljou/) and used as an explanatory variable.? Results
Radial growth was mainly controlled by soil water availability during the growing season, and differed by crown class. The growth rates of dominant trees were significantly higher than the growth rates of suppressed trees. Removal of 30 % and 45 % of the initial basal area produced a growth release in both dominant and suppressed trees that did not occur under less intense thinning treatments.? Conclusions
Soil water availability was the main driver of radial growth during the growing season. Forest management confirmed its value for ameliorating the effects of water limitations on individual tree growth. These results may help managers understand how altering stand density will differentially affect diameter growth responses of Aleppo pine to short-term climatic fluctuations, promoting forests that are resilient to future climatic conditions. 相似文献20.