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1.
The Influence of Desiccation, Rough Handling and Cold Storage on the Quality and Establishment of Sitka Spruce Planting Stock 总被引:2,自引:0,他引:2
DEANS J. D.; LUNDBERG C.; TABBUSH P. M.; CANNELL M. G. R.; SHEPPARD L. J.; MURRAY M. B. 《Forestry》1990,63(2):129-141
In January 3-year-old Sitka spruce were lifted and treated eithercarefully or roughly. Plants in each treatment were either transferredto the laboratory for testing, or transferred to cold storageat 1°C. In the laboratory, half of the plants ineach treatment were desiccated to shoot water potentials ofless than 2.0 MPa. Plants were then tested for root growthpotential (RGP), bud dormancy, frost hardiness and carbohydratecontent before transplanting at a field site, Additional plantswere removed from the nursery and cold store at approximatelymonthly intervals until late April, and then treated as above.In April, plants in each treatment were transferred to on-sitefield storage (sheughs), for 2 or 4 weeks prior to testing asabove. Carefully handled plants maintained large RGP which was positivelycorrelated with plant water potential, root water content, shootrelative growth rate and field survival. Overall RGP was reduced: 59 per cent by desiccation; 85 percent by rough handling; and 98 per cent by desiccation and roughhandling in combination. On site storage for 4 weeks in April/May reduced RGP. Whereplants produced more than 30 new roots > 1 cm long in RGPtests, field survival exceeded 90 per cent. Cold storage at1°C maintained RGP, (of carefully handled plants),bud dormancy and shoot frost hardiness at approximately pre-storagevalues. In contrast, total nonstructural carbohydrate contentdeclined in cold store but increased in March/April for plantsin the nursery. 相似文献
2.
Seasonal Changes in the Frost Hardiness of Provenances of Picea sitchensis in Scotland 总被引:6,自引:0,他引:6
Changes in the natural level of frost hardiness of shoots offour provenances of Picea sitchensis were monitored over twogrowing seasons by detaching shoots from 7 to 10-year-old treesgrowing in a nursery in Scotland, and subjecting them to freezingtemperatures under conditions which simulated night frosts. Six seasonal phases of frost hardiness were identified (Fig.3).
- During each autumn, killing temperatures (the level of hardiness)decreased from 5°C to below 20°C, beginningseveral weeks after shoot elongation ceased. Alaskan provenanceshardened in September, apparently in response to shorteningday lengths alone, whereas an Oregon provenance did not hardenuntil November, after repeated frosts. Queen Charlotte Islandsprovenances were intermediate.
- From November to March allprovenances were hardy to below 20°C,which is adequateto prevent direct freezing injury at mostplantation sites.
- In March-April, several weeks before bud-burst, old shootsdehardenedto killing temperatures of about 10°Cin responseto warm temperatures, and southerly provenancesdid so beforenortherly ones.
- During bud-burst the newly-emergingshoots were hardy to only3°C to 5°C untilthey were about 3.5 cmlong. All provenances burst bud at thesame time and were equallyfrost susceptible at this time.
- DuringMay-July the elongating shoots fluctuated in hardinessbetween5°C and 10°C apparently in responsetofluctuating ambient temperatures.
- In August 1980 there wasa period of late summer dehardeningto killing temperaturesof about 3°C.
3.
At two English forest nurseries, transplants of five coniferspeciesPicea sitchensis, Picea abies, Tsuga heterophylla,Abies grandis, and Pinus contortawere grown with fertilizersupplying N, P, K, and Mg in amounts intended to be adequatefor producing healthy green trees with nutrient concentrationsin the sufficiency range as determined by earlierexperiments. Luxury uptake of nitrogen was obtainedwith top-dressings of Nitro-Chalk applied in thenursery during early September, when top growth had nearly ceased.Tests of the effect of this extra N on forest establishmentwere repeated in four successive years under a wide range ofsoil and climatic conditions, keeping the trees in a cold storeduring each winter and planting them on forest sites in England,Scotland, or Wales during the following spring. Except for Grandfir, nitrogen advanced bud-break of all species during the firstsummer after planting and had no deleterious influence on survival.It tended to increase growth of Sitka spruce during the seasonafter planting, but in later years the differences became smallin relation to tree size. The effects on other species weresmall, except for one considerable decrease in the growth responseof Grand fir at a single site. Frost damage of Sitka spruce of Washington origin was severeon a Welsh and a Scottish site where this frost-sensitive provenancewould not normally be grown. At the Welsh, but not the Scottishsite, the nitrogen treatment increased the damage. In the few experiments (confined to Picea sitchensis) whichtested late-season potassium in the nursery, K concentrationswere increased from deficiency to barely sufficiency level;growth in the forest was increased in two of the four experiments.The extra K had no effect on frost damage. 相似文献
4.
In a 14-week study, 1-year-old Aleppo pine seedlings were grownin two growth chambers. Seedlings were artificially hardenedby decreasing photoperiod and temperature. In each chamber halfof the seedlings were fertilized with nitrogen (8.4 mg seedling1).In order to determine the relative importance of the hardeningenvironment versus fertilization, each chamber was programmedto decrease night temperatures down to a low of 8 or 4°C.Chlorophyll fluorescence and frost hardiness was measured fivetimes during the experiment. A sample of seedlings from eachtreatment was exposed to an artificial frost at 5°Cand the freezing effects were assessed by measurements of chlorophyllfluorescence and visual evaluation of needle damage. Seedlingsincreased their frost hardiness during the experiment in allthe treatments but the ratio of variable to maximal chlorophyllfluorescence (Fv/Fm) measured before freezing did not vary duringthe experiment. This indicates that Aleppo pine maintains itsphotosynthetic ability during hardening in contrast to otherconiferous species from colder climates. The effect of nitrogenfertilization on frost hardiness was small in comparison withchamber effect. Nitrogen fertilization slightly delayed theacquisition of hardening in the coldest chamber. Seedlings inthe warmest chamber did not become fully resistant to 5°C,but in the coldest chamber, where night temperature reached4°C, all the seedlings were resistant to the frost. Severedamage caused by frost could be related to a rapid rise of minimalfluorescence (F0) but the best index of damage was the dropof Fv/Fm after freezing. 相似文献
5.
Frequent bud frost damage in cultivation of Abies procera Rehderand pending climate changes are the background for this studyof cold hardiness under varying acclimation regime (in closed-topchambers) and experimental warm spells during the cold season.LT50 values were established by freezing tests at differenttimes of year. Damage and deaths were assessed on leader buds,subapical lateral buds, needles and cambium. Minor parallelexperiments involved Abies nordmanniana, Picea abies and Piceasitchensis. Lower acclimation temperatures resulted in deeperfrost hardiness during late autumn but less during spring, comparedwith ambient temperature controls. Elevated temperatures resultedin less deep frost resistance. Apical buds generally developeddeeper frost hardiness than lateral buds but less deep thanthe cambium, varying with species, however. Frost damage inbuds ranged from death over partially destroyed bud contentsresulting in distorted shoots to buds seemingly remaining dormant.Responses to warm spells differed with duration, timing andspecies, ranging from dramatic decrease in frost hardiness withor without subsequent recovery to no reaction. Furthermore,the reactions did not show any clear relation to dormancy level.For A. procera, exposure to fluctuating temperatures appearedto be particularly problematic. This explains why this speciesdevelops best in coastal climates, and in sites sheltered fromtemperature extremes either by hedging, a winter snow cover,or topography. The Christmas tree production will suffer severelyon sites with harsh temperatures due to losses of lateral andterminal buds, which destroy the crown symmetry. Clipping ofgreenery is less influenced by frost damages, although the developmentof normal branch whorls is often disturbed. 相似文献
6.
Morphological and physiological differences in Scots pine seedlings of six seed origins 总被引:1,自引:0,他引:1
One-year-old seedlings of Scots pine (Pinus sylvestris L.) offour native seed origins (Loch Maree Islands, Glengarry/GlenMorriston, Glen Affric and Abernethy), a commercial Britishseedlot, and a seedlot from Hedesunda, in middle Sweden, werecompared at monthly intervals from October 1993 to April 1994.Seedling morphology, root condition, root frost hardiness andbud dry matter were determined at each date. There were clear morphological differences among seed origins.Seedlings raised from the commercial seedlot (A70) were largerbut had a poorer root:shoot ratio than the other seed origins.Of the native pines tested, the Loch Maree Islands origin allocateda larger proportion of its photosynthate to fine roots and needlesand smaller proportion to woody structures. Seedlings raised from the commercial British seedlot tendedto have poorer bud lignification than the other origins andalso, in autumn, higher electrolyte leakage rates from its fineroots. During winter, the Swedish origin had the lowest fineroot electrolyte leakage. Seedlings of all origins showed aprogressive increase in fine root hardiness towards mid-winterwith maximum hardiness (7°C) in January. Dehardeningoccurred over subsequent months reaching 3°C in April.Differences among origins were evident. The Swedish seedlotdeveloped greater frost resistance than the other origins, hardeningbegan earlier in autumn and dehardening began later in spring.The commercial seedlot hardened later than the other originsbut reached a similar level of frost hardiness by January. Ofthe native pines, seedlings of the Loch Maree Islands originwere slowest to develop root hardiness. 相似文献
7.
Gary A. Ritchie 《New Forests》1986,1(1):29-42
An experiment was conducted to ascertain what relationships might exist among dormancy status, cold hardiness and stress resistance in 2+0 Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), seedlings. Seedlings were lifted from a western Washington nursery on six dates spanning the 1980–81 lifting season. On each date samples of seedlings were subjected to the following treatment: (1) tumbling for 5 minutes, (2) desiccation of roots for 30 minutes at 30°C and 2.1 kPa vapor pressure deficit, (3) exposure of shoots to temperatures of –10°C, –15°C or –20°C for two hours and (4) unstressed control. On two lift dates sub-samples of seedlings were placed into –1°C storage and held for two months before the above stress treatments were administered. Bud dormancy status was determined, using a bud break test, on seedlings from each lift date before and after storage.After one growing season in the field percent survival, vigor, height growth and shoot and root weight were determined on stressed and unstressed seedlings. Survival and vigor were less affected by the stress treatments than were height and weight. Severity of stress was in the order –20°C > –15°C > desiccation > handling > –10°C. Degree of cold injury was directly related to seedling dormancy status whether dormancy status had been attained in the nursery from natural chilling or in frozen storage. Seedlings in a mid-range of dormancy release (between deep rest and quiescence) were most resistant to all imposed stresses. 相似文献
8.
In an 11-year-old stand of Sitka spruce (Picea sitchensis [Bong.]Carr.) application of nitrogen fertilizers, at a rate of 10kgN ha1 month1, increased mean diameter incrementby 12 per cent, while the further addition of phosphorus, at5 kg ha1 month1, resulted in a 23 per cent increase.An attack by the green spruce aphid (Elatobium albietinum Walker)occurred during the period of fertilizer addition. The mostseverely affected trees showed a reduction in diameter growthof 50 to 56 per cent but the severity of the attack betweentrees was unrelated to the treatments applied. However, fertilizerapplication did hasten the recovery of diameter growth afterdefoliation. 相似文献
9.
Laboratory freezing tests were used to determine seasonal changesin the frost hardiness of detached shoots of young trees ofEucalyptus gunnii (from central Tasmania), and E. niphophilaand E. debeuzevillei (snow gums from the SnowyMountains, Australian Capital Territory). The trees were growingat the Bush Estate. No difference was found between the speciesor between seedlots, all of which were from high altitudes nearthe tree line. In midwinter (February) the shoots tolerated 16°Cwithout suffering damage, and many shoots survived temperaturesas low as 18°C to 22°C. This result agreedwith Evans' (1986) observation that some trees within theseseedlots survived temperatures in the range 19°Cto 23°C in field planting during the winter 1981/82.In their native habitats the trees rarely experience temperaturesbelow 20°C. The shoots did not harden appreciably before they experiencedfrosts (in late October/early November) and so may be proneto autumn frost damage. By contrast, they were slow to dehardenin late winter and spring and did not seem prone to spring frostdamage. There were no killing air frosts during the period of this study(winter 1985/6), but many trees died, possibly as a result ofground freezing, producing root injury and/or shoot desiccation.In subalpine regions of Tasmania and the Snowy Mountains theground is covered by snow throughout the winter. Variation in frost hardiness within these hardy seedlots couldbe exploited. 相似文献
10.
In direct-seeded woodlands and nursery seed-beds, weeds canrapidly invade newly sown areas, leading to death or suppressionof tree seedlings. Hand weeding is usually expensive, and thesafe use of broad-spectrum contact herbicides is seldom possible.Hence in the work reported here, the tolerance of young treeseedlings to the potentially more selective, post-emergenceherbicides clopyralid, cycloxydim and metazachlor, was tested.Seedlings of Fraxinus excelsior L. (ash) with two to four expandedtrue leaves (24 ETL), Fagus sylvatica L. (beech) (24ETL), Acer pseudoplatanus L. (sycamore) (46 ETL), Prunusavium L. (cherry) (68 ETL) and Quercus robur L. (oak)(68 ETL) appeared to tolerate applications of cycloxydimat a rate of 0.45 kg a.i. ha1. Applications of 0.2 kga.i. ha1 clopyralid or 1.25 kg a.i. ha1 metazachloralso appeared to be generally tolerated, but did cause somesuppression of annual growth increment in Q. robur, P. avium,F. sylvatica and A. pseudoplatanus, particularly where repeatapplications were made to the earliest growth stages. Seedlingsurvival was unaffected by any herbicide treatment. Mixturesof all three herbicides were no more damaging than the sameherbicides applied separately. Therefore, depending on treespecies and growth stage, it appears that clopyralid could potentiallybe safely used to control a range of herbaceous weed species,metazachlor a variety of seedling or germinating weeds and cycloxydima range of established grass weed species, in direct sown woodlandsor nursery seed-beds, although further research is advisableto confirm crop safety. 相似文献
11.
Difficulties overwintering container stock in northern Ontario led to the development of the extended greenhouse culture hardening regime for spruce seedlings. Laboratories to measure shoot frost hardiness and evaluate terminal bud development were established to monitor nursery crops being hardened using this regime. Information on frost hardiness and bud development provided by these laboratories has been used by nursery managers to determine readiness of container seedlings for overwintering. Since 1982, over 200 stock lots have been monitored by these operational laboratories. This database can be used to determine the importance of nursery cultural factors and seed source on frost hardening. The database shows large differences between nurseries in approach to hardening seedlings which were reflected in levels of freezing damage, winter desiccation and overwintering success. Rates of frost hardening (i.e., the interval between terminal bud initiation and attainment of a --15°C level of shoot frost hardiness) of crops produced in north central Ontario failed to show significant seed source effects. The rate of frost hardening was faster in crops producing fewer needle primordia in terminal buds. 相似文献
12.
Sitka spruce and Douglas fir transplants were kept under threecold storage regimes: (1) bare-root in a humidified cold store(H); (2) sealed in polythene bags in a humidified cold store(HP); (3) sealed in polythene bags in a direct cold store (D).Regimes H and D are routinely used in current practice. Plants of both species had significantly (P < 0.001) lowerroot moisture content (RMC) in treatment H within 4 weeks ofstorage; this difference was maintained throughout the periodof storage and potentially damaging levels of RMC were reachedwithin 8 weeks. Sitka spruce had significantly (P < 0.001)higher root growth potential (RGP) than Douglas fir throughoutthe storage period. Differences in RGP between storage treatmentswere not significant until 12 weeks after storage when treatmentH was poorer (P < 0.05) than treatments HP and D. The survival of plants from treatment H after 1 year in thefield was significantly poorer (P < 0.001) than the othertreatments in both species. Height increment was also less intreatment H than the other treatments (P < 0.001) after 1year. However, this difference disappeared in the second season.After 1 year there were no differences in root collar diameter(RCD) increment between storage treatments but Douglas fir showedgreater increment (P < 0.001) than Sitka spruce. Rapid assessment of RMC in slightly over 48 h makes this a potentiallyuseful test of planting stock quality during and after coldstorage. 相似文献
13.
The phenology and frost hardiness of shoots of 15 provenancesof Alnus rubra growing in Scotland were measured over one autumn,winter and spring. Dates of budset (in September) and the onsetof rapid frost hardening (in October-November) occurred about2 days earlier for each degree latitude of origin northwards,except for an Idaho provenance. However, all provenances dehardenedat about the same time in March and burst their buds between8 and 14 April. Assuming that rapid frost hardening in the autumnwas triggered primarily by shortening daylengths, Alaskan provenancesof A. rubra seemed better adapted to British conditions thansouthern British Columbian provenances, which have been mostcommonly planted. However, even Alaskan provenances are proneto spring frost damage. Scottish A. glutinosa and Alaskan A.sinuata set buds and frost hardened 12 weeks before eventhe Alaskan A. rubra, and burst their buds 23 weeks laterin April-May. All three species were hardy to below 30°Cfrom December to mid-March. 相似文献
14.
Greenhouse-cultured, container-grown ponderosa pine (Pinus ponderosa var. scopulorum Engelm.), interior Douglas-fir (Pseudotsuga menziesii var. glauca (Beissn.) Franco) and Engelmann spruce (Picea engelmannii (Parry) Engelm.) were cold acclimated and deacclimated in growth chambers over 19 weeks. Stem cold hardiness, total new root length at 14 days and days to bud break were measured weekly. Relationships among cold hardiness, root growth potential (RGP) and bud dormancy suggest that cold hardiness, which can be measured quickly, could provide a useful basis for estimating the two other parameters. During cold acclimation, there was a lag period in which stem cold hardiness remained at -15 degrees C and RGP was at a minimum, in all three species. Douglas-fir and Engelmann spruce buds remained fully dormant during this lag period. Ponderosa pine buds had no chilling requirement for the loss of dormancy, and reached quiescence during the lag period. Immediately following the lag period, as stem cold hardiness progressed to -22 degrees C, RGP increased to a high plateau in all three species, and Douglas-fir and Engelmann spruce buds approached quiescence. Cold deacclimation and bud development began immediately on exposure to warm, long days, but RGP remained high until stem cold hardiness returned to approximately -15 degrees C. At bud break, cold hardiness and RGP were at the minimum. 相似文献
15.
Cuttings of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) from three open-pollinated families were rooted in two types of tray, and then grown for 1.5 years in a bareroot nursery. During their second winter they were sampled periodically and tested for cold hardiness, dormancy status, root growth potential and various morphological characteristics. Two-year-old seedlings and transplants were tested concurrently for comparison. Rooted cuttings, seedlings and transplants cold hardened at similar rates during early winter, achieving the same level of midwinter hardiness (LT(50) = -18 degrees C) in early January. However, rooted cuttings remained hardier later into the spring than did seedlings or transplants. Rooted cuttings exhibited deeper dormancy in early winter than seedlings or transplants but these differences disappeared after January. Root growth potentials of all three stock types remained above threshold values established for transplants throughout winter. Rooted cuttings had greater stem diameter, higher stem diameter to height ratio, and greater root weight than either seedlings or transplants. This may reflect lower growing densities for the rooted cuttings. Root/shoot ratios of rooted cuttings were greater than for seedlings and similar to those of transplants. Rooted cuttings also had deeper and coarser root systems, which probably reflects lack of wrenching at the nursery. 相似文献
16.
The growth of seven Picea sitchensis x Picea glauca hybridswas compared with the growth of two P. sitchensis provenancesand Picea glauca var.albertiana at two sites in northern Scotland.The sites were at Aultmore (an exposed, dry site with a mineralsoil) and Shin (a frosty, wet site with deep peat). They wereof the type considered more suited to Pinus contorta than P.sitchensis. At age 10, in 1984, most of the hybrids, at both sites, wereabout 10 and 20 per cent taller than P. sitchensis of Masset(Q.C.I.) and Ketchikan (Alaska) provenance, respectively. P.glauca var. albertiana grew very poorly, especially at Aultmore. At Aultmore, the frost hardiness of three of the tallest hybrids,the two P. sitchensis provenances, and P. glauca var.albertiana,was tested at about 3-weekly intervals throughout 1982 and 1983.Detached shoots were subjected to artificial frosts in a programmablechamber. P. glauca var. albertiana was frost susceptible atbudburst, but at all other times it was relatively very frosthardy (eg. to10°C in mid-August). Also, the hybridswere consistently more hardy than P. sitchensis of even Ketchikan(Alaska) provenance from July onwards. However, the hybridswere less frost hardy than P. sitchensis of Masset (Q.C.I.)provenance in early spring (they dehardened a week earlier inMarch-April) and their buds were equally as frost susceptibleat the time of budburst. In 1983, trees of P. glauca var. albertianaburst their buds about a week sooner than P. sitchensis. It was concluded that P. sitchensis x P. glauca hybrids canperform better than P. sitchensis at sites considered marginalfor P. sitchensis, and that their good performance may be partlyattributed to, or associated with, their greater summer andautumn frost hardiness. A programme of inter-specific hybridizationis being pursued. 相似文献
17.
Burial of seed in moist sand for several weeks during winter(pit stratification), and early sowing have been common practicesin forest nurseries for breaking dormancy of conifer seed. However,in the experiments reported here the emergence of Sitka sprucewas greatly reduced when seeds naturallly infected by Geniculodendronpyriforme, were moist-chilled for 6 weeks in a stratificationpit or in moist sand in a refrigerator at about 4° C. Reductionin emergence was less when seed was refrigerated without sand(naked), and negligible when the seed was treated with Thiramor Captan before chilling. Sowing 6 weeks early produced erraticresults, often giving fewer but larger seedlings. Moist-chillingof seed did not increase total emergence but normally made it12 weeks earlier so that the seedlings were larger byautumn. Addition of extra water to the seedbeds did not increaseemergence or growth, even in a dry season. 相似文献
18.
Beechnut Storage: A Review and Practical Interpretation of the Scientific Literature 总被引:1,自引:0,他引:1
Beechnuts are not easy to store or long-lived in storage. Theyexhibit storage characteristics between those of orthodoxand recalcitrant seeds. Superimposed upon thisis a pronounced dormancy which is overcome by between 4 and20 weeks of moist chilling. Pretreatment periods of 1520weeks occupy most of the time between collection in Octoberand sowing the following April; hence short-term storage isa combination of moist storage and pretreatment. Most evidencepoints to the best short-term storage/pretreatment conditionsbeing 05°C at 2832 per cent moisture content(m.c.) (fresh weight basis). Long-term storage requires a periodat low moisture content and the best conditions appear to be10 per cent m.c. and 10°C or below. The relative merits of overcoming seed dormancy before or afterdry storage are reviewed. There is no consensus about whichsequence is best but the former procedure is likely to be preferredby nursery managers because it allows storage of non-dormantnuts which can be withdrawn from store and sown immediately. Received 13 February 1990. 相似文献
19.
Recent ideas on silage and fuelforestry call for more information on the total harvestablewoody dry matter produced by hardwoods grown at very close spacingsin fertile soils and coppiced every few years. Yields of oven-driedstems and branches (S and B) are presented here for Populustrichocarpa Torr. and Gray, clone Fritzi Pauley.Plantings in Bedfordshire at 21 600 trees ha1 had a meanannual increment (M.A.I.SB) of 5.2 t ha1 y1 overfive years, and plantings in the Cambridgeshire fens at 1480trees ha1 produced 4.8 t ha1 y1 over sixyears. Fan-shaped spacing experiments, established in Midlothianby inserting cuttings through black polythene into nursery soilwith added fertilizers, gave 4.6 t ha1 y1 at theend of the first year and about 7 t ha1 y1 oneyear after coppicing, but only with over 250 000 stems ha1producing closed canopies with leaf area indices of about 4.Similar spacing experiments planted without fertilizer on farmlandin Gloucestershire, Suffolk, Argyll and Midlothian gave averageM.A.I.SB values of 6.57.0 t ha1 y1 afterthree years with over 25 000 trees ha1 and similar valuesafter five years with over 10 000 trees ha1. Peak currentannual increments (C.A.I.SB) averaged 1012 t ha1y1. The maximum M.A.I.SB, attained in Gloucestershire,was 10.0 t ha1 y1 at age 5 with over 20 000 treesha1, with maximum C.A.I.SB values of about 14 t ha1y1 at age 4; M.A.I.SB values of about 11.5 t ha1y1 were anticipated at this site by age 68. Equivalentstem volumes are given. As expected, trees subjected to competitionaccumulated greater proportions of their woody biomass in stemsrather than branches. Biomass yields of fully-stocked young hardwood stands are independentof planting density. In Britain, M.A.I.SB values of 68t ha1 y1 can be obtained over 1 or 25 years byplanting 250 000 or 2000 trees ha1, using vigorous Populusspp, Salix spp or Nothofagus procera on good sites. Advantages and problems of silage forestry arediscussed, and it is considered that hardwood fuel coppicescould not meet more than about 2% of national energy needs. The reciprocals of individual tree weights were linearly relatedto planting density. 相似文献
20.
The roots of second-rotation Sitka spruce (Picea sitchensis(Bong.) Carr.) planted on peaty gley sites are restricted tothe old litter (LFH) layer and are dependent on its decompositionfor availability of nutrients. A series of these sites of increasingage from felling were sampled to estimate changes in the nutrientcapital of the LFH horizon over time at Kielder Forest, Northumberland.Previous stand histories were reconstructed from stump data.Geographical, climatic, soil and mensurational data suggestedthat the use of a time series was justified. Nutrient capital in the LFH horizon generally declined overa 5 year period after clearfelling from approximately 997, 51and 83 kg ha1 to 676, 30 and 31 kg ha1 of N, Pand K, respectively. However, N concentration increased overa 5 year period from 11 mg g1 to 14 mg g1, P concentrationremained constant at about 0.6 mg g1, and K concentrationdecreased from 1.0 mg g1 to 0.7 mg g1. Nutrientconcentrations and contents of the LFH horizon were higher underthe brash (slash) swathes that resulted from the use of organizedfelling techniques than under clear strips devoid of brash. The patterned input of nutrient capital in brash as a resultof organized felling was also determined. Brash containing 219,20 and 71 kg ha1 of N, Pand K, respectively, was systematicallydistributed at a rate of 491 ha1 over 66 per cent ofthe site after harvesting. The needles and small branch fractionscontained 71 per cent of the N and 80 per cent of the P andK present in the brash. 相似文献