Different xylogenesis responses to atmospheric water demand contribute to species coexistence in a mixed pine–oak forest

Seasonal patterns of wood formation(xylogenesis) remain understudied in mixed pine-oak forests despite their contribution to tree coexistence through temporal niche complementarity.Xylogenesis was assessed in three pine species(Pinus cembroides,Pinus leiophylla,Pinus engelmannii) and one oak(Quercus grisea) coexisting in a semi-arid Mexican forest.The main xylogenesis phases(production of cambium cells,radial enlargement,cell-wall thickening and maturation) were related to climate data consideri...     

Commercial thinning that maintained species diversity of a mixed black spruce–jack pine stand enhanced productivity

Commercial thinning enables forest managers to meet timber production objectives. Thinning reduces tree density to alleviate competition for resources and favour growth of selected tree species. However, in doing so, thinning can homogenize the composition of mixed-species forests and raise biodiversity issues. There is increasing evidence that species richness can lead to higher productivity through a complementarity effect. Hence, thinning that would maintain species diversity of mixed-species forests could enhance stand productivity and help forest managers to reconcile timber production objectives and biodiversity issues. The objective of this study was to compare post-thinning stand production, experimentally over 10 years, in mixed and monospecific stands of black spruce (Picea mariana [Mill.] B.S.P.) and jack pine (Pinus banksiana Lamb.). The post-thinning stand production curve of the mixed stand converged toward that of the unthinned mixed stand while the production curves of the thinned and unthinned monospecific stands remained parallel. The convergent productivity of the mixed stand could be explained by a positive interaction between effects of thinning and niche complementarity. We propose that thinning that maintains species diversity of mixed stands could help forest managers who are implementing ecosystem management to reconcile timber production objectives with biodiversity issues.     

Response of selected water chemical quality parameters to slight thinning in a mature oak–beech forest ecosystem under sub-humid climate conditions

The objective of this study was to determine the effect of 18% thinning on streamflow nutrient flux from a mature oak–beech forest ecosystem by paired watershed approach. Two experimental watersheds including control (W-I) and treatment (W-IV) watersheds were used in the study. The experimental watersheds were monitored about 6 years from 2006 to 2011 for the calibration period and 4 years from 2012 to 2015 for the treatment period. The forest in the treatment watershed was thinned between October and December in 2011, and the forest in the control watershed was left untreated. Water grab samples were collected from the streams in the watersheds on weekly basis during both the calibration and treatment periods and analyzed for calcium (Ca²⁺), magnesium (Mg²⁺), Kjeldahl nitrogen (KN), sodium (Na⁺), potassium (K⁺), iron (Fe³⁺), aluminum (Al³⁺), ammonium nitrogen (NH₄ ⁺-N), and sulfate (SO₄ ²⁻). The simple linear regression equations were developed between mean monthly nutrient fluxes of two watersheds in the calibration period with significantly high correlation coefficients, and they were used to estimate nutrient fluxes from the treatment watershed during the treatment period as if thinning had not been applied. The changes in the monthly nutrient fluxes were estimated as the differences between measured and values calculated with the linear regression equations. Results showed that removal of 18% standing timber volume did not significantly change nutrient exports except for KN and Na⁺ from the treatment watershed.

     

Multipurpose conversion management of Scots pine towards mixed oak–birch stands—A long-term simulation approach

The forest growth model 4C was used to investigate how conversion management of a Scots pine (Pinus sylvestris L.) stand towards a mixed oak–birch stand would affect stand structural development – and hence biodiversity and productivity – in the long term. For this purpose the 4C model was parameterised for natural regeneration of light demanding species and extended for management of multi-layered stands. A series of structural indicators was selected to describe key factors of forest biodiversity at the stand scale. Two consecutive aspects of Scots pine conversion were tested: (1) the choice of conversion strategy between thinning and gap creation and (2) the choice of conversion regime in terms of cutting cycle, thinning type and pine tree retention. Three simulated conversion strategies aim at the gradual removal of the pine canopy but differ in the spatial organisation of pine cuttings and hence result in different light conditions for regeneration. Only the directed gap creation strategy was able to maintain and increase birch admixture to the stand and to approach natural stand structural development. Simulation of 12 conversion regimes for the directed gap creation strategy indicated that thinning type (from above or from below), pine tree retention at final felling (50% of the standing volume or none) and cutting cycle (6, 9 or 12 years) all significantly influence stand structural development. These effects were clearest for oak development. Birch occurred in a few mixed clusters, but tended to disappear when longer cutting cycles were used. Based on a multi-criteria analysis we conclude that the optimal conversion regime – in which both stand productivity and biodiversity objectives can be combined – implies thinning from above, pine tree retention, and cutting cycles of 6 years. The conceptual validity of the model as well as the applicability of the results are discussed.     

Analysis of forest structural complexity using airborne LiDAR data and aerial photography in a mixed conifer–broadleaf forest in northern Japan

Determining forest structural complexity,i.e.,a measure of the number of different attributes of a forest and the relative abundance of each attribute,is important for forest management and conservation.In this study,we examined the structural complexity of mixed conifer–broadleaf forests by integrating multiple forest structural attributes derived from airborne Li DAR data and aerial photography.We sampled 76 plots from an unmanaged mixed conifer–broadleaf forest reserve in northern Japan.Plot-level metrics were computed for all plots using both field and remote sensing data to assess their ability to capture the vertical and horizontal variations of forest structure.A multivariate set of forest structural attributes that included three Li DAR metrics(95 th percentile canopy height,canopy density and surface area ratio) and one image metric(proportion of broadleaf cover),was used to classify forest structure into structural complexity classes.Our results revealed significant correlation between field and remote sensing metrics,indicating that these two sets of measurements captured similar patterns of structure in mixed conifer–broadleaf forests.Further,cluster analysis identified six forest structural complexity classes includingtwo low-complexity classes and four high-complexity classes that were distributed in different elevation ranges.In this study,we could reliably analyze the structural complexity of mixed conifer–broadleaf forests using a simple and easy to calculate set of forest structural attributes derived from airborne Li DAR data and high-resolution aerial photography.This study provides a good example of the use of airborne Li DAR data sets for wider purposes in forest ecology as well as in forest management.     

Functional traits explain growth–mortality trade-offs in a mixed broadleaf-conifer forest in northeastern China

The trade-offs between tree growth and mortality are important indicators of the life-history strategies of species and have been demonstrated to be maintained by inherent fitness differences among species. There is a growing consensus that functional traits which inflect the ecological performances of individuals may be correlated with growth–mortality trade-offs within communities. In this study, the aforementioned trade-offs were detected and their relationships with the functional traits were examined, based on data collected in a typical temperate forested area in northeastern China. Principal components analyses were performed for growth–mortality trade-off relationships. The factor scores on the first principal component were used to represent the species’ positions at an axis of life-history variations among different species. Correlation and multiple regression analyses were performed for the purpose of detecting the relationships between functional traits and the growth–mortality trade-off. The results showed that there were significant trade-offs between the tree growth and mortality rates in the examined plot. Among the six functional traits which were considered, the H_max was found to have the strongest effects on the demographic rates and trade-offs. The observed negative effects on the tree growth and mortality rates suggested that the large species often grow more slowly and living longer, while species at small adult size grow fast but die easily. It was also observed that the specific leaf area and the leaf carbon content level were also very important to the growth and mortality trade-offs. The traits considered in this study explained 68% of interspecific variations in the species positions in the growth–mortality trade-off. In addition, wood density was found to show no effect on the trade-off. This was likely due to the fact that the density of the wood may influence the demographic rates more strongly during the early life stages of the trees. The results of discrimination analysis further confirmed the explanations abilities of the functional traits on the variations of life-history strategies among the species. Due to the fact that the relationships between the functional traits and the trade-offs were found to be relatively variable, it will be necessary to further consider other traits which are more closely and mechanistically linked to individual performances in future investigations, such as leaf life span, leaf tissue density, photosynthetic capacity, and dark respiration rates.

     

Wood density of three oak species in Tennessee, 1940–45 and 1970–75

Three oak species from East Tennessee were core-sampled to determine whether the density of wood produced in a pre-pollution period (1940–45) was different from that of a pollution period (1970–75) using gamma-densitometry. Component densities of average earlywood, average latewood, minimum earlywood, maximum latewood and average periodic density for Quercus alba L. and Quercus prinus L. were significantly greater for 1940–45 than for 1970–75. Differences between the two time periods for mean earlywood (P = 0.05), maximum latewood and mean periodic densities were found for Q. velutina.     

Fuel loads,snag abundance,and snag recruitment in an unmanaged Jeffrey pine–mixed conifer forest in Northwestern Mexico

Management of downed woody fuels and snags (standing dead trees) is receiving increasing attention because of their ecosystem values and effects on potential fire behavior. Research has correlated the abundance of many wildlife species with snags and downed woody material but very little information exists of the abundance and arrangement of these forest structures, particularly in unmanaged forests. Conifer forests in northwestern Mexico have not experienced systematic fire suppression or harvesting making them unique in western North America. In 1998, average snag density in Jeffrey pine–mixed conifer forests in the Sierra San Pedro Martir (SSPM) National Park was 3.95 snags/ha but 35% of inventoried plots had no snags. In 2002, average snag density significantly increased to 5.10 snags/ha after a multiple-year drought. Average surface and ground fuel loads were 15.8 and 8.7 t/ha, respectively. High variability characterized all snag and fuel attributes measured in this forest. This high amount of variation is probably the result of the relatively intact frequent surface fire regime and because no harvesting has occurred in the sampled area. The patchy distribution of snags observed argues against the application of uniform targets for snag retention across similar forested landscapes. An improvement in management guidelines would be to manage for snag density and large fuels over moderate spatial scales (hundreds of hectares) instead of on a per hectare basis. Forest fragmentation and diverse ownerships in many western United States forests complicates this recommendation. Conservation of the forests in the SSPM is critical because it is the last landscape-scale, old-growth mixed conifer forest in western North America with a relatively intact frequent fire regime.     

Spatial distribution of late-successional coniferous species regeneration following disturbance in southwestern Québec boreal forest

Remnant tree stands left intact following a disturbance constitute the sole seed banks available for regeneration of coniferous species that neither bear serotinous cones nor reproduce vegetatively. The success of regeneration of tree species on these disturbed sites is, therefore, dependent on the distance from potential seed sources. The regeneration of balsam fir (Abies balsamea (L.) Mill.), white spruce (Picea glauca (Moench) Voss) and white cedar (Thuja occidentalis L.) was studied at two sites in Québec’s southwestern boreal forest in order to quantify the influence of remnant stands on spatial distribution of regeneration. The first site is located in an area that burned in 1944 while the second site is located in an area that was clear-cut in the mid-1980s. Canonical correspondence analyses were used to determine the respective contributions of environmental data and spatial variables to the pattern of spatial distribution of regeneration. The results reveal that distance from a remnant stand is the most important variable in explaining spatial distribution of regeneration when compared to environmental variables such as soil type, drainage, slope and altitude. The plots of regeneration density against distance from a remnant stand for both the burned site and logged site show that regeneration density decreases abruptly with distance from a remnant stand. Furthermore, spatial autocorrelation analyses (Moran’s I) indicate that even small remnant zones can significantly influence the pattern of spatial distribution of regeneration for the three species studied. The results presented here suggest that where preestablished regeneration is not abundant enough, alternative silvicultural systems such as strip clear-cutting or seed-tree systems could be used instead of cuts with protection of regeneration and soils (CPRS).     

Estimating the age–diameter relationship of oak species in Switzerland using nonlinear mixed-effects models

Tree growth plays a key role in forest dynamics, yet little attention has been paid to quantifying tree age–diameter relationships. Predicting diameter growth of oaks is especially important due to their role in nature conservation and adaptive forest management under climate change. Thus, we (1) identified environmental variables that shape age–diameter relationships of oaks and (2) quantified the accuracy of predictions based on these variables. We determined the age–diameter relationship of 243 oaks (Quercus spp.) growing in Switzerland by using tree-ring samples. Nonlinear mixed-effects models based on a modified Chapman-Richards equation were fitted with environmental variables included as covariates. The fixed effects elevation, slope and water-holding capacity were most important in shaping the age–diameter relationships. Lower elevations, steeper slopes, north-facing aspects, higher water-holding capacities and moister summers resulted in larger maximum diameters. For 75 % of the oaks, age–diameter relationships predicted by the fixed effects matched fairly well the observations (root mean square error between predictions and observations <6 cm); the inclusion of random effects reduced root mean square errors for 86 % of the trees. These results suggest that water runoff plays a key role for the age–diameter relationships, accompanied by limiting temperature effects at higher elevations. The fixed effects covered variability in site quality, whereas the random effects included tree-specific deviations from expected age–diameter relationships, potentially due to neighbourhood effects such as stand density and competition.     

Spatial pattern analysis of forest trees based on the vectorial markOA

Analysis of spatial patterns to describe the spatial correlation between a tree location and marks(i.e., structural variables), can reveal stand history, population dynamics, competition and symbiosis. However, most studies of spatial patterns have concentrated on tree location and tree sizes rather than on crown asymmetry especially with direct analysis among marks characterizing facilitation and competition among of trees, and thus cannot reveal the cause of the distributions of tree locations...     

Mangrove structure,litter and macroalgal productivity in a northern‐most forest of Florida

Rhizophora mangle L. dominated 10 overwash islands within Tampa Bay forming the northernmost mangrove forests on the west coast of Florida. The mean number of trees and basal area were 5040 trees ha^-1 and 20.5m²ha^-1, respectively. Basal areas ranged from 1.1 (Avicennia germinans (L.) Stern), to 2.6m²ha^-1 (Laguncularia racemosa Gaertner), to 16.8 (R. mangle). Cockroach Bay mangroves are small (5.8–7.0m tall) versus coastal forests of south Florida and the Caribbean. Total litter production for a 12 month period was maximal in September for fringing (7.4gdwtm^-2d^-1) and interior (8.7gdwtm^-2d^-1) areas with the two zones not being significantly different. Average litter fall (3.1gdwtm^-2d^-1) was similar to more tropical Caribbean mangals. Leaves accounted for 68 of the litter and reproductive material for 6, being similar to tropical riverine and overwash marine angiosperm communities on the Mexican coast. Although mangrove forests in Tampa Bay are small in stature and experience cold damage and occasional frosts, their litter fall is similar in biomass to that of more tropical mangals. However, their reproductive output is low based on litter fall versus more tropical mangals suggesting that the northern extension is less than optimum. Macroalgal diversity was low (10 species) as compared to Caribbean mangals. Macroalgal turf (0.8gCm^-2d^-1) and epiphyte communities of A.germinans pneumatophores (2.7gCm^-2d^-1) have productivity levels that are equal to or greater than those of Caribbean mangals.     

Biodiversity of arbuscular mycorrhizal fungi in different trees of madhupur forest, Bangladesh

Roots and rhizosphere soils of Acacia auriculiformis A. Cunn. ex Benth., A. mangium Wild., Artocarpus heterophyUus Lamk. C., Dalbergia sissoo Roxb. ex A. P. D., Eucalyptus camaldulensis Dehnn., Hevea brasiliensis （Wild. ex Juss） Muell. Arg., Swietenia macrophylla King. and Tectona grandis L. were collected from different locations of Madhupur forest area to study the biodiversity of Arbuscular Mycorrhizal （AM） fungal colonization and spore population. All the plants showed AM colonization. Out of eight selected plants, mycelial colonization was lowest in the roots of A. heterophyllus （22%） and the highest was in the roots of H. brasiliensis （78%）. Mycelial intensity was observed poor （25%-77%） and moderate （23%-57%） in all plants species and abundant （11%-40%） was in most of the plant species. Vesicular colonization was observed in five plant species. The lowest was recorded in E. camaldulensis （4%） and the highest was in H. brasiliensis （21%）. Poor （24%-56%）, moderate （16%-100%） and abundant （11%-40%） type of vesicular intensity were observed. Arbuscular colonization was observed in three plants. The highest was in A. mangium （72%） and the lowest was in S. macrophylla （17%）. Arbuscular intensity was recorded as poor （12%-44%）, moderate （22%-100%） and abundant （4%-47%）. The highest AM fungal spore population was in A. auriculiformis （714） and the lowest was in D. sissoo （102）. Five AM fungal genera were recorded. Glomus was found to be dominant. A few spores remained unidentified. Significant correlation was observed between percent coloniza- tion and spore population. The results of the present study indicate the occurrence of AM fungi and the mycotrophism of the plants of Madhupur forest area and the applicability of AM technology in the forest management of Madhupur forest.     

The tree species matrix,influence on the level of herbivore browsing in mixed forest stands in southwest Sweden†

Herbivory pressure on a forest stand and each plant individual may be affected by the spatial distribution of conspecific and allospecific plants on the site; whether the plants are standing in solitary or groupwise settings; or by the differences in the preference of plants in relation to each other. The study was conducted in southwest Sweden, where 120 plots (1508?m²) were visited and 1280 individual woody plants evaluated and measured. We tested the hypothesis that preferred woody plants can protect unpreferred conspecific and allospecific woody species against herbivory in a system with one dominant, generalist herbivore, the fallow deer (Dama L.), and that the effectiveness of this protection varies depending on the relative preference of the neighboring conspecific and allospecific woody species placed in conspecific groups or standing solitary. Our results support the hypothesis that preferred woody plants can protect unpreferred conspecific and allospecific woody species against herbivory in the case of Picea abies dominating stands.     

Dynamics of dominant tree species in a forest ecotone on the northern slop of Changbai Mountain

The competition and dynamics of dominant trees species in the forest ecotone between the broad-leaved/Korean pine （Pinus koraiensis） mixed forest and the spruce-fir forest （also known as dark conifer forest） in Changbai Mountain, Jilin Province in Northeast China were studied by using Lotka-Volterra model, based on the data from twenty-eight sample plots with area of 20 mx90 m for each one. Results showed that under natural condition, differentiation of communities followed two directions： one would be Spruce （Picea jezoensis and few P. koraiensis） and fir （Abies nephrolepis） co-dominant conifer forest, and at the equilibrium fir was absolutely preponderant （77.1% of relative dominance （RD））; the other would be the conifer and broad-leaved mixed forest, and at equilibrium, the broad-leaved tree species was 50% of RD in the broad-leaved/Korean pine mixed forest and 66% of RD in the broad-leaved and spruce-fir mixed forest. The study demonstrated that both broad-leaved/Korean pine mixed forest and dark conifer forest were climax community, the ecotone had transitional characteristics, and the diversification of the forest communities suggested that the direction of succession was affected by local habitat.     

Allelopathic effect of larch leaching on ash growth in mixed forest

lntroductionConcerningthemechanismoftheinteractionbe-tWeentreespeciesinthelarch(LafisgmelinI)-ash(PraxiPusmandShuFbe)mixedforest,wehavestudiedtheaIIelopathiceff6ctsoftheIarchvolatilematter,litterandrootsecretion.onthegroWthofash.TheobtainedresultsshowedclearIythatthelarchvoIatiIematt6rplayedaninhibitivepart,andboththeIarchIitt6randrootsecretionpIayedanobviousfystimuIatingpaFtontheashgroWth(Wu2oooa,2ooob,2oooc).Thispapermakesastudyofanotherallelo-pathicway(Ieaching)betWeenlarchandash.T…     

Not seeing the forest for the trees? The environmental effectiveness of forest certification in Sweden

Forest certification can be conceived as one of many rapidly growing non-state market driven (NSMD) modes of governance. The environmental effectiveness of forest certification is oftentimes evaluated by indicators such as stringency of standards, degree of participation by key stakeholders, certified area, etc. In political science, forest certification as an NSMD governance arrangement is usually evaluated in terms of the quality of the decision-making procedures (input legitimacy) rather than for its problem solving capacity, i.e. its environmental performance or effectiveness. We conceptualize environmental effectiveness as a function of a standard's environmental stringency and the area covered by the standard, the latter dependent on the degree of social acceptance. Accordingly, the environmental effectiveness of different certification schemes ought to be evaluated taking both the standard stringency and the area certified into account. The forest certification process in Sweden illustrates how forestry history and regional differences affect the development, acceptance and adoption of different certification schemes. Industrial and Northern forestry owners favour the NGO led Forest Stewardship Council (FSC) standards whereas Southern small-scale private forest owners preferred to develop an alternative scheme the Programme for Endorsement of Forest Certification (PEFC). We demonstrate that there is a bifurcated geographical coverage of the two certification schemes along a north–south divide coupled with a similarity in standard stringency and a high degree of acceptance in their different areas of dominance. Both forest certification schemes display a similar degree of environmental effectiveness — but in different parts of the country and for different types of ownership.     

Effect of cleaning methods in mixed pine–deciduous stands on moose damage to Scots pines in southern Finland

Abstract

The effect of different cleaning treatments on moose (Alces alces L.) damage to Scots pines (Pinus sylvestris L.) was examined using a generalized linear mixed model (GLMM) with a binomial response (logit-link function). The treatments were the following: no cleaning, point-cleaning of broadleaves (mainly birch, Betula pendula Roth and B. pubescens Ehrh.) within a radius of 1 m from the pine, point-cleaning delayed 2 years, topping of competing broadleaves (main stem cut at a height of 1.3 m) and total cleaning of broadleaves. A randomized complete block design with three replicates was established in three planted pine stands (mean height of pines 1.5 m). Measurements taken four growing seasons later (mean height of pines 3.5 m) showed that all cleaning treatments reduced moose damage, although the difference was not significant (p=0.078) in the case of topping. The proportion of damaged pines was predicted as a function of the characteristics of pines and broadleaves in two untreated, planted pine stands with a height of 3 m. Within these untreated stands the number of damaged pines increased in plots with a higher number of broadleaves and taller broadleaves. In conclusion, the results show that on relatively fertile growing sites of pine cleaning treatments reduced moose damage on pine. It is recommended that cleaning is performed before broadleaved trees occur as overgrowth above pines.