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1.
Fire suppression over the last century has increased conifer expansion and dominance in aspen-conifer forests, which appears to be a driving force behind aspen decline in some areas. The primary objective of this study was to examine how increasing conifer dominance affects aspen regeneration vigor following the return of fire. The influence of physiographic features and herbivory on aspen regeneration vigor were also examined. The study was conducted in the Sanford fire complex located in the Dixie National Forest in southern Utah, USA, where more than 31,000 hectares burned in the summer of 2002. Seven years after the burn (at 66 locations) we measured aspen regeneration density and height as response variables and former stand composition and density (the burned trees were still standing), soil characteristics, slope, aspect and presence or absence of herbivory as independent variables. Aspen regeneration (root suckering) densities ranged from <500 to 228,000 stems/hectare with an average of 37,000 stems/hectare. Post-fire aspen regeneration density was most strongly correlated with pre-fire stand successional status (as measured by stand composition and species abundance), with percent conifer abundance (R2 = −0.55) and overstory aspen density (R2 = + 0.50) being the most important. Average aspen suckering densities ranged from approximately 60,000 stems/hectare in what were relatively pure aspen stands (>90% aspen) to less than 5000 stems/hectare in stands where conifer abundance was greater than 90%. Soil C, N, and P showed positive correlations (R2 = 0.07 to 0.17) with aspen regeneration vigor, while soil texture had a relatively weak influence on sucker regeneration. Aspen regeneration densities were 15% lower on north facing aspects compared to east, west and south facing aspects with slope steepness showing no correlation with regeneration vigor. Regeneration density was significantly lower (8%) at sites with evidence of herbivory versus sites where herbivory was absent. Overall, the aspen regeneration response in the Sanford fire complex was strong despite high wildlife densities, which may be related to disturbance size. Where the maintenance of aspen is desired in the landscape we recommend promoting fire when the percentage of overstory conifer stems is greater than 80% or overstory aspen density is less than 200 overstory stems/hectare.  相似文献   

2.
Season of harvest has often been suggested as a driver for the erratic success of aspen (Populus tremuloides) sucker regeneration, partially due to root carbohydrate reserves and soil conditions at the time of harvest. A field experiment in western Manitoba, Canada, assessed root suckering and root carbohydrates of aspen in response to season of harvest and machine traffic. Six sites (120 m × 120 m) were selected within two large mature aspen stands slated for summer harvest. Plots (50 m × 50 m) were hand-felled (without machine traffic) in mid-summer, late summer, winter, and one plot was left uncut as a control. Season of cut with no traffic had no effect on sucker density, height or leaf dry mass per sucker. During the dormant season, root starch reserves were highest in the winter cut plots, however, just prior to suckering, this difference in carbohydrate reserves among the three seasons of harvest disappeared and by the end of the first growing season root reserves in all three seasons of cut had recovered to near control levels. Adjacent plots that were conventionally harvested in the summer and impacted by logging traffic had similar sucker densities but had 19% less height growth of suckers and 29% less leaf dry mass per sucker compared to suckers in plots harvested at the same time without traffic. After one growing season, root carbohydrate levels were similar whether or not machine traffic was used; however, the reduction in leaf dry mass in plots with machine traffic could have negative implications for carbohydrate accumulation and growth. The study suggests that the phenological state of the mature aspen plays a very small role in aspen regeneration and that harvesting practices and site conditions are likely the main drivers of aspen regeneration success.  相似文献   

3.
Natural regeneration of Abies sachalinensis on soil was studied in a natural sub-boreal forest managed by single tree selection cutting located within the Hokkaido Tokyo University Forest. First, seedlings of A. sachalinensis on skid trails, soil mounds, soil pits, and undisturbed soil were counted, and their areas were measured. Most seedlings were found on skid trails. Seedling densities of A. sachalinensis on sites with soil disturbance, with the exception of soil pits, were significantly greater than that on undisturbed soil. Second, densities of Sasa senanensis, heights of S. senanensis, and seedling densities of A. sachalinensis on skid trails and undisturbed soil in both a closed-canopy stand and in a canopy gap were compared. Seedling density of A. sachalinensis on skid trails in the closed-canopy stand was significantly greater than that in the canopy gap. Average density and average height of S. senanensis, which is known to interfere with regeneration of many woody species, were significantly greater on skid trails and undisturbed soil in the canopy gap than those in the closed-canopy stand. It could be concluded from this study that both crown closure and soil disturbance are essential for natural regeneration of A. sachalinensis on soil in sub-boreal forests.  相似文献   

4.
We inventoried plant regeneration and soil compaction along mule trails to evaluate damage to forest stands and regeneration follow-ing mule hauling before and after operations in Kheyrud Forest in the...  相似文献   

5.
One of the main threats to the sustainability of community forestry in the Selva Maya is insufficient regeneration of commercial tree species. We evaluated the regeneration status of 22 commercial tree species in a managed semideciduous tropical rain forest in Southern Mexico. The study was carried out in six harvesting areas along a 16-year chronosequence. In each area, 10 transects (1000 m2) were established and all trees >50 cm height and <10 cm diameter were recorded. We evaluated the relationships between seedling and sapling abundance, and canopy cover and disturbance condition (closed forest, canopy gap, log landing, skid road, primary road and secondary road). The area occupied by closed forest canopy increased with age of harvesting area (65–91% of sampled area), while the area occupied by canopy gaps decreased (22–9%). Log landings occupied less than 1% of the sampled area. The predominant canopy cover was 75–80% in all harvesting areas, even in the most recently harvested areas. The highest densities of seedlings and saplings, of both shade tolerant and intolerant species, were found in log landing and skid trails, followed by secondary roads. Even Simarouba glauca, a shade tolerant species, displayed higher densities in sites with ≤65% of canopy cover. Our results support previous findings and indicate that the levels of disturbance caused by existing harvesting procedures may be inadequate to promote sufficient regeneration of not only light demanding desirable species but also for some of the evaluated shade tolerant species of commercial interest. Seedling and sapling densities exhibited by Swietenia macrophylla, for example, are insufficient to support current harvesting rates. The application of a spatial mixed system with patch-cuts of different sizes, a consequence of group felling, could be applied to provide the necessary conditions for the regeneration of the main commercial species.  相似文献   

6.
In the southern Rocky Mountains, current mountain pine beetle (Dendroctonus ponderosae Hopkins) outbreaks and associated harvesting have set millions of hectares of lodgepole pine (Pinus contorta var. latifolia Engelm. ex Wats.) forest onto new stand development trajectories. Information about immediate, post-disturbance tree regeneration will provide insight on dynamics of future stand composition and structure. We compared tree regeneration in eight paired harvested and untreated lodgepole pine stands in the Fraser Experimental Forest that experienced more than 70% overstory mortality due to beetles. New seedlings colonized both harvested and untreated stands in the first years after the beetle outbreak. In harvested areas the density of new seedlings, predominantly lodgepole pine and aspen, was four times higher than in untreated stands. Annual height growth of pine and fir advance regeneration (e.g., trees established prior to the onset of the outbreak) has doubled following overstory mortality in untreated stands. Growth simulations based on our regeneration data suggest that stand basal area and stem density will return to pre-beetle levels in untreated and harvested stands within 80-105 years. Furthermore, lodgepole pine will remain the dominant species in harvested stands over the next century, but subalpine fir will become the most abundant species in untreated areas. Owing to terrain, economic and administrative limitations, active management will treat a small fraction (<15%) of the forests killed by pine beetle. Our findings suggest that the long-term consequences of the outbreak will be most dramatic in untreated forests where the shift in tree species composition will influence timber and water production, wildfire behavior, wildlife habitat and other forest attributes.  相似文献   

7.
Overstory mortality, understory tree recruitment, and vegetation development were assessed in trembling aspen (Populus tremuloides Michx.) stands following two recent episodes of forest tent caterpillar defoliation (Malacosoma disstria Hbn.) in northeastern Ontario. The results suggest that poplar (aspen and balsam poplar (Populus balsamifera L.)) mortality increased with consecutive years of insect defoliation occurring from the mid-1980s to mid-2000s and the proportion of poplars in the overstory, but decreased with improved pre-defoliation tree vigour (DBH increment). The first outbreak, which lasted from the mid-1980s to early 1990s, was more severe in terms of insect defoliation and contributed more to poplar mortality and decline. The decline began in the late 1990s and peaked in early 2000s. Poplar regeneration and understory shrubs responded rapidly to foliage loss to insect defoliation and mortality of overstory poplars. The regenerated poplars were able to maintain their growth under developing shrubs and residual overstory canopy and numbers were sufficient to compensate for the poplar trees lost to insect infestation. The defoliation-induced overstory decline will accelerate the transition of aspen stands to conifer dominance through enhanced conifer recruitment and growth, and reduced hardwood overstory in aspen-dominated stands, while hardwood dominance will persist in pure aspen stands. From a timber supply perspective, the decline caused by forest tent caterpillar defoliation could delay the availability of aspen stands for harvesting by 40–50 years.  相似文献   

8.
Growth in height, diameter, and volume was measured on 9- to 18-year-old ponderosa pine (Pinus ponderosa) and 10- to 13-year-old lodgepole pine (Pinus contorta) trees growing on or near compacted skid trails in the Yakima Indian Reservation in south-central Washington. Soil bulk density of the 0- to 30.5-cm deep layer was measured with a single-probe nuclear densimeter on two sides of each sample tree and in adjacent undisturbed soil. On three ponderos pine sites logged 23 years before the study, average bulk density on skid trails was 15% greater than on adjacent undisturbed soil. On a lodgepole pine site logged 14 years before the study, soil on skid trails averaged 28% greater bulk density than undisturbed soil.Total growth of ponderosa pine and the last 5 years of growth were significantly related (P = 0.07) by regression analysis to age of trees, site index, basal area of the adjacent overstory, and the percentage of increase in soil bulk density. At the mean increase in soil bulk density, total height, diameter, and volume growth were reduced 5%, 8%, and 20%, respectively.Total growth of lodgepole pine and the last 5 years of height, diameter, and volume growth were significantly related to tree age and the percentage of soil organic matter. Increase in soil bulk density was not significantly related to growth of this species.  相似文献   

9.
We examined the extent of soil disturbance associated with bulldozer yarding and the regrowth of woody vegetation on bulldozer paths (skid trails) in selectively logged dipterocarp forest. In an area logged in 1993, using conventional, i.e., uncontrolled, harvesting methods, about 17% of the area was covered by roads and skid trails. In contrast, in a 450-ha experimental area where reduced-impact logging guidelines were implemented, 6% of the area was similarly disturbed. Skid trails in the reduced-impact logging areas were less severely disturbed than those in conventional logging areas; the proportion of skid trails with subsoil disturbance was less than half that in conventional logging areas. Four years after logging, woody plant recovery on skid trails was greater in areas logged by reduced-impact than by conventional methods. Skid trails where topsoil had been bladed off had less woody vegetation than skid trails with intact topsoil. In a chronosequence of logging areas (3, 6, and 18 years after logging), species richness and stem densities of woody plants (>1 m tall, <5 cm dbh) were lower on skid trail tracks than on skid trail edges or in adjacent forest. Both richness and density increased with time since logging, but even 18 years after logging, abandoned skid trails were impoverished in small woody stems compared with adjacent forest. Minimizing soil and stand disturbance during logging appears to allow a more rapid recovery of vegetation on bulldozed soils, but the long-term fate of trees growing on compacted soils remains uncertain.  相似文献   

10.
Sudden aspen decline (SAD), affecting Populus tremuloides, was first observed in Colorado in 2004. By 2008 it affected at least 220,000 ha, an estimated 17% of the aspen cover type in the state. In southwestern Colorado, we examined site and stand features in paired healthy and damaged plots to assess the effects of SAD on aspen and to identify factors associated with decline. Root mortality increased significantly with recent crown loss. Consequently, density of regeneration did not increase as the overstory deteriorated, and regeneration that originated since 2002 decreased significantly in stands with moderate to severe SAD. However, mortality of regeneration did not increase with that of the overstory. Remeasurement of a subset of plots after 1–2 yrs showed significant increases in severity. Contrary to expectations, overstory age and diameter were not related to SAD severity as measured by recent crown loss or mortality. Severity of SAD was inversely, but weakly, related to basal area, stem slenderness, and site index, and positively related to upper slope positions. This is consistent with moisture stress as an underlying factor. To test the role of climate as an inciting factor for SAD, a landscape-scale climate model was used to compare moisture status of declining and healthy aspen at the height of the warm drought in water year 2002. Polygons identified as damaged aspen in the 2008 aerial survey had greater moisture deficits than healthy aspen in the 2002 water year. SAD has led to loss of aspen cover in some stands, and is occurring in areas where early loss of aspen due to climate change has been predicted. Further warm, dry growing seasons will likely lead to recurrence of SAD.  相似文献   

11.
Limiting surface soil disturbance caused by forest harvesting machines is an important task and is influenced by the selection of efficient and reliable predictors of such disturbance. Our objective was to determine whether soil moisture content affects soil load bearing capacity and the formation of ruts. Measurements were conducted in six forest stands where various machines operated. We measured the formation of ruts along skid trails in connection with varying soil moisture content. Soil moisture content was determined through the gravimetric sampling method. Our results showed that severe(rut depth16–25 cm) to very severe disturbance(rut depth [26 cm)occurred in forest stands where the instantaneous soil moisture exceeded its plasticity limits defined through Atterberg limits. Atterberg limits of soil plasticity ranged from 26 to 32 % in individual stands. Regression and correlation analysis confirmed a moderately strong relationship(R = 0.52; p \ 0.05) between soil moisture content and average rut depth. This confirmed that soil moisture is a suitable and effective predictor of soil disturbance.  相似文献   

12.
Excessive slash loading could pose a problem for the regeneration of the serotinous lodgpole pine especially in forests at higher elevation where soil temperature is limiting. In the past, these forests have commonly been harvested using full-tree harvesting where trees are processed at roadside; however, recently cut-to-length harvesting has become a more frequent harvesting method. In cut-to-length harvesting the harvested trees are processed in the block, as a result slash accumulation is much higher on these cutblocks. In an experimental field trial, the cone distribution, natural lodgepole pine regeneration, and the growth and establishment of planted lodgepole pine were evaluated in response to slash load, drag scarification, and mounding after cut-to-length harvesting of high elevation lodgepole pine stands in the Rocky Mountains. Twelve sites were established, each contained six plots which were randomly assigned to six treatment combinations of two slash removal (slash and slash removed) and three mechanical soil preparation treatments (no soil preparation, drag scarifying, and mounding). The slash removal reduced slash volume by more than 50% but also reduced the number of lodgepole pine cones available for regeneration by over 33%. However, soil mechanical treatments offset this effect as fewer cones were necessary to achieve high natural pine regeneration densities. Drag scarification of plots resulted in 12 times the number of pine seedlings compared to the non-prepared plots. Although slash removal did not have an effect on the number of naturally regenerated lodgepole pine seedlings, it had a positive effect on their growth performance. Conversely, planted pine seedlings had lower mortality and better growth in soils that had been mechanically prepared and had the slash removed; however, the growth effects became only apparent 4 years after planting. While slash removal and mechanical soil preparation did increase soil temperatures; the slash removal treatment had a more transient effect on soil temperatures than soil preparation. Differences in soil temperature decreased over time which appeared to be mostly driven by a warming of the soils in the plots with no soil preparation, likely a result of the decomposition of the finer slash and feathermosses. Overall, it appears that surface disturbance on these high elevation sites had a far greater effect on lodgepole pine regeneration and growth than the increased accumulation of slash as a result of cut-to-length harvesting.  相似文献   

13.
14.
Quantifying tree damage and mortality caused by single-tree selection harvesting is critical to understanding postharvest forest dynamics and management. In this study, we quantified the effects of tree size and species and the distance from residual trees to felled trees and skid trails on damage to residual trees and mortality in mixed coniferous–broadleaved stands of Hokkaido, northern Japan. Among the 4,961 trees that we studied, 373 (7.5 %) were damaged, and 148 of these trees (3.0 %) died during or immediately after logging. Hierarchical Bayesian modeling showed that the risk of damage to residual trees increased with increasing size of the felled trees and with increasing proximity to felled trees and skid trails. Smaller residual trees had the greatest risk of damage. Species differed in their susceptibility to damage; Abies sachalinensis (Fr. Schm.) Masters and Picea jezoensis (Sieb. et Zucc.) Carr. were the most susceptible species in our sample plots. Smaller damaged trees had the highest risk of mortality. The damaged trees that did not die at the time of logging had a higher risk of postharvest mortality than undamaged trees. Our results indicate that, to minimize logging-induced damage and mortality: (1) the spatial arrangement of skid trails should be optimized and fixed, (2) the risk of skidding damage should be considered before marking the trees to be harvested and choosing the felling direction, and (3) logging operators should be instructed to avoid damaging small trees, especially those of the species that are most susceptible to damage.  相似文献   

15.
Trembling aspen (Populus tremuloides Michx.) density and growth were assessed 9–12 years after stand establishment to determine whether mechanical site preparation (MSP) affects crop tree quality. Study sites were either treated with disc trenching or ripper ploughing and planted with white spruce (Picea glauca (Moench) Voss) seedlings immediately after harvest (treated) or were undisturbed since harvest (control). Stands were surveyed during the summer of 2002 with standard regeneration survey plots. Results show that aspen stem density and height were lower in MSP-treated areas relative to untreated areas. Diameter growth rates were unaffected by treatment, however the percentage of stem discolouration was higher in untreated control stands compared to site prepared areas. The results of this study indicate that there are no long-term benefits to carry out MSP for aspen promotion. However, as MSP does not appear to seriously harm the aspen crop, we suggest that this treatment can still be used on sites where aspen densities may be low without treatment (e.g., sites with extremely low soil temperatures, poor soil aeration, or vigorous competitive vegetation) or where a mixture of aspen and planted spruce are desired.  相似文献   

16.
In this study, we quantified the effects of local neighbourhood competition, light availability, and proximity to skid trails on the growth of sugar maple (Acer saccharum Marsh.) trees following selection harvest. We hypothesized that growth would increase with decreasing competition and increasing light availability, but that proximity to skid trails would negatively affect growth. A total of 300 sugar maples were sampled ∼10 years after selection harvesting in 18 stands in Témiscamingue (Québec, Canada). Detailed tree and skid trail maps were obtained in one 0.4 ha plot per stand. Square-root transformed radial growth data were fitted to a linear mixed model that included tree diameter, crown position, a neighbourhood competition index, light availability (estimated using the SORTIE light model), and distance to the nearest skid trail as explanatory variables. We considered various distance-dependent or -independent indices based on neighbourhood radii ranging from 6 to 12 m. The competition index that provided the best fit to the data was a distance-dependent index computed in a 6 m search radius, but a distance-independent version of the competition index provided an almost equivalent fit to data. Models corresponding to all combinations of main effects were fit to data using maximum likelihood, and weighted averages of parameter estimates were obtained using multimodel inference. All predictors had an influence on growth, with the exception of light. Radial growth decreased with increasing tree diameter, level of competition and proximity to skid trails, and varied among crown positions with trees in suppressed and intermediate positions having lower growth rates than codominants and dominants. Our results indicate that in selection managed stands, the radial growth of sugar maple trees depends on competition from close (≤6 m) conspecific neighbours, and is still affected by proximity to skid trails ∼10 years after harvesting. Such results underscore the importance of minimizing the extent of skid trail networks by careful pre-harvest planning of trail layout. We also conclude that the impact of heterogeneity among individual-tree neighbourhoods, such as those resulting from alternative spatial patterns of harvest, can usefully be integrated into models of post-harvest tree growth.  相似文献   

17.
Soil compaction due to the use of heavy machinery for timber harvesting has become a widespread problem in forestry. However, only few studies deal with the regeneration of compacted forest soils. In the present study, we examined the potential of accelerating soil regeneration by planting black alder trees (Alnus glutinosa (L.) Gaertn.) in skid lane tracks. In 2003, seedlings were planted into the rut beds of severely compacted skid lanes in two Swiss forest sites. In addition, some of the ruts were filled with compost. In 2009 and 2010, we assessed the success of these measures by analysing physical parameters of soil structure (bulk density, total and coarse porosity and air permeability), root densities and tree growth. Tree growth was exceptionally strong on the skid lanes. Total and coarse soil porosity and air permeability showed significant increase in planted skid lanes as compared to untreated control subplots, approaching values found for untrafficked soil in the immediate vicinity. All soil physical parameters were closely correlated to root mass density. Compost application enhanced tree growth and soil structure regeneration on one site, but had a retarding effect on the other site. Planting black alders has great potential as an environmentally friendly measure to accelerate the structural regeneration of compacted forest soils in temperate humid climates.  相似文献   

18.
Catastrophic wind events, including tornado, hurricane, and linear winds, are significant disturbances in temperate forested wetlands. Information is lacking on how post-disturbance salvage logging may impact short and long-term objectives in conservation areas where natural stands are typically managed passively. Woody regeneration and herbaceous cover were assessed for three years in a bottomland hardwood forest across a gradient of damage from an F4 tornado, with and without subsequent salvage logging. Soil disturbance intensity and recovery associated with salvage logging within wind-disturbed sites were also assessed. Woody stem density and proportion of potential overstory species (species with the potential to occupy a position in the canopy) increased as a function of wind disturbance intensity. Stem density, proportion of overstory trees, or species diversity did not differ between wind + salvage and wind-disturbed-only plots. Significant dissimilarity occurred among soil disturbance classes within salvaged sites. By the third growing season, vegetation in soil disturbance classes in wind + salvage areas was converging toward undisturbed conditions and bottomland hardwood forest recovery was underway in all vegetation disturbance types and soil disturbance classes. Post-tornado salvage logging, applied judiciously, may contribute to microsite and vegetation diversity.  相似文献   

19.

One of the basic requirements for sustainable forest management in mixed broadleaved stands is to provide suitable conditions for natural regeneration of trees. These conditions include, but are not limited to, the frequency, quality and the composition of seedling species. Compacted soil layers caused by forest machinery traffic are the most common problem affecting seedling establishment and growth after skidding operations. In this study, we evaluated the frequency, quality, diversity of seedling species and physicochemical properties of soil on 10-, 20- and 30-year-old abandoned skid trails. Further comparison to the values of the mentioned above parameters in control areas allowed for the evaluation in terms of natural recovery processes, under varying traffic intensity and the longitudinal slope of the skid trails in a mixed broadleaved forest. Results showed that there is a significant positive trend of recovery for soil physicochemical properties and ecological characteristics (density and quality) of seedling growing on the skid trails. The time required to recover soil properties and ecological attributes of seedlings increased with increasing traffic intensity and slope of the skid trail. Our results showed that it takes 20 and 30 years to fully recover the chemical and physical properties of the soil, respectively. On the 30-year-old skid trails, density and species diversity indices were fully recovered, but the quality of the seedlings was not restored to the control conditions. The results suggest the importance of a proper planning of the skid trail network, avoiding in particular skid trails with a slope gradient higher than 20%.

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20.
Establishing white spruce (Picea glauca (Moench) Voss) by planting it under established aspen (Populus tremuloides Michx.), stands has substantial potential as a technique for regenerating boreal mixedwood stands. The presence of an aspen overstory serves to ameliorate frost and winter injury problems and suppresses understory vegetation that may compete with white spruce. In this study we examine the growth of white spruce during the first 10 years after being planted underneath a 39 year-old stand of trembling aspen following thinning and fertilization. Results indicate successful establishment and reasonable growth rates of white spruce planted under thinned and unthinned aspen stands, even with aspen basal area of 51 m2 ha−1. Thinning of overstory aspen to 1000 or 2000 stems ha−1 did not increase light reaching seedlings, but did result in improvements in light above the shrub layer and in diameter and height growth of the underplanted seedlings. However, these increases in growth of underplanted spruce may not justify the expense of thinnings. Fertilization of these stands prior to planting had no effect on spruce growth. Growth of spruce underplanted at this site near Fort Nelson was similar to that at two other stands near Dawson Creek, B.C.  相似文献   

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