Utilization of non-timber forest products in humid tropics: Implications for management and livelihood

Community dependence on forest resources for diverse needs has high implications for long term management of forests. Sustainable extraction of non-timber forest products (NTFPs) is considered best feasible strategy for forest conservation in biodiversity rich areas. This paper examines the heterogeneity of NTFPs use by tribal communities in northeast India, a global hot spot by examining diversity of NTFPs at-large, its consumption pattern, and contribution to rural income and forest revenue. A total of 343 NTFPs recorded used for diverse purposes by tribal communities. When species choice grouped as per use category, utilization for medicinal (163 species), edible fruits (75 species) and vegetables (65 species) purposes was reasonably high. Community dependence on forest resources was 100% for firewood and house construction material. 76 plant species were sold in three major local markets while an additional 22 species traded at commercial scale mainly outside the state. NTFPs contributed 19-32% of total household income for different tribal communities, which was significant. Illicium griffithii, Rubia cordifolia, Oroxylum indicum, Swertia chirayita, Litsea sebifera, Taxus wallichiana, Valeriana jatamansii, Thalictrum foliolosum, Picrorhiza kurrooa, Everniastrum cirrhatum, Cordyceps sinensis, Aconitum fletcherianum, Nardostachys jatamansi, Picrorhiza kurrooa, Gymnadenia orchidis, Calamus, Quercus and Pinus roxbughii were important commercial species. NTFPs also generated substantial revenue to the State government, though it is falling year after year, which is alarming. To meet community livelihoods, income and forest revenue from NTFPs, it desires a thorough management plan and policy guidelines for these resources from all line departments. The knowledge on diversity, its consumption pattern, and contribution to rural income and forest revenue may enable planners to accurately plan sustainable management of NTFP resources and community development in near future.     

Assessing the relationships between stand development and understory vegetation using a 420-year chronosequence

The large-scale conversion of old forests to tree plantations has made it increasingly important to understand how understory vegetation responds to such landscape changes. For instance, in some forest types a reduction in understory richness and cover is thought to result from the development of canopy closure in plantations, although there is a paucity of empirical data demonstrating this relationship. We used a 420-year forest chronosequence as a case study to assess the relationship between stand age, tree canopy cover and understory vascular plant richness and composition in the Siskiyou Mountains of Oregon. The chronosequence consisted of six young managed (age 7–44) and nine older unmanaged (age 90–427) stands. All stands were similar in underlying geology, slope, elevation, and aspect. We found a non-linear relationship between stand age and richness, in which richness was highest in the youngest stands, reached a low in mid-aged stands (∼55 years), then increased in the oldest stands. We also found that percent tree canopy cover was correlated with total understory cover, richness, diversity, and species composition. In general, young stands were characterized by high shrub and graminoid cover and old stands were characterized by an abundant herb layer. Our work suggests that a major component of our study landscape is currently entering the forest stage (canopy closure) characterized by low levels of vascular plant species richness and cover. We use our results to discuss the potential effects of future forest management on understory plants.     

近自然化改造对杉木人工林物种多样性的影响

在广西凭祥市热林中心青山实验场设置杉木人工林近自然化改造样地,进行3种间伐强度处理(75%、55%、35%),均匀套种4种阔叶树。改造5年后,研究不同间伐强度近自然经营对群落组成、物种多样性和林分生长状况的影响。结果表明:近自然化改造5年后,乔、灌、草3层的物种数明显增加,灌草层的优势种有所变化,但各间伐处理的优势物种大致相同;各间伐处理的灌木层物种多样性指数无显著差异;改造后各处理草本层物种多样性指数均高于未改造的纯林,各处理草本层的Simpon指数和均匀度指数均显著高于改造前的纯林。杉木胸径随间伐强度的增大而增加且差异显著,但各处理间树高无显著差异;套种的4种阔叶树的胸径、树高随间伐强度的增大而增加,大叶栎、红椎的胸径、树高生长量在不同间伐处理林分中有显著差异;高间伐强度改造的杉木人工林适合套种阳性树种大叶栎、米老排和中性树种红椎,中等间伐强度改造的杉木人工林适合套种耐阴性树种润楠。     

The compatibility of timber and non-timber forest product extraction and management

We discuss the extent of compatibility of timber and non-timber forest product (NTFP) extraction in two neotropical localities from biophysical, social, and institutional perspectives: the community concession forests of the Petén, Guatemala, and extractivist communities in northern Bolivia. In both localities, timber is harvested via reduced-impact logging (RIL) practices. In Guatemala, the main NTFP extracted is foliage from the understory xate palm (Chamaedorea spp.); in northern Bolivia, the fruits of the Brazil nut tree (Bertholletia excelsa). The Guatemalan case suggests a relatively high degree of compatibility due to low timber harvesting intensities, coupled with temporal, spatial, and social segregation of xate extraction in a well-defined land tenure system. In northern Bolivia, forest management regulations pay little attention to NTFPs and land tenure issues and related conflict complicate efforts to limit timber harvesting impacts on Brazil nut trees. The introduction of timber management plans overlain on customary property rights systems in which the extraction of Brazil nut is organized could undercut its management system. The two case studies suggest that specific legislative, education, and project interventions may help to promote the compatibility of timber and NTFP extraction and management. These include formal training of foresters on NTFP ecology and management while taking into account the perspectives of multiple stakeholders in the design of management plans. Tropical forest users, research managers, and policy makers will also better understand the need for integrated management of timber and NTFPs, if the trade-offs and potential economic benefits from NTFP extraction are clarified.     

Short-term effects of selection and clear cutting on the shrub and herb layer vegetation during the conversion of even-aged Norway spruce stands into mixed stands

The conversion of even-aged Norway spruce (Picea abies (L.) H. Karst.) stands into more site adapted mixed stands is the main silvicultural goal in Germany and many other European countries. The conversion will primarily be achieved with the help of selection cutting, removing single target diameter trees and creating small gaps. At sites highly susceptible to windthrow, however, small scale clear cutting (up to 1 ha) might be an appropriate alternative. Both selection and clear cutting will affect the understorey vegetation, an important component of forest biodiversity and ecosystem functioning. Thus, with the help of a BACI-experiment (before–after/control–impact), the short-term effects of selection and small scale clear cutting on the herb and shrub layer vegetation were studied at two sites in the Solling Hills (Lower Saxony, Germany). Both treatments led to an increase in herb and shrub layer species numbers, due to persistent and invading (mainly pioneer tree species and indicators of disturbance) species and to a shift in plant traits. Most responsible for the changes in vegetation was a higher light availability, and to a lesser degree soil disturbance. The number of annuals and ruderals, however, increased only immediately after clear cutting, whereas on selection cutting plots the numbers remained high. Gaining in number and coverage were competitive perennials (Epilobium angustifolium, Juncus effusus, Rubus idaeus) especially on the clear cuttings, fulfilling a nutrient retention function. The competitives did not deter tree species from regenerating in the clear cut plots, whereas on selection cutting plots the large expansion of R. idaeus appeared to be hindering tree colonisation. Suffering under clear cut conditions were predominantly shade-tolerant, closed-forest species, but no loss of these species could be detected. This indicates the potential for a fast recovery of these forest species, as soon as, pioneer trees expand.     

Diversity of non-timber forest products in Cachar District,Assam, India

We identified 67 and 21 NTFP-yielding plant and animal species, respectively, in a reserve forest in Cachar district of Assam. We recorded globally threatened species listed by IUCN as Critically Endangered （plant species Dipterocarpus turbinatus）, Vulnerable （plant species Aquillaria malaccensis）, Endangered （animal species Hoolock hoolock, Indotestudo elongata and Manis pentadactyla）, and Vulnerable （animal species Nilssonia hurum and Rusa unicolor）. The whole plant or animal and/or their various parts were used as food or medicine, in house construction, magico-religious activities and others. While some NTFPs were harvested throughout the year, others were harvested seasonally. A comprehensive NTFP policy, along with scientific measures for regen- eration, restoration and augmentation of NTFP-yielding plants and animals, would help in addressing the conflicting demands of conservation and livelihood in the forests of this area.     

Research in Non-Timber Forest Products

Abstract

Summary Non-timber forest products (NTFPs) have emerged as a complex set of issues reflecting changes in society and how natural resources are regarded. These issues range from the sustainability of forest management practices to the relationship of diverse cultures and communities to public lands and their resources. Research and its relationship to this set of issues is a relatively unknown aspect of NTFPs.

This paper reports on early NTFP research by scientists in the USDA Forest Service's Pacific Northwest Research Station. It characterizes efforts over approximately five years and identifies their key elements. It also discusses the role research has and could play in addressing the problems and questions associated with NTFPs and sustainable forestry .     

The Non-Timber Forest Products Sector in Nepal: Emerging Policy Issues in Plant Conservation and Utilization for Sustainable Development

The non-timber forest products (NTFPs) sector in Nepal is being promoted with the concept of sustainable development as articulated by the Convention on Biological Diversity (CBD). Traditionally, people have been dependent on NTFPs for subsistence; however, international trade is growing. To promote this sector, Nepal adopted the 2004 Herbs and NTFP Development Policy. Our goal is to assess the effectiveness of the policy along with the other forestry and natural resource policies in Nepal concerning the conservation and use of NTFPs. We conducted open-ended semi-structured interviews with 28 key informants in summer 2006 in Nepal and collected relevant documents and publications. Qualitative analyses of data yielded important issues that should be addressed to promote the sector as envisioned by the Government. The most important is the need for inventory and research on NTFP species in widespread use. There are some issues regarding species banned under various other National Acts; the lack of marketing information and capacity building; administrative barriers; royalty rates; and illegal harvesting. The most important immediate need is the development of an NTFP Strategy and Action Plan. Our findings should help further implement the policy and promote the sector along the stated mission of conservation and sustainable development.     

Fuel and stand conditions in an isolated, unmanaged forest landscape in central Oregon

? Mature, unmanaged forests in western North America provide important reference conditions for managers, yet little is known about fuel composition and the factors that influence fuel accumulation in such stands. Our objectives were to characterize fuels in a passively managed landscape of dry forests in central Oregon and identify environmental factors influencing fuel accumulation.

? Ordination techniques and analysis of variance revealed no statistical differences in total fuel loads across a wide range of environmental conditions.

? Individual fuel size classes, however, did vary by stand location and composition. Interior stands had more 1- to 100-h fuels and snags than stands at or near the edge, stands dominated by ponderosa pine had fewer small-diameter fuels and snags, lodgepole pine stands had more 1-h fuels and snags, and white fir stands had more 10-h fuels, duff, and snag basal area.

? Tree species, density and age, and years since last fire were the most important environmental variables, explaining 23% of the variation in fuels.

? Our results present further evidence that fuel composition is highly variable at many spatial and temporal scales. They also provide useful baseline information for managers of fire-prone western forests.

     

Forest management conditioning ground ant community structure and composition in temperate conifer forests in the Pyrenees Mountains

The search for indicators to monitor management impact on biodiversity is a crucial question because management practices promote changes in community structure and composition of different animal groups. This study explores the effect of widely conducted management practices (forest logging and livestock) in Pinus uncinata forests in the Pyrenees range (NE Spain) on the structure and composition of ground ant communities compared to those of old-growth stands. Forest structure clearly differed in stands with different forest managements. These stands managed for different uses also showed marked differences in structure and composition of ground ant communities. There was a great dominance of a single species, Formica lugubris, which accounted for 99% of ants collected in pitfall traps. Rarefaction curves indicated that species richness was highest in old-growth stands and lowest in even-aged ones, with woodland pasture stands showing an intermediate value. Classification methods allowed us to identify two groups of species: six species related to old-growth plots and three species (including F. lugubris) associated to managed stands. Habitat structure played an important role in determining the structure of ant communities: forests with high tree density but low basal area were the most favourable forest type for F. lugubris, while the abundance of the remaining ant species was negatively affected by the abundance of F. lugubris and by tree cover.     

Options for Non-Timber Forest Product Management in the Condor Bioreserve,Ecuador

Abstract

This paper describes and evaluates the current role that non-timber forest products (NTFPs) and their users play on the conservation of the four protected areas-Reserva Ecológica Cayambe-Coca, Reserva Ecológica Antisana, Parque Nacional Cotopaxi, and Parque Nacional Sumaco-Napo Galeras-that comprise the Condor Bioreserve. The category non-timber forest product (also referred to as non-wood forest product) encompasses a myriad of resources that reside in forests and can be useful to people. Plants from which NTFPs are derived play essential roles in tropical forest ecosystem processes such as regeneration and nutrient cycling. In the Condor Bioreserve, certain species of such plants are commercially valuable while others are endangered. People are extracting these NTFPs from the Bioreserve, yet the magnitude and frequency of such extraction is unknown. This paper examines how the three groups of land users that have been identified by The Nature Conservancy-colonists, indigenous peoples, and hacienda owners-may utilize NTFPs. Understanding the role that these plants play in the lives of its collectors is imperative for designing effective strategies to alleviate pressure that extraction may place on the Condor Bioreserve ecosystems. Data on who harvests NTFPs, which plant parts they use, what quantities they harvest, and harvest frequency must be complemented by insight into how each group benefits from these activities. Moreover, an unbiased approach must examine how NTFP extraction may enhance certain ecosystems in addition to ways in which it may hurt Bioreserve resource ecosystems.     

模拟林冠损伤对藜蒴栲群落林下植被生物多样性的影响

以2008年冰雪灾害对森林生态系统的破坏为背景,通过人工去除林冠层保留树干模拟森林的机械损伤,研究粤北小坑流域藜蒴栲群落林下植被的变化.结果表明:(1)试验1.5年后,去除林冠层、林下添加枯枝落叶的处理,林下灌木、草本科、种及乔木幼苗种的数量显著增加(P＜0.05);去除林冠层且地表枯落物层保持不变的处理,林下草本科、种数量增加显著,乔木幼苗及灌木科、种差异不显著(P＞0.05);林下添加枯枝落叶,林冠层不做处理,林下植被各生活型物种数有所减少,但差异不显著.(2)去除林冠层后,一些阳生种如野桐、山乌桕、山苍子、红紫珠、蕨状苔草、广东蛇葡萄等大量入侵并占据优势地位,林下植被盖度显著提高.(3)去除林冠层在短期内可显著增加林下植被生物多样性,添加枯落物对林下植被生物多样性影响不显著.     

Population distribution and threats to sustainable management of selected non-timber forest products in tropical lowland rainforests of south western Nigeria

Uncontrolled harvesting of non-timber forest products (NTFPs) poses a serious risk of extermination to several of these species in Nigeria. Yet, there is a paucity of information on the distribution, population status and sustainable management of NTFPs in most of the tropical lowland rainforests. We, therefore, assessed the population, distribution and threats to sustainable management of NTFPs within the tropical lowland rainforests of Omo and Shasha Forest Reserves, south western Nigeria. Data were obtained through inventory surveys on five top priority species including: bush mango (Irvingia gabonensis (Aubry-Lecomte ex O’Rorke) Baill), African walnut (Tetracarpidium conophorum (Mull. Arg.) Hutch. & Dalziel syn. Plukenetia conophora), chew-stick (Massularia acuminata (G. Don) Bullock), fever bark (Annickia chlorantha Setten & P.J.Maas syn. Enantia chloranta) and bush pepper (Piper guineense Schumach. & Thonn.). Purposive and stratified random sampling techniques were used for the inventory. Each forest reserve was stratified into three, viz: less disturbed natural forest (for areas that have been rested for at least ten years), recently disturbed natural forest (for areas that have suffered one form of human perturba- tion or the other in the last five years), and plantation forest (for areas carrying forest plantation). Data were collected from eighteen 10m×500m belt transects located in the above strata. The species were gener- ally fewer in both plantation and recently disturbed natural forest than the less disturbed natural forest, suggesting that forest disturbances (habitat modification) for other uses may have an effect on the occurrence and densities of the NTFPs. Exceptions to this trend were found for P. guineense and T. conophorum, which were fairly common in both planta-tion and recently disturbed natural forest. Among three tree NTFP species (i.e. I. gabonensis, M. acuminata and A. chlorantha), only I. gabonensis showed a significant difference in overall DBH size classes for both reserves (t=-2.404; df =21; p=0.026). Three tree NTFP species in both reserves further showed differences from the regular patterns of distribution of trees. The fairly regular reverse J-shaped size class distribution observed for M. acuminata in the study sites, however, suggests a recuperating population. In general, destructive harvesting of species, logging operations, low population size, narrow distribution ranges and habitat degradation are the major threats to the population of NTFPs in the study area. The implications of our findings for sustainable management of NTFPs in the study area are discussed and recommendations are made for a feasible approach towards enhancing the status of the species.     

Early height development and species stratification across five climax series in the eastern Washington Cascade range

Reconstruction in twenty single-cohort stands across five climax series on the eastern slope of the Washington Cascade Range found a variety of species combinations and development patterns. Western larch (Larix occidentalis Nutt.) and lodgepole pine (Pinus contorta Dougl. ex Loud) were found to be very competitive species that usually occupied a dominant position in stands in the Abies grandis, Abies lasiocarpa, and Tsuga mertensiana climax series. Interior Douglas-fir (Pseudotsuga menziesii var. glauca (Mirb.) Franco) was found in all five climax series and, although its height growth was less than that of western larch or the lodgepole pine, it was usually found in the upper stratum. These results suggest that site classification based on climax potential should be used cautiously when applied to young stand management decisions regarding seral species. For example, the difference between an Abies grandis, and a Tsuga mertensiana climax series is quite large in terms of potential productivity but in either case western larch, if present, will likely dominate these stands.     

Effects of the overstory on the diversity of the herb and shrub layers of Anatolian black pine forests

Understory plants are important components of forests because they are responsible for the majority of the vascular plant diversity of forest ecosystems. The richness and composition of understory communities are closely related to the tree layer diversity, structure and composition. The aim of this study was to examine the understory diversity of Anatolian black pine (Pinus nigra Arnold subsp. pallasiana (Lamb.) Holmboe)-dominated forests on the Kazda?? Mountains of West Turkey. To describe the overstory structure and composition in a numerically and quantitatively well-defined manner, cumulative abundance profiles (CAPs) of the tree species were used. The resemblance of the sampling plots was classified into five stand types assessing the CAP through the Fuzzy C-Means clustering method. A permutational multivariate analysis of variance (PERMANOVA) was performed to test the variance of the community ecological distance between the five stand types, and the results showed significant differences in these clusters. Many shade-tolerant plants were associated with the mixed stands of Anatolian black pine–Kazda?? fir. The composition of the herb and shrub layer could not be explained by the environmental variables but by differences in the overstory structure of the stands. Pure or nearly pure Anatolian Black pine stands were more diverse than mixed oak–Anatolian black pine and Kazda?? fir–Anatolian black pine stands. However, although dense and young pure Anatolian black pine stands had the most diverse plant species in the shrub layer, they were ranked third in terms of the herb layer diversity. The Anatolian black pine–Kazda?? fir mixed stands had the lowest herb and shrub layer diversity. These results allow us to comprehend the relationship between the overstory structure and composition, and the understory diversity. Understanding this relationship is important for the conservation of understory plant diversity in the management of forest ecosystems.     

Impact of silviculture on dead wood and on the distribution and frequency of tree microhabitats in montane beech-fir forests of the Pyrenees

In forest ecosystems, the level of biodiversity is strongly linked to dead wood and tree microhabitats. To evaluate the influence of current forest management on the availability of dead wood and on the abundance and distribution of microhabitats, we studied the volume and diversity of dead wood objects and the distribution and frequency of cavities, dendrothelms, cracks, bark losses and sporophores of saproxylic fungi in montane beech-fir stands. We compared stands unmanaged for 50 or 100 years with continuously managed stands. A total of 1,204 live trees and 460 dead wood objects were observed. Total dead wood volume, snag volume and microhabitat diversity were lower in the managed stands, but the total number of microhabitats per ha was not significantly different between managed and unmanaged stands. Cavities were always the most frequent microhabitat and cracks the least frequent. Dendrothelm and bark loss were favored by management. Beech (Fagus sylvatica) carried many more microhabitats than silver fir (Abies alba), especially cavities, dendrothelms and bark losses. Fir very scarcely formed dendrothelms. Secondary tree species played an important role by providing cracks and bark losses. The proportion of microhabitat-bearing trees increased dramatically above circumference thresholds of 225 cm for beech and 215 cm for fir. Firs with a circumference of less than 135 cm did not carry microhabitats. In order to conserve microhabitat-providing trees and to increase the volume of dead wood in managed stands, we recommend conserving trees that finish their natural cycle over 10–20% of the surface area.     

Comparison of tree growth and undergrowth development in aerially seeded and planted Pinus tabulaeformis forests

Direct seeding is a less expensive practice than planting and has the potential to become a viable alternative to transplanting for afforestation and regeneration purposes. As an effective and a less costly regeneration method, aerial seeding has been applied with several tree species. As early as 1956, Chinese people engaged in aerial seeding and stands with a total of 2.97×10⁷ hm² have been developed up to 2004. Our study tested whether the growth of planted Chinese pine (Pinus tabulaeformis Carr.) seedlings and its undergrowth development in northwest aspects differ from that of aerially sown seedlings on the northern and northwestern aspects of slopes. In 2007, we collected data such as height, diameter at breast height (DBH), clear bole height and canopy widths of trees, abundance, coverage, and frequency of shrubs and herbs from 21-year-old planted Chinese pine stands on a northwestern aspect (PNW), aerially sown stands in a northwest aspect (ANW) and aerially sown stands in a northern aspect (AN). Results showed that the relation of crown area and mean DBH was best fitted by a double inverse model for the ANW and AN forests and by a quadratic model for the PNW forest. There was no difference in the growth between ANW and AN forests, while growth was significantly higher in the PNW forest than in the ANW and AN forests. That was consistent with the Sorenson diversity indices in the shrub and herb layers, indicating that there was a large number of the same species in both aerially seeded stands, although their locations were different. Both the number of species in the undergrowth and the Shannon-Wiener index in the shrub layer were higher in the PNW stands than in the ANW and AN stands. Dominant families for all three stands were Rosaceae and Compositae in the shrub and herb layer, respectively. The dominant species for all three stands was Spiraea pubescens in the shrub layer, while the dominant species was different from each other in the three stands. The discrepancy in diversity and composition of species in the herb layer show that herbs are sensitive to shrubs in the three forests. High mortality and skewed diameter distributions reflect severe competition and too high a density in the aerially seeded forests. Thus, aerial seeding is a viable and effective regeneration technique, but management practices, such as thinning, should be applied to these forests.     

Demand for non-timber forest products: Surveys of urban consumers and sellers in Switzerland

In recent years, worldwide socioeconomic, ecological, and cultural changes have led to a paradigm shift in forestry and forest management. Some years ago, most forest enterprises exclusively focused on wood production, whereas today, forest management is expected to be multifunctional and economically as well as socially and ecologically sustainable [Harrison, S., Herbohn, J., Niskanen, A., 2002. Non-industrial, smallholder, small-scale and family forestry: what's in a name? Small-scale Forest Economics. Management and Policy 1 (1), 1–11; Schmithuesen, F., 2004. Forest policy developments in changing societies — political trends and challenges to research. Towards the sustainable use of Europe's forests: Forest ecosystems and landscape research: Joensuu: European Forest Institute, Proc., vol. 49, pp. 87–99]. In this context, non-timber forest products (NTFPs) such as forest honey, mushrooms, or berries could gain importance as an additional source of income for the forestry sector. To assess the market potential of NTFPs in Switzerland, two corresponding surveys were conducted. A consumer survey (N = 897) conducted in six cities in Switzerland investigated the purchase of Swiss NTFPs, consumer preferences, and demand potentials. An additional survey of sellers of NTFPs (N = 105) looked at these aspects from a complementary perspective. The results showed that there is a high esteem and considerable demand potential for such products among urban consumers. Food products were mentioned most frequently. Both consumers and sellers considered quality and environmental friendliness the most important attributes of NTFPs. Structural and organizational innovations and considerable marketing efforts and sales promotion would be required to expand the market for Swiss NTFP, however. Our results indicate that certification of the products' origin, environmental performance, and quality could support such efforts and that recreational and ecological forest services have a synergetic potential for the marketing of NTFPs.