Tree and livestock productivity in relation to tree planting configuration in a silvopastoral system in North Wales, UK

Silvopastoral systems in Europe offer the potential of introducing environmental benefits while at the same time increasing the diversity of farm outputs. The establishment of new silvopastoral systems by planting young trees into existing pasture was investigated at a site in North Wales, UK. Two tree species, sycamore (Acer pseudo-platanus L.) and red alder(Alnus rubra Bong.), were planted into pasture at a range of densities and planting arrangements. Growth of trees planted in farm woodland blocks (2500 stems ha^–1) was compared with the growth of trees planted at 400 stems ha^–1 in clumps and dispersed throughout the plot and at 100 stems ha^–1 (dispersed). Over the first six years after planting, alder trees were significantly taller and larger in diameter than sycamore. Sycamore trees planted at close spacing in farm woodland or clumped arrangements were significantly larger in diameter than widely spaced sycamore at 100 and 400 stems ha^–1. Livestock productivity was unaffected by the presence of trees during the six-year establishment phase of the system. The planting of trees in a clumped pattern appears to combine silvicultural benefits to tree growth with agricultural benefits of maintaining livestock production while trees are established.This revised version was published online in November 2005 with corrections to the Cover Date.     

The establishment phase of a silvopastoral national network experiment in the UK

Silvopastoral agroforestry can satisfy some objectives required of European land-use systems: reduced agricultural production, increased timber production, increased product diversity and environmental enhancement. A national network experiment was set up on four sites, each representative of a UK grassland farming area, with three replicates of common treatments: sycamore (Acer pseudoplatanus L.) at two silvopastoral planting densities [100 (S100) and 400 (S400) stems ha^–1] protected by tree shelters and an agricultural control (ACONT) all with grazing sheep; a woodland control (WCONT, 2500 trees ha^–1) without grazing. Common management protocols were applied and common measurements recorded. Results are provided for the first six years. There were no significant differences between S100, S400 and ACONT in agricultural productivity, though there were significant differences between sites (P < 0.001). There were no significant differences in tree survival between the silvopastoral treatments and WCONT (mean 92.5% ± 0.74) but there was a difference between S100 and S400 (90.8 vs 94.7%: P < 0.001). There were significant differences between the sites (range 86.5–96.2%: P < 0.001) and between the first three years, when replacement of dead trees took place, (82.5, 95.1 and 96.9% for years 1, 2 and 3 respectively: P < 0.001). There were significant differences in the total height of the trees in years two to four between WCONT, S100 and S400 (113.5, 154.1 and 194.5 cm respectively in year four: P < 0.001). However, by year six WCONT and S100 were similar (180.7 ± 17.31 cm) while S400 were taller (219.0 ± 22.80 cm: P < 0.05). It is concluded that tree shelters maintained silvopastoral tree survival at the level of conventional woodland. Tree height extension was compromised on S100 where a higher animal:tree ratio resulted in greater animal activity and soil compaction around trees compared to S400. The site with poorly-drained soil proved to be unsuitable for sycamore-based silvopastoral agroforestry.This revised version was published online in November 2005 with corrections to the Cover Date.     

Effect of shading by baobab (Adansonia digitata) and néré (Parkia biglobosa) on yields of millet (Pennisetum glaucum) and taro (Colocasia esculenta) in parkland systems in Burkina Faso, West Africa

An experiment was conducted in Nobéré, Burkina Faso to assess the effect of shade of two indigenous fruit trees, Adansonia digitata (Baobab) and Parkia biglobosa (Néré), on a shade-tolerant crop called taro (Colocasia esculenta) in comparison with millet (Pennisetum glaucum) a shade-intolerant crop. Photosynthetically active radiation (PAR) and performance of crops under trees and in the open field were assessed during three cropping seasons. Millet performed better under baobab (806.1?±?121.48?kg?ha^?1) compared to the control plot (595.8?±?83.43?kg?ha^?1) and néré (320.2?±?59.91?kg?ha^?1). In contrast, the yield of taro was higher under néré (4124.0?±?469.05?kg?ha^?1) compared to the control plot (2336.9?±?617.04?kg?ha^?1) and baobab (2738.3?±?595.61?kg?ha^?1). There was a strong relationship between the amount of PAR intercepted by trees and crop yields under trees. As PAR decreased the yield of millet decreased whereas the yield of taro increased. Hence, it was concluded that parkland productivity could be enhanced by cropping taro under néré where light reduction was 83, 56 and 18% in zones A, B and C, respectively. An efficient association of baobab with crops could be the production of taro in zone A and millet in zones B and C where PAR reduction was 62, 38 and 15%, respectively.     

Ailanthus triphysa at different density and fertiliser levels in Kerala, India: tree growth, light transmittance and understorey ginger yield

Ailanthus triphysa (Family – Simaroubaceae) growth is known to vary in response to different stocking and fertiliser levels. Understorey productivity related to these differences remain elusive, yet are important for optimising the combined production of tree and crop components. A split plot experiment to evaluate the effect of different stocking levels and fertiliser regimes on ailanthus growth, stand leaf area index (LAI) and understorey PAR (photosynthetically active radiation) transmittance was started at Vellanikkara, India in June 1991. Main plot treatments included four densities (3,333, 2,500, 1,660 and 1,111 trees ha⁻¹), replicated thrice. Four fertiliser levels (0:0:0, 50:25:25, 100:50:50 and 150:75:75 kg N:P₂O₅:K₂O ha⁻¹) formed the sub plot treatments. Ginger (Zingiber officinale) was planted as an understorey crop in May 1994 with contiguous treeless control plots. Soil nutrient availability before and after ginger was assessed. Higher densities stimulated ailanthus growth modestly, while fertiliser response of tree and ginger was inconsistent. PAR transmittance below the canopy was related to tree density, LAI and time of measurement. Midday PAR flux having low standard deviations is ideal for evaluating canopy effects on understorey light availability. Ginger in the interspaces exhibited better growth compared to sole crop. Highest rhizome yield was observed in the 2,500 trees ha⁻¹ stocking level, which is optimum for below five year-old ailanthus stands on good sites. It represents 52% mean daily PAR flux or 73% midday PAR flux. Ailanthus+ginger combinations improved the site nutrient capital when ginger was adequately fertilised, despite treeless controls having relatively higher initial soil nutrient availability. This revised version was published online in June 2006 with corrections to the Cover Date.     

Nitrogen-fixation dynamics in a cut-and-carry silvopastoral system in the subhumid conditions of Guadeloupe, French Antilles

This paper summarizes several studies on N recycling in a tropical silvopastoral system for assessing the ability of the system to increase soil fertility and insure sustainability. We analyzed the N₂ fixation pattern of the woody legume component (Gliricidia sepium), estimated the recycling rate of the fixed N in the soil, and measured N outputs in tree pruning and cut grass (Dichanthium aristatum). With this information, we estimated the N balance of the silvopastoral system at the plot scale. The studies were conducted in an 11-year-old silvopastoral plot established by planting G. sepium cuttings at 0.3 m × 2 m spacing in natural grassland. The plot was managed as a cut-and-carry system where all the tree pruning residues (every 2-4 months) and cut grass (every 40-50 days) were removed and animals were excluded. No N fertilizer was applied. Dinitrogen fixation, as estimated by the ¹⁵N natural abundance method, ranged from 60-90% of the total N in aboveground tree biomass depending on season. On average, 76% of the N exports from the plot in tree pruning (194 kg [N] ha^–1 yr^–1) originated from N₂ fixation. Grass production averaged 13 Mg ha^–1 yr^–1 and N export in cut grass was 195 kg [N] ha^–1 yr^–1. The total N fixed by G. sepium, as estimated from the tree and grass N exports and the increase in soil N content, was about 555 kg [N] ha^–1 yr^–1. Carbon sequestration averaged 1.9 Mg [C] ha^–1 yr^–1 and soil organic N in the 0-0.2 m layer increased at a rate of 166 kg [N] ha^–1 yr^–1, corresponding to 30% of N₂ fixation by the tree. Nitrogen released in nodule turnover (10 kg [N] ha^–1 yr^–1) and litter decomposition (40 kg [N] ha^–1 yr^–1) contributed slightly to this increase, and most of the recycled N came from the turnover or the activity of other below-ground tree biomass than nodules. This revised version was published online in June 2006 with corrections to the Cover Date.     

Competition for light between hedgerows and maize in an alley cropping system in Hawaii, USA

Successful agroforestry systems depend on minimizing tree-cropcompetition. In this study, field experiments and a simulation model were usedto distinguish between tree-crop competition for light and belowgroundcompetition in an alley cropping system. Maize (Zea maysL.) was harvested periodically in three treatments: between vertical barriers ofshade cloth, hedgerows of Flemingia macrophylla (Willd.)Merr., and sole maize. Radiation intercepted by the maize was calculated using asimulation model based on measured values for direct and diffuse light, hedgerowdimensions and leaf area, and solar trajectory. Radiation use efficiency wascalculated as biomass production per unit of intercepted radiation. Maizebiomass and yield in both the alley crop and the shade cloth treatment weregreatest in the center of the alleys. Grain yield between hedgerows was 3.5Mg ha⁻¹ (averaged across the alley), significantlyless than in the shade cloth (7.4 Mg ha⁻¹) or thesole maize (7.7 Mg ha⁻¹) treatments. Lightintercepted by the maize in the alley crop was about half that intercepted bythe maize in the sole crop. The shade cloth intercepted less light than thehedgerows because it did not have an appreciable width. Radiation use efficiencyin the three treatments was 0.75 g mol⁻¹ PAR anddid not differ significantly among treatments. Tree-crop competition wasoverwhelmingly for light. This revised version was published online in June 2006 with corrections to the Cover Date.     

Carbon storage and nitrogen cycling in silvopastoral systems on a sodic in northwestern India

Tree-based land-use systems could sequester carbon in soil and vegetation and improve nutrient cycling within the systems. The present investigation was aimed at analyzing the role of tree and grass species on biomass productivity, carbon sequestration and nitrogen cycling in silvopastoral systems in a highly sodic soil. The silvopastoral systems (located at Saraswati Reserved Forest, Kurukshetra, 29°4prime; to 30°15prime; N and 75°15prime; to 77°16prime; E) consisted of about six-year-old-tree species of Acacia nilotica, Dalbergia sissoo and Prosopis juliflora in the mainplots of a split-plot experiment with two species of grasses, Desmostachya bipinnata and Sporobolus marginatus, in the subplots. The total carbon storage in the trees + grass systems was 1.18 to 18.55 Mg C ha⁻¹ and carbon input in net primary production varied between 0.98 to 6.50 Mg C ha⁻¹ yr⁻¹. Carbon flux in net primary productivity increased significantly due to integration of Prosopis and Dalbergia with grasses. Compared to 'grass-only' systems, soil organic matter, biological productivity and carbon storage were greater in the silvopastoral systems. Of the total nitrogen uptake by the plants, 4 to 21 per cent was retained in the perennial tree components. Nitrogen cycling in the soil-plant system was found to be efficient. Thus, It is suggested that the silvopastoral systems, integrating trees and grasses hold promise as a strategy for improving highly sodic soils. This revised version was published online in June 2006 with corrections to the Cover Date.     

Foliar Responses of Understorey Abies lasiocarpa to Different Degrees of Release Cutting in a Betula papyrifera and Conifer Mixed Species Stand

Foliar responses of subalpine fir [Abies lasiocarpa (Hook.) Nutt.] to thinning were studied in a 35-yr-old mixed stand of paper birch (Betula papyrifera Marsh.) and conifers. The stand regenerated naturally after a wildfire with a canopy dominated by paper birch (average height 9.8 m) and an understorey dominated by subalpine fir (average height 1.6 m). The stand was thinned to four densities of birch: 0, 600 and 1200 stems ha-¹ and control (unthinned at 2300-6400 stems ha-¹) in the autumn of 1995. The understorey conifers, mainly subalpine fir, were thinned to 1200 stems ha-¹. The study used a completely randomized split-plot design. Three sample trees were systematically selected from each treatment replicate and each tree stratum (upper, intermediate and lower understorey). One-year-old and older age class needles were collected from one south-facing branch within the fifth whorl from the tree top. Thinning of paper birch significantly (p <0.001) increased leaf area and dry weight per 100 needles for intermediate and short trees except in the 0 birch treatment. Understorey subalpine fir trees in 600 stems ha¹ birch (T3) had the largest leaf area and leaf dry weight per 100 1-yr-old needles. Specific leaf area (SLA) decreased from unthinned (T1) to 0 birch (T4). Lower understorey trees had the largest SLA. One-year-old needles had significantly higher N, P and K concentrations in all the thinning treatments. These responses are consistent with the shade tolerance of subalpine fir. The results suggest that when managing a paper birch-conifers mixed-wood forest it may be of benefit to understorey conifers to leave a birch canopy as a nursing crop.     

Pasture production under different tree species and densities in an Atlantic silvopastoral system

We studied the effect of six tree species planted at six different densities on pasture production seven years after establishment. Annual and seasonal pasture production was studied every six months, over three years. Pasture production was lower under conifer trees (Pseudotsuga menziesii (Mirb.) Franco, Pinus pinaster Aiton, Pinus radiata D. Don) than under broadleaved trees (Betula alba L., Quercus rubra L. and Castanea sativa Mill.). Annual pasture production under Pseudotsuga menziesii and Pinus pinaster decreased progressively starting from 952 trees ha⁻¹, while decline in herbage production under Pinus radiata began to occur at 427 trees ha⁻¹. Tree density effect on pasture production was detected at 2,000 trees ha⁻¹ for all of the deciduous species studied. This effect on pasture production was more important in the first six months of the year (June sampling), while from June to December herbage production was less affected by tree density. The tree effect became more noticeable over time, with the last sampling showing the inverse relationship between tree density and herbage production most clearly. Seven years after tree establishment, pasture production was quite consistent under tree densities between 190 trees ha⁻¹ and 556 trees ha⁻¹ and declined remarkably from 556 trees ha⁻¹ to 2,500 trees ha⁻¹. The study also indicated that by the sixth growing season, annual pasture production under different tree species is inversely correlated with tree leaf area index.     

Radiation-use efficiency and dry matter partitioning of a young olive (Olea europaea) orchard

Radiation-use efficiency (RUE) relates biomass production to the photosynthetically active radiation (PAR) intercepted by a plant or crop. We determined RUE and biomass partitioning coefficients of young olive (Olea europaea) trees for use in a general growth model. In 1995, 1-year-old olive trees var. 'Picual' were planted at a density of either 0.5 or 2.0 trees m(-2) near Córdoba, Spain, at a site providing favorable growth conditions. During the experiment (1995-1997), both PAR interception by the canopy and plant area index (PAI) were measured with radiation sensors. Regular harvests were performed to determine leaf area and biomass accumulation in roots, wood (stem, branches and trunk) and leaves. Leaf, wood and root biomass partitioning coefficients were calculated. The leaf area partitioning coefficients were also estimated. Dry matter production was linearly related to cumulative intercepted PAR. Seasonal RUE, calculated as the slope of the regression of aboveground biomass and cumulative intercepted PAR, was 1.35 g (MJ PAR)(-1). Radiation-use efficiency appeared to respond to environmental conditions, but was independent of planting density and PAI. The young olive trees allocated 0.26 of their total biomass to roots. Partitioning of aboveground dry matter was 0.60 to wood and 0.37 to leaves. As competition increased, dry matter partitioning to wood increased to 0.70.     

Afforestation of savannah with cocoa agroforestry systems: a small-farmer innovation in central Cameroon

Cocoa cultivation is generally considered to foster deforestation. Contrary to this view, in the forest–savannah interface area in Cameroon, farmers have planted cocoa agroforestry systems on Imperata cylindrica grasslands, a soil-climate zone generally considered unsuitable for cocoa cultivation. We undertook a survey to understand the agricultural and ecological bases of this innovation. Age, cropping history and marketable cocoa yield were assessed in a sample of 157 cocoa plantations established on grasslands and 182 cocoa plantations established in gallery forests. In a sub-sample of 47 grassland cocoa plantations, we inventoried tree species associated with cocoa trees and measured soil organic matter levels. Marketable cocoa yields were similar for the two types of cocoa plantations, regardless of their age: 321?kg?ha^?1 in cocoa plantations on grasslands and 354?kg?ha^?1 in cocoa plantations in gallery forests. Two strategies were used by farmers to eliminate I. cylindrica prior to the establishment of cocoa plantations, i.e., cropping oil palms in dense stands and planting annual crops. Farmers then planted cocoa trees and fruit tree species, while preserving specific forest trees. The fruit tree and forest tree densities respectively averaged 223 and 68 trees?ha^?1 in plantations under 10?years old, and 44 and 27 trees?ha^?1 in plantations over 40?years old, whereas the cocoa tree density remained stable at 1,315 trees?ha^?1. The Shannon–Weaver index increased from 1.97 to 2.26 over the same period although the difference was not statistically significant. The soil organic matter level was 3.13?% in old cocoa plantations, as compared to 1.7?% in grasslands. In conclusion, our results show that the occupation of grasslands by cocoa agroforestry systems is both an important example of ecological intensification and a significant farmer innovation in the history of cocoa growing.     

Lime,sewage sludge and mineral fertilization in a silvopastoral system developed in very acid soils

In recent years, in the European Union, sewage sludge production has been increased as a result of EU policy (European directive 91/271/EEC). Organic matter and nutrient sewage sludge contents, principally nitrogen, indicate it can be used as fertilizer. The objective of the experiment was to compare the effect of no fertilization, three doses of sewage sludge, with or without liming, and the fertilization usually used in the region applied over a period of 3 years on pasture production and tree growth in a silvopastoral system. The experiment was conducted in the northwest of Spain. The soil was very acid (soil pH = 4.5) and had very low nutrient levels, especially P, that is related to site index. It was sown with a grass mixture (25 kg ha⁻¹ of Lolium perenne L. 10 kg ha⁻¹ of Dactylis glomerata L. and 4 kg ha⁻¹ of Trifolium repens L.) in Autumn 1997 under a plantation of 5-year-old Pinus radiata D. Don at a density of 1,667 trees ha⁻¹. Liming and sewage sludge fertilization increased soil pH and reduced saturated aluminium percentage in the interchange complex (Al/IC) in the soil, coming up the effect before with liming. Medium and high sewage sludge doses increased pasture production in the two first years. In a silvopastoral system, positive tree growth response to different fertilization treatments depended on tree age, initial soil fertility, soil pH, the relationship of competition with pasture production and previous liming application.     

Growth, herbivore distribution, and herbivore damage of timber trees in a tropical silvopastoral reforestation system

Context

The reforestation of degraded tropical pastures requires innovative tree planting systems that consider land user needs.

Aim

We established a silvopastoral reforestation system and assessed the effects of companion trees on the native timber tree Tabebuia rosea in Panama. Timber tree individuals were established in (1) solitary plantings (TSol) or in companion plantings with (2) Guazuma ulmifolia (TGua) or (3) the nitrogen-fixing Gliricidia sepium (TGli).

Methods

We quantified growth characteristics and herbivory of T. rosea, and analyzed leaf chemistry (including the stable isotopes ??¹⁵N and ??¹³C) and structure (leaf mass per area).

Results

Companion trees significantly affected stem diameter growth of T. rosea. Stem diameter growth was as high in TGli trees as in TSol trees but was reduced in TGua trees. Furthermore, TGua trees had higher leaf water content, and lower ??¹³C and lower leaf mass-to-area ratio than TGli trees, suggesting there were effects of shading by G. ulmifolia on T. rosea. Herbivory was high but not affected by planting regimes. Leafing phenology did not differ between planting regimes and G. sepium did not increase nitrogen content in T. rosea leaves.

Conclusion

Companion tree planting can support timber tree growth in silvopastoral reforestations, but adequate species selection is crucial for successful implementation of this planting system. Tree?Ctree interactions seem to be more relevant for timber tree growth than herbivory in the studied system.     

Wood production in a natural stand of Acacia karroo in Zimbabwe

Trees are important components of agricultural systems in the dry tropics of Africa. The assumption that exotic species are more productive than the indigenous woodland in these situations is seldom supported by quantitative data. In this study a new technique of recognising annual growth rings from the bands of marginal parenchyma has been used to model growth in a natural population of Acacia karroo Hayne, one of the most widespread and useful trees in the savanna woodlands of southern Africa. The study showed both basal area and sectional area at ankle height to be excellent predictors of total tree volume. Stands may yield from about 1 to 4 m³ ha⁻¹ year⁻¹ dependent upon spacing and site; individual trees produced up to 1 m³ of wood in 26 years. Maximum current annual increment was reached about 12 years after the trees approached breast height, thereafter declining until they became moribund about 8 years later. The economic rotation for wood production was close to the lifespan of the tree.     

Insights from full-rotation Nelder spacing trials with Eucalyptus in São Paulo,Brazil

The choice of spacing among trees for operational plantations is typically based on one or more experimental plantations that test for the response of tree and stand growth to a range of tree-to-tree distances. The most common design for spacing experiments entails rectangular plots that test different distances between rows, and between trees within rows, and with replication of plots covering one to several hectares within a single stand. Other designs may offer more information with simpler layouts, and we examined the insights that could be obtained from a Nelder (fan-shape) design, where spacing among trees varies with the radial distance from a central point. The response of Eucalyptus dunnii seed-origin trees to spacing was essentially similar between a classic plot design replicated in four blocks (tree spacing from 4.5 to 9.2 m² tree^?1, covering 1.4 ha of land), and a Nelder design (tree spacing from 2.1 to 44.0 m² tree^?1, covering 0.6 ha of land). The Nelder design showed slightly higher volume increment through 10 years of stand development than the block design (for the range of overlap in spacing), but the treatment effect of spacing was essentially identical between the designs at the level of both trees and stands. A second Nelder experiment used clonal-origin trees from six clones, testing for differences in responses to spacing among clones. In all three spacing experiments and for all clones, individual-tree growth was greatest at the widest spacing and stand-level growth was highest at the tightest spacing. These trends were much clearer across the wider range of spacing tested in the Nelder plots (228 to 4 760 trees ha^?1) than in the narrower range of spacing tested in the block design (1 111 to 2 222 trees ha^?1). Current annual increment reached a higher, earlier peak at narrow spacing. At 8.5 years, the light use efficiency (stem volume growth per unit of light intercepted) was about twice as great for trees at narrow spacing than at wider spacing. Overall, the Nelder designs provided the same information on responses to spacing as the classic block design. The simplicity and small size of Nelder designs provide valuable insights for basic decisions on spacing for operational plantations, particularly when forestry extends into new geographic areas, new genotypes, and new silvicultural techniques.     

Preliminary estimation of carbon stock in a logging concession with a forest management plan in East Cameroon

ABSTRACT

Logging operations in Cameroon are based on the extraction of wood from natural forests. In this article, we assessed the carbon stock in a forest management unit (FMU) located in East Cameroon from field inventory to postfelling operations up to sawmill and export terminals. Tree basal area and aboveground biomass were calculated based on trees inventoried in the annual allowable cut. We observed that from an exploitable tree potential of 0.696 trees ha^?1 inventoried within a diameter range of 50–110 cm, 0.141 tree ha^?1 (i.e., 20% of the inventoried trees) were logged. In other words, out of 6.78 tC ha^?1 inventoried, 1.84 tC ha^?1 (i.e., 27% was logged), 1.62 tC ha^?1 arrived in the log yard and 1.3 tC ha^?1 arrived in sawmill, while 0.32 tC ha^?1 reached the export terminal. In terms of damages caused on vegetation, 4.45% of all the annual allowance cut (AAC) were affected during logging activities, this represents almost 33,188.07 tons of carbon. These findings show that the implementation of reduced-impact logging (RIL) could reduce these losses throughout the logging steps and help propose a process for the valuation of wood waste in the forest and sawmill. In this context, reducing emissions from deforestation and degradation will be engaged with the right approach.     

Mechanized Direct-Seeding of Native Forests in Xingu,Central Brazil

The Y Ikatu Xingu Campaign brought together indigenous people, farmers, researchers, governmental, and non-governmental organizations seeking riparian forest restoration in the Xingu watershed, in west-central Brazil. Forest restoration is challenging in the region because of scarce nurseries, long distances, and high costs associated with the usual technique of planting nursery-raised seedlings. This article describes mechanized direct seeding and compares it with the planting of seedlings, in terms of cost and tree densities at ages of 0.5 until 5.5 yr after planting. Direct-seeding was mechanized using common agricultural machines designed for sowing cereals or grasses, which were loaded with 200,000 seeds of native trees and 150,000 seeds of annual and sub-perennial legumes, plus 50–150 kg sand ha^?1. The Campaign restored more than 900 ha by direct-seeding and 300 ha by planting seedlings. The great demand for native seeds was met by the Xingu Seed Network, formed by Indians, small landholders, and peasants, which commercialized 98 tons of native seeds and earned US$500,000 since 2006. Direct-seeding costs less per hectare than planting seedlings (US$1,845 ha^?1 against US$5,106 ha^?1), results in higher tree densities (2,500–32,250 trees ha^?1 against 1,500–1,650 trees ha^?1), is more practical, and creates layers of dense vegetation that better resembles natural forest succession.     

Process studies in aPinus radiata-pasture agroforestry system in a subhumid temperature environment. II. Analysis of dry matter yields in the third year

In this paper we analyzed the dry matter yields (DM) produced in an agroforestry trial consisting of pine trees grown over (1) Control (bare ground), (2) ryegrass/clovers (Lolium perene/Trifolium spp.), (3) ryegrass only, and (4) lucerne (Medicago sativa) during the third growing season between 1992 and 1993. In addition these pastures were grown alone in separate plots in the open. The results show that:

Dry matter production,morphology and nutritive value of <Emphasis Type="Italic">Dactylis glomerata</Emphasis> growing under different light regimes

Plant growth, morphology and nutritive value under shade can differ between temperate grasses. Therefore, the aim of this study was to quantify the dry matter (DM) production, sward morphology, crude protein (CP%), organic matter digestibility (OMD) and macro-nutrient concentrations (P, K, Mg, Ca and S) in a grazed cocksfoot (Dactylis glomerata L.) pasture under 10-year-old Pinus radiata D. Don forest. Four levels of light intensity were compared: full sunlight (100% photosynthetic photon flux density-PPFD), open + wooden slats (∼43% PPFD), trees (∼58% PPFD) and tree + slats (∼24% PPFD). The mean total DM production was 8.2 t DM ha⁻¹ yr⁻¹ in the open and 3.8 t DM ha⁻¹ yr⁻¹ in the trees + slats treatment. The changes in cocksfoot leaf area index (LAI) were related to variations in morphological aspects of the sward such as canopy height and tiller population. CP% increased as PPFD declined with mean values of 18.6% in open and 22.5% in the trees + slats treatment. In contrast, the intensity of fluctuating shade had little effect on OMD with a mean value of 79 ± 3.2%. The mean annual macro-nutrient concentrations in leaves increased as the PPFD level declined mainly between the open and the trees + slats treatments. It therefore appears that heavily shaded dominant temperate pastures in silvopastoral systems limit animal production per hectare through lower DM production rates and per animal through reduced pre-grazing pasture mass of lower bulk density from the etiolated pasture.     

Effects of understorey vegetation on tree establishment and growth in a silvopastoral system in northern Greece

The interaction between understorey plants and trees in a young silvopastoral system was studied in a sub-humid Mediterranean environment with cold winters in northern Greece. The experimental design was a split-split plot with three replications and included: three understorey treatments (grass, legume and control), two tree species (Acer pseudoplatanus and Pinus sylvestris) and two spacings (2.5×2.5 m and 3.5×3.5 m). The competition between herbaceous plants and trees resulted in significant differentiation in tree growth while their spacing did not produce any significant differences. The influence of trees on herbage yield either by plant species or spacing was not significant. During the early stages of establishment, a significant positive correlation was observed between sycamore growth and soil moisture, leaf weight, leaf area and leaf number as well as nitrogen and potassium concentration in leaves. Six years after planting the height increase of the sycamore trees was largest in the control treatment (415%), lowest in the grass treatment (134%) and intermediate (192%) in the legume treatment. Much higher was the increase obtained for the diameter, 161%, 207% and 536% respectively for the grass, legume and control treatments. The Scots pine trees grew faster than sycamore over the course of the experiment. Height increased by 397%, 351% and 400% and diameter by 518%, 443% and 683% respectively for the grass, legume and control treatments. This revised version was published online in June 2006 with corrections to the Cover Date.