Potential of natural and improved fallow using indigenous trees to facilitate cacao replanting in Ghana

Poor establishment, due to loss of soil fertility, weeds and lack of appropriate shade, is a major constraint to replanting cacao on previously used land. Spathodea campanulata, Newbouldia laevis and Ricinodendron heudelotii planted as monospecific improved fallow and Terminalia ivorensis, T. superba and Antiaris toxicaria planted as a multispecies improved fallow and a natural tree fallow were assessed for their potential to facilitate cacao replanting in a randomized complete block design experiment. Simpson and Shannon diversity indices and species richness in the natural tree fallow were 0.6, 1.6 and 20, respectively, at 4 years after trial inception. The Multispecies and the R. heudelotii improved fallows had better height growth, crown development and light transmission characteristics, which are desirable for cacao shade. However, these were not comparable to S. campanulata or the natural tree fallow in terms of improving microsite topsoil pH, % organic carbon and % total nitrogen and site capture. Since optimum fallow period is shortened by growing fast-growing trees, the height growth rate >2.0 m per annum in all the treatments except N. laevis indicates the suitability of these species for improved fallow. The trees species showed different and complementary characteristics and from a standpoint of biodiversity conservation and the future floristic composition of the landscape the natural tree fallow with its diversity of tree species may be recommended as a rehabilitation technique to facilitate the replanting of cacao with a diverse overhead shade.     

Coarse root growth of Veronese poplar trees varies with position on an erodible slope in New Zealand

Poplars are commonly planted on moist, unstable pastoral hill country to prevent or reduce soil erosion, thereby maintaining hillslope integrity and pasture production. Mechanical reinforcement by poplar root systems aids slope stabilisation, particularly when the roots are anchored into the fragipan or underlying rock. Total root length, mass and distribution of coarse roots (≥2 mm diameter) were determined for three Populus deltoides × nigra ‘Veronese’ trees in their 12th growing season after being planted as 3 m poles at upper slope (TU), mid-slope (TM) and lower slope (TL) positions on an erodible hillslope near Palmerston North in the southern North Island. Most of the roots were distributed in the top 40 cm of soil. Depth of penetration of vertical roots was dependent on slope position and limited by the available depth of the soil above a fragipan (0.35 m at the upper slope to 1.4 m at the lower slope). Roots penetrated the fragipan at the upper slope position where the soil depth was shallowest, and at the mid-slope, but not the lower slope position. Total coarse root length was 287.9 m for TU, 1,131.3 m for TM and 1,611.3 m for TL, and total coarse root dry mass (excluding root crown) was 8.15 kg for TU, 38.77 kg for TM and 81.35 kg for TL.     

Relationship between tree canopy height and the production of pasture species in a silvopastoral system based on alder trees

Silvopastoral systems in New Zealand that incorporate trees planted to control soil erosion on hills largely rely on the productivity of the pastoral system for financial returns. The effect on pasture productivity of increasing the tree canopy height by pruning Italian gray alder (Alnus cordata) was investigated by measuring the response of light, soil moisture, soil temperature, pasture production of major pasture species, and grazing behaviour of sheep. A split-plot design with four replicates was used. The main plot treatments were three levels of shade (81, 23, and 12% of available photosynthetic photon flux (PPF)), created by pruning 11 year old alder grown at the same density. The sub-plot treatments were four pasture mixes: perennial ryegrass (Lolium perenne), Yorkshire fog (Holcus lanatus), and cocksfoot (Dactylis glomerata), each sown with white clover (Trifolium repens), and cocksfoot sown with lotus (Lotus pedunculatus). Soil temperature was highest under light shade. Total herbage yield at 50 mm stubble height from October to May under heavy and medium shade was 60 and 80%, respectively, of the total herbage harvested under light shade. Cocksfoot had the greatest herbage yield, either with lotus or white clover. The tillering of perennial ryegrass was suppressed by shade more than for the other grass species making ryegrass unsuitable for use in this silvopastoral system. More sheep grazed in the light shade than in the heavy shade, but there was no difference in sheep preference for cocksfoot or Yorkshire fog. Lotus was grazed more frequently than white clover. Pruning of alder to increase canopy height has the potential to improve the productivity of the understorey pasture and its acceptability to sheep.     

Comparison of soil physical properties in evergreen and deciduous forests in central Cambodia

We investigated soil physical properties in three forest types in tropical lowland monsoon forests in central Cambodia under the same climatic conditions, i.e., Kanhaplic Haplustults in dry evergreen forest (KH-E), Arenic Haplustults in dry deciduous forest (AH-D), and Arenic Ultic Alorthods in mixed evergreen–deciduous forest (AA-M), to clarify the relationship between forest types and soil physical properties. The clay content was correlated with water content at ψ = −9.8 and −1500 kPa (WC10 and WC1500), available water capacity (AWC), and the van Genuchten (vG) parameter N (P < 0.01). vG parameter N was in the order AH-D > AA-M > KH-E whereas vG parameter α had a high value in KH-E soil at 0–100 cm in depth. The cumulative AWC (AWCcl, mm) at a soil depth of 0–200 cm was higher in the AH-D than in the KH-E, and was not considered a major factor affecting the distribution of different forest types under the same climatic conditions. The unsaturated hydraulic conductivity (K) at 0–100 cm in depth, estimated by use of models, was higher in AH-D than in KH-E mostly at matric potential ψ > −10 kPa. The low K in KH-E at ψ > −10 kPa was considered favorable for evergreen trees to retain the soil water for the transpiration in the dry season, and the matric potential in KH-E showed more gentle decreases in the early dry seasons than AH-D. Thus the differences in K among generally sandy soil types could possibly affect the establishment of different forest types in the study area with the same climate.     

Soil properties and nitrogen availability in silvopastoral plantings of Acacia melanoxylon in North Island, New Zealand

Acacia melanoxylon, a N₂-fixing timber tree occurring naturally in eastern Australia, is now promoted as a component of silvopastoral systems; but the interaction of the tree with pasture and soils has not been adequately studied. This study investigated the effects of Acacia melanoxylon on soil nitrogen (N) levels, N availability, soil pH, bulk density, organic carbon, C:N ratios and soil moisture in three separate silvopastoral sites with contrasting soil types in the North Island of New Zealand. At each site four tree stocking rates were studied (0, 500, 800, and 1700 stems ha^–1). The trees were nine years old at the time of the study. Soil samples from each study site were taken once at three depths (0 to 75 mm, 75 to 150 mm, and 150 to 300 mm), with three replicates per tree stocking rate. Soil analyses showed that although there were differences between soil types, few statistically significant differences occurred due to tree stocking rate. A greenhouse pot trial growing ryegrass (Lolium multiflorum L. cv. Concord) in soil from the A horizon of each soil type from under the trees and the open pasture found that ryegrass yield, N uptake and N supply increased with increasing tree stocking rate. Increased N supply under the trees, coupled with greater soil moisture compared to the open pasture may have accounted for the higher pasture yield under Acacia melanoxylon compared to non dinitrogen fixing tree species. This study suggested that Acacia melanoxylon in a silvopastoral system had the potential to increase soil N availability.This revised version was published online in November 2005 with corrections to the Cover Date.     

Establishment of native tropical timber trees in monoculture and mixed-species plantations: Small-scale effects on tree performance and insect herbivory

Plantations of native timber species established on former pasture are a potential strategy to reduce the logging pressure on remnant natural forests in the tropics. Such plantations may help to mitigate or reverse the negative impacts of land degradation, and they may contribute to the long-term livelihood of livestock farmers. Planting native trees is, however, perceived as a risky activity due to limited knowledge of their performance and due to marked losses of newly established seedlings attributed to insect pests. Our study focuses on the small-scale effects of environmental heterogeneity, tree diversity and insecticide treatment on the performance of three native Central American timber species two years after establishment, and on damage inflicted by insect herbivores. Growth, survival and herbivore leaf damage were quantified for Anacardium excelsum (Anacardiaceae), Cedrela odorata (Meliaceae) and Tabebuia rosea (Bignoniaceae) planted in an experiment in Panama as (1) monocultures, (2) mixed stands, and (3) mixed stands protected by insecticides. Our study revealed that small-scale effects can have a substantial impact on the success of timber trees planted on former pasture. Growth performance and survival of the three species was strongly affected by small-scale environmental heterogeneity, which was expressed as significant differences in growth and survival among different plantation plots at the same study site. Establishment of trees in mixed stands did not have significant effects on tree survival and growth compared to pure stands, although it reduced herbivore pressure in one of the studied tree species. All tree species grew best and suffered lowest leaf damage when protected by insecticides, indicating a general influence of insect herbivory on growth of newly established trees. In contrast to growth performance, survival was not significantly affected by different management practices or herbivory. The large variability among plots in tree survival and growth, and also in the effects of management practices such as planting design and insecticide treatment, emphasizes the importance of small-scale environmental heterogeneity on tree survival and growth.     

Sugar maple and yellow birch regeneration in response to canopy opening,liming and vegetation control in a temperate deciduous forest of Quebec

We examined how the density, growth and survival of sugar maple (Acer saccharum Marsh.) and yellow birch (Betula alleghaniensis Britton) regeneration are influenced by gap size, soil nutrient availability and understory vegetation. We used a factorial combination of (1) three gap sizes (small: <100 m²; medium: 100–300 m²; large: ∼1000 m²); (2) presence/absence of liming (92% CaCO₃ at 500 kg ha⁻¹, 1st year post-harvest); and (3) presence/absence of vegetation control (weeding twice a year; 1st to 3rd year post-harvest). We monitored height increment and survival of 1500 seedlings and saplings of both species from the 3rd to the 6th year post-harvest, and assessed density 6 years post-harvest. Both species exhibited a complex set of density, growth and survival responses across the combination of treatments. Compared to sugar maple, yellow birch had an overall lower density, greater growth, and similar survival rate; the two species attained maximum values in different gap size for density, and similar gap size for growth and survival. Liming had very little or no effect on the species. The growth of yellow birch was slightly but significantly greater when understory vegetation was controlled, particularly in medium and large gaps. These results suggest that a variety of canopy gap sizes can provide the right combination of understory conditions for regenerating these two functionally different tree species.     

Tree-grass relationships in open eucalypt woodlands of northeastern Australia: influence of trees on pasture productivity, forage quality and species distribution

Pasture yield, quality and species distributions were compared between zones around live and killed eucalypt trees at two woodland sites in northeast Queensland which differed markedly in soil fertility. Trees affected pasture quality and yield on an individual tree basis: N concentration and dry matter digestibility tended to be higher under trees than in inter-tree areas at both sites and pasture yields declined with distance from killed trees at the lower fertility site. However, the distribution of individual species did not vary markedly with distance from trees. Trees also affected pasture yield on a woodland basis: yields were greater where the trees were killed than under intact woodland. Soil under trees had higher levels of organic carbon and greater litter cover than soil in inter-tree areas. However, pasture yields did not generally reflect this fertility gradient since growth was limited by moisture availability due to drought conditions during the study period. Pasture N concentration was higher under trees that in inter-tree areas since plants under trees produced a similar amount of biomass as plants in inter-tree areas, but had access to higher nutrient levels. Trees appeared to have a greater effect on soil nutrient availability at the low fertility site. Live trees depressed pasture yields to a lesser degree at the low fertility site, demonstrating that the effects of trees on soil water availability (on a woodland scale) are less important when soil nutrients are more limiting to growth. These results indicate that, while removing trees may enhance pasture productivity, this benefit may be offset by a reduction in pasture quality. Given the beneficial effect of trees on soil nutrients, tree removal may also have longer term implications for soil nutrient dynamics. This revised version was published online in August 2006 with corrections to the Cover Date.     

林豆复合生态系统土壤理化性质的研究

本文对混种大豆的落叶松.大豆、水曲柳-大豆复合生态系统土壤理化性质进行研究。结果表明,在一个生长季内混种后的土壤物理性质得到了改善。落叶松.大豆与水曲柳一大豆复合生态系统的土壤容重为1.112g/cm3和1.058g/cm3均低于相对应的纯林:混种大豆后土壤总孔隙度增加。两种林-豆复合生态系统土壤有机质分别比对应纯林高1.77倍和1.09倍:落叶松一大豆复合生态系统全氮和水解氮含量分别高于落叶松纯林4.2%和53.0/%,水曲柳-大豆复合生态系统全氮和水解氮含量分别高于水曲柳纯林75.5%和3.3%:混种大豆后全磷含量降低,而有效磷含量则增加:落叶松-大豆复合生态系统全钾和有效钾比落叶松纯林高0.6%和17.5%,水曲柳-大豆复合生态系统全钾和有效钾分别比水曲柳纯林高56.4%和21.8%。图1表3参27。     

Effects of intercropping systems of trees with soybean on soil physicochemical properties in juvenile plantations

The intercropping system of tree with soybean in juvenile plantations, as a short-term practice, was applied at Lao Shan Experimental Station in Mao’er Shan Forest of Northeast Forestry University, Harbin, China. The larch (Larix gmelinii)/soybean (Glycine max.) and ash (Fraxinus mandshurica) intercropping systems were studied in the field to assess the effects of the intercropping on soil physicochemical properties. The results showed that soil physical properties were improved after soybean intercropping with larch and ash in one growing season. The soil bulk density in larch/soybean and ash/soybean systems was 1.112 g·cm⁻³ and 1.058 g·cm⁻³, respectively, which was lower than that in the pure larch or ash plantation without intercropping. The total soil porosity also increased after intercropping. The organic matter amount in larch/soybean system was 1.77 times higher than that in the pure larch plantation, and it was 1.09 times higher in ash/soybean system than that in the pure ash plantation. Contents of total nitrogen and hydrolyzable nitrogen in larch/soybean system were 4.2% and 53.0% higher than those in the pure larch stand. Total nitrogen and hydrolyzable nitrogen contents in ash/soybean system were 75.5% and 3.3% higher than those in the pure ash plantation. Total phosphorus content decreased after intercropping, while change of available phosphorus showed an increasing trend. Total potassium and available potassium contents in the larch/soybean system were 0.6% and 17.5% higher than those in the pure larch stand. Total potassium and available potassium contents in the ash/soybean system were 56.4% and 21.8% higher than those in the pure ash plantation. Biography: FAN A-nan (1972–), female, Ph. Doctor in Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, P. R. China.     

Structural root growth of young Veronese poplars on erodible slopes in the southern North Island,New Zealand

In New Zealand poplars are commonly planted on moist, unstable pastoral hill country to prevent or reduce soil erosion, thereby maintaining hillslope integrity and pasture production. Mechanical reinforcement by poplar root systems aids slope stabilisation. Root mass and distribution were determined for three Populus deltoides × nigra ‘Veronese’ trees aged 5, 7 and 9.5 year planted as 3 m poles at 8 m × 8 m spacing on a hillslope near Palmerston North in the southern North Island. Most of the structural roots (≥2 mm diameter) were distributed in the top 40 cm of soil. Vertical roots penetrated to about 1.0 m, being the depth of the soil above a fragipan. Total structural root dry masses (excluding root crown) were 0.57, 7.8 and 17.90 kg for the trees aged 5, 7 and 9.5 year, respectively. Total structural root length was 79.4 m for the 5 year tree and 663.5 m for the 9.5 year tree. Surrounding trees were estimated to increase root mass density to 3 times and root length density to 4–5 times the contribution of the single tree at 9.5 year. The study indicated that root development of wide-spaced poplar trees on hillslopes was minimal in the first 5 years but then increased rapidly. These results suggest that poplar trees established from poles may take at least 5 years to develop a structural root network that will effectively bind soil.     

南京地区极端低温对常绿绿化树种景观的影响及分析

通过调查2004年冬季南京市7个不同地域常绿绿化树种所发生的冻害情况,分析了该地当年温度、降水等气候因子变化对常绿绿化树种景观效果的影响.     

不同经营措施马尾松林生物量和土壤性质研究

通过对永春本地和从广东罗定县引进的马尾松造林后采取的集约经营和一般经营两种措施比较,结果表明,集约经营的本地松和罗定松10年生时其平均木生物量与一般经营的本地松相比提高20%以上,集约经营的马尾松与一般经营相比,土壤容重下降而土壤持水量和孔隙度增加,土壤有机质和速效磷含量提高,集约经营的马尾松林土壤理化性质得到改善。     

Community ecology and spatial distribution of trees in a tropical wet evergreen forest in Kaptai national park in Chittagong Hill Tracts,Bangladesh

We investigated the floristic composition, woody species diversity and spatial distribution of trees in a tropical wet evergreen forest in Kaptai national park in Chittagong Hill Tracks, Bangladesh. We recorded 25 families, 37 genera, 40 species and 1771 woody individuals in a 0.09 ha plot. Euphorbiaceae and Moraceae were the most speciesrich families, and Castanopsis, Ficus and Terminalia were the most species-rich genera. Bursera serrata Wall. ex Colebr. was the dominant species in terms of highest importance value(13%). Trema orientalis(L.)Bl was typically a light demanding species as it appeared in the top canopy with only one individual having the seventh highest IV, but had no regeneration. The expected maximum number of species(Smax) was 140,indicating that many species may invade the forest as the Smax is greater than the recorded total number of species. The nature of the disappearance and appearance of species in the present forest reflects instability of floristic composition. The values of Shannon's index H ′ and Pielou's index J ′(evenness) were 3.36 bit and 0.63, respectively. These values show moderately high species diversity as compared to other subtropical forests in the tropics. In addition, a sample area of 200 m2 in this forest would be sufficient for measuring the diversity indices H ′ and J ′,whereas the trend of J ′may indicate the rate of equality of individuals among the different species decreased with increasing area. The distribution pattern for the total stand was completely random. However, the dominant species showed aggregate distribution for small areas, but random distribution for large areas. The spatial association between species showed that the strongest positive interspecific association occurred between Streblus asper Lour. and Castanea indica Roxb.(ω =0.51). As a whole, most species were weakly associated with each other,of which 58% species associations were completely negative. The result of cluster analysis showed that species pairs were spatially independent at all or most small clusters; stands of species from all clusters are mosaics of complete habitat and pioneer habitat. All patches in this forest community have similar habitat and regeneration niches, which could be a phenomenon for a young growth forest. Thus, the existence of habitat and regeneration niches may be an important factor in the maintenance of diversity in this forest.     

Postharvest organic matter removal effects on FH layer and mineral soil characteristics in four New Zealand Pinus radiata plantations

The effects of three postharvest organic matter removal treatments on fermentation-humus (FH) layer and soil characteristics were compared in replicated trials in four second rotation New Zealand Pinus radiata plantation forests, planted 8–16 years prior to sampling. All sites were sampled in early 2002 and 2003. Increasing organic matter removal significantly decreased the mass of the FH layer in the treatment plots, the moisture content in the FH layer and mineral soil, the concentration of carbon in the FH layer and mineral soil, the pool of carbon stored in the FH layer, the concentration of nitrogen in the mineral soil and the pool of nitrogen stored in the FH layer. Mineral soil pH was significantly increased with increasing organic matter removal. The persistence of the significant differences in the FH layer and mineral soil characteristics strongly suggested that variations in organic matter removal have long-term effects on forest floor properties, and significantly influences carbon storage over the life of the rotation.     

Effects of widely spaced trees and livestock grazing on understory environments in tropical savannas

The effects of widely spaced trees ofAcacia tortilis andAdansonia digitata on their understory environments were investigated in four savannas located along a gradient of increasing livestock utilization in Tsavo National Park (West), Kenya. Plant species composition and biomass and the physical and chemical properties of soils that occur below tree crowns were compared to open grasslands. The tree-crown zones at lightly and moderately grazed sites had a unique understory flora and higher plant biomass, lower temperatures and bulk densities, and higher levels of P, K, Ca and mineralizable N than their associated opengrassland zones. In the heavily grazed savanna, few differences between tree-crown and grassland zones were found. The beneficial effects of savanna trees on their understory environments appear to diminish with increasing livestock utilization.     

Nitrogen fertiliser effects on litter fall, FH layer and mineral soil characteristics in New Zealand Pinus radiata plantations

The effects of nitrogenous fertilisation on litter fall, FH layer and soil characteristics were investigated in replicated trials in six second rotation New Zealand Pinus radiata plantation forests. Four trial sites also incorporated three different post-harvest organic matter removal treatments. All sites were sampled in early 2002 and 2003. Fertilisation significantly increased the nitrogen content and decreased the carbon:nitrogen ratio of the litter fall. Fertilisation significantly increased the mass of the FH layer in the treatment plots, moisture content in the FH layer, the concentration of nitrogen in the FH layer and the pool of carbon and nitrogen stored in the FH layer. Fertilisation significantly increased the nitrogen concentration of the mineral soil, and decreased the mineral soil carbon:nitrogen ratio and pH. Several significant site × fertilisation interaction terms indicated that variations in the fertilisation regimes and site characteristics substantially influenced the effects of fertilisation. Fertilisation did not significantly decrease the relative differences between the organic matter removal treatments. The significant differences in the litter fall, FH layer and mineral soil characteristics strongly suggest that nitrogenous fertilisation has the capacity to significantly alter the forest floor environment, and may be able to increase carbon storage over the life of the rotation.     

Relative effects of irrigation and intense shade on productivity of alley-cropped tall fescue herbage

The relative effects of irradiance and soil water on alley-cropped herbage are poorly understood. Our objective was to determine effects of irrigation on herbage productivity when tall fescue [Lolium arundinaceum (Schreb.) Darbysh.] was grown in two sites, a meadow and a loblolly pine (Pinus taeda L.) alley (620 trees ha⁻¹), near Booneville, Arkansas. Three tall fescue entries were space planted in the meadow and pine alley with or without irrigation. Herbage mass and nutritive value were determined at 8-week intervals for 2 years. Mean daily PAR was 33.9 (2004) and 37.5 mol m⁻² d⁻¹ (2005) in the meadow, while the pine alley received 5.6 mol m⁻² d⁻¹ (17% of the meadow) in 2004 and 4.3 mol m⁻² d⁻¹ (11% of meadow) in 2005. Effect of tall fescue entry tended to be small relative to site and irrigation. Irrigation compensated for evapotranspiration in the meadow, but not in the pine alley when summer rainfall was about normal (2004) or low (2005). Nevertheless, site (PAR) had a greater effect on herbage specific leaf weight, leaf elongation rate, tillers plant⁻¹, mass tiller⁻¹, and total nonstructural carbohydrate concentration than soil water. Irrigation might have had greater impact on herbage productivity if more water had been applied or if canopy cover had been less extreme. Silvicultural practices should be imposed to improve penetration of solar irradiance to the alley crop.     

Identification of key soil indicators influencing plantation productivity and sustainability across a national trial series in New Zealand

New Zealand is committed to developing sustainable forest management practices as evidenced through Government involvement in international forestry agreements such as the Montreal Process, and the forestry sector's adoption of forest certification mechanisms. Despite this, there is little quantitative evidence of how plantation forestry affects site quality and long-term site productivity. To address this issue, a nationwide study of site quality was established at 31 key sites covering the range of edaphic and environmental conditions for New Zealand plantation forests. At each location, eight short-term site quality plots were planted at a very high stand density (40,000 stems ha⁻¹) to rapidly identify key soil indicators of growth which may be useful for determining site sustainability. The plots at each site were arranged in a factorial design with the following three factors: species (Pinus radiata D. Don and Cupressus lusitanica Mill.), fertiliser (no fertiliser and nutrients supplied in excess of crop demands) and disturbance (low and high disturbance). These plots were harvested after 4 years and data was used to (i) examine treatment and site effects on volume mean annual increment (MAI) and (ii) identify key soil properties that influence volume MAI for the two species.Volume MAI significantly ranged 7-fold across sites. For the treatments, species accounted for most of the variance in volume MAI, with values for P. radiata significantly exceeding those of C. lusitanica by 95%. Volume MAI was significantly affected by fertilisation which induced gains of 33%. Disturbance did not significantly affect volume MAI. After correction had been made for climatic variables, soil properties that were most strongly related to volume MAI for both species included C:N ratio, total phosphorus (P), and organic P. When soil properties were included in combination, the best predictive models of volume MAI formulated for P. radiata included total P and C:N ratio while the best predictive model for C. lusitanica included C:N ratio and Olsen P. Variation in species sensitivity to total P and C:N ratio is likely to be attributable to their different mycorrhizal associations. For both species, the most important soil physical property influenced by management operations was total porosity. This research suggests that total P, C:N ratio, Olsen P and total porosity should be used as indicator properties for determining sustainability of plantation grown P. radiata and C. lusitanica.