Silvicultural systems for southern bottomland hardwood forests

Silvicultural systems integrate both regeneration and intermediate operations in an orderly process for managing forest stands. The clearcutting method of regeneration favors the development of species that are moderately intolerant to intolerant of shade. In fact, clearcutting is the most proven and widely used method of successfully regenerating bottomland oak species in the South. The seed-tree method of regeneration favors the establishment of light-seeded species. Mechanical soil scarification may be necessary if the desired species requires bare mineral soil for establishment. The shelterwood method of regeneration can provide for the development of heavy-seeded species, but has produced highly variable results with southern bottomland oaks. The single-tree selection method of regeneration favors the development of shade-tolerant species. When single-tree selection is applied repeatedly to stands containing commercially valuable shade-intolerant species, composition will gradually shift to less-valuable, more-tolerant species. Consequently, the single-tree selection method of regeneration is not recommended for any commercially valuable bottomland hardwood tree species. Group selection, in its strictest application, creates only small openings that usually fail to allow sufficient light to the forest floor for satisfactory establishment and development of shade-intolerant bottomland species. Patch cutting, a combination of uneven-aged (group selection) and even-aged (clearcutting) silviculture, designed to create larger openings, has been successfully used to produce an uneven-aged stand that consists of many small, irregularly shaped, even-aged groups. Silvicultural systems should include a planned program of intermediate operations designed to enhance the growth and development of those species favored during the regeneration process. Improvement cutting and commercial thinning are increasingly common in southern bottomland hardwood forests. Other partial cuttings employed today in bottomland hardwood forests typically involve some form of crop-tree release. Specific recommendations for the selection of silvicultural systems are presented for the eight most important species groups found in southern bottomland hardwood forests.     

Species composition and dynamics in two hardwood stands in Vermont: a disturbance history

Stand composition and structure utilizing stem analysis was studied in two hardwood stands in Vermont. In a mixed hardwood stand with some white pine and hemlock, a major entry of new trees in the main canopy seems associated with harvesting coincident with land exchange. More recent partial cuttings have promoted establishment of new seedlings or development of suppressed advanced-growth shade-tolerant beech, (Fagus grandifolia, Ehrh.), hophornbeam (Ostrya virginiana, (Mill) K. Koch), and striped maple (Acer pensylvanicum, L.). Very few sugar maple and red oak seedlings and saplings are present.

In a northern hardwood stand some red spruce (Picea rubens, Sarg.), that were 240 to 306 years old, became established before any known harvest, and exhibited release following harvests of the mid-1800s. This major harvest, coupled with the differential growth between spruce and hardwoods, and seed/seedling availability, resulted in a major change in stand composition. Trees now in the main canopy of sampled stands appear to have either been released or newly established following various harvests. Harvests have been of such frequency that natural disturbances seem insignificant. Many of the competitive understory species have become abundant following harvests of the 1960s and 1980s and may have been present as advanced growth and responded to the release. Following the harvest of 1981–1982, abundant yellow birch became established on skid trails. Elsewhere in the stand, yellow birch seedlings and saplings are only in great abundance in areas that were possibly sizable gaps following earlier harvesting.

The dynamics of tree entry and growth in gaps of small or large size probably occur in a similar way in many other stands of the region. Though the sampling of this study is limited, there is no suggestion of continuous tree establishment at any particular location, the new age classes seem associated with either a gap or stand replacing disturbance attributed to harvesting.     

Influence of harvesting on biogeochemical exchange in sheetflow and soil processes in a eutrophic floodplain forest

Floodplain forests contribute to the maintenance of water quality as a result of various biogeochemical transformations which occur within them. In particular, they can serve as sinks for nutrient run-off from adjacent uplands or as nutrient transformers as water moves downstream. However, little is known about the potential that land management activities may have for alteration of these biogeochemical functions. This paper examines the effects of three harvesting regimes (unharvested control, clearcut, and partial cut) on the physical and chemical parameters within the Flint River floodplain located in southwestern Georgia, USA. Data presented in this paper were collected during the year following initiation of the harvesting treatments which occurred in September of 1993. Sheetflow water chemistry (total suspended solids (TSS), total dissolved solids (TDS), nitrate (NO₃⁻), phosphate (PO₄³⁻), sulfate (SO₄²⁻), calcium (Ca²⁺), potassium (K⁺), magnesium (Mg²⁺), ammonium (NH₄⁺), total phosphorous (P), total nitrogen (N), total carbon (C), dissolved organic carbon (DOC)), sedimentation rates, depth of soil oxidation after flooding, saturated hydraulic conductivity, and bulk density were measured. During the year immediately after treatment installation, alterations in some of the physical and chemical properties (TDS, NO₃⁻, total P, and K⁺) of floodwaters crossing harvest plots were detected. Soil oxidation depths, saturated hydraulic conductivity and bulk density also changed with treatment. The meaning of the changes detected is uncertain but they suggest the nature of potential changes in nutrient spiralling and non-point source cumulative effects that may occur within a managed watershed. Second-year data may offer an interesting comparison of sheetflow chemistry and sedimentation changes between vegetated and non-vegetated conditions.     

Ground-layer plant community responses to even-age and uneven-age silvicultural treatments in Wisconsin northern hardwood forests

We evaluated ground-layer plant diversity and community composition in northern hardwood forests among uncut controls and stands managed with even-age or uneven-age silvicultural systems. Even-age treatments included diameter-limit cuttings (20-cm diameter at 30-cm stem height) in 1952 and shelterwood removals in 1964. Uneven-age treatments included three intensities of selection harvest (light, 20.6 m²/ha residual basal area after harvest; medium, 17.2 m²/ha residual basal area; and heavy, 13.8 m²/ha residual basal area) that were applied in 1952, 1962, 1972, and 1982. All treatments were winter logged over snow pack. In 1991, plant diversity and community composition were examined. Species richness for spring ephemerals ranged from 1 to 6 species/150-m², spring ephemeral diversity (Shannon's Index of Diversity) averaged 0.57 ± 0.04 and evenness averaged 0.45 ± 0.03. Summer flowering species richness ranged from 1 to 18 species/1-m², with an average diversity of 0.71 ± 0.07 and evenness of 0.42 ± 0.03. We found no significant differences among treatments for any of these variables, although power to detect a difference (at p = 0.05) was low in all cases (0.15–0.55) due to high variance and low replication. Community composition was not significantly different among the treatments, for either spring (p = 0.09) or summer (p = 0.79) flora. Few exotic species were found in any treatment. Lack of exotic invasions and minimal differences in plant diversity or composition among treatments may be due, in part, to the negligible amount of soil disturbance that resulted from winter logging. While other (unmeasured) ecosystem components may differ among these silvicultural treatments, our results suggest that ground-layer plant communities in northern hardwood ecosystems are either resistant to change or have recovered within the 40 years since disturbance in the even-age treatments and within 10 years since disturbance in the uneven-age treatments.     

The bottomland hardwood forest of the southern United States

Bottomland hardwood forests are valued for timber production, water storage, enhanced water quality, nutrient cycling, erosion control and wildlife habitat. However, the majority of southern bottomland stands, 90% of which are in private ownership, are occupied by a degraded mixture of tree species, caused largely by repeated, incomplete harvests. They can be naturally regenerated to a stand of favorable species composition by removing the residual stand of merchantable and non-merchantable trees. For timber production, these systems respond best to even-aged management such as clearcutting, shelterwood cutting and patch clearcutting. Plant succession under such systems allows shade intolerant species to occupy the site, followed by species of increasing shade tolerance. Uneven-aged systems, such as individual-tree selection and group selection, are also viable regeneration methods, but they require great attention to detail and frequent stand entry. In areas of high sensitivity to timber harvesting, a two-aged system such as a leave-tree is recommended. Stand disturbance, either planned or unplanned, is needed to keep southern hardwood forest stands viable.     

Immediate impacts of partial cutting strategies on stand characteristics and value

This study evaluated the impacts of partial cutting on stand characteristics, product recovery, and financial return in mature black spruce-balsam fir stands in Quebec. Four harvesting strategies (clearcut with advance growth protection, irregular shelterwood cutting leaving small merchantable stems, and two patterns of selection cutting) were each applied four times in 20 ha harvest blocks representing irregular black spruce-balsam fir stands. Before the four harvesting strategies were applied, there were no significant differences in stand characteristics (i.e., quadratic mean DBH, basal area, and merchantable stem volume) or expected product recoveries (i.e., lumber volume and value, chip volume and value, and total product recovery) estimated using the Optitek sawing simulation package. There was no significant difference in stand characteristics or product recovery values of the harvested stems between the selection cutting approaches (p > 0.05). However, significant differences in stand characteristics and product recovery values of the harvested stems existed between these treatments and both of the two other treatments. After cutting, the two selection cutting treatments had the lowest impacts on stand characteristics, as compared to the two other treatments. The selection cutting approach which used temporary skidding trails and where cutting was initially concentrated over half of the stand resulted in the highest benefit/cost ratio, relatively high net income and high total product value of residual trees.     

Effects of strip wise tillage in combination with liming on chemical and physical properties of acidic spruce forest soils after clear cutting

We investigated the initial effects of strip wise soil loosening (0–35 cm depth) on soil chemical and physical parameters by using a deeply working rotary cultivator in combination with liming and mixing of the dolomite with the soil material of acidic forests. The investigations took place 8 months after the treatment. pH values and contents of exchangeable Ca and Mg increased significantly at the tilled depth whereas the content of exchangeable Al and easily soluble P decreased. The rate of mineralisation increased at this depth which was shown by a loss of Corg, Ntot and short-term loss of NO₃-N. The treatment led to a mobilisation of Mn at the tilled depth. However, the content of exchangeable Pb decreased due to an increased pH value. Below the tillage depth of 35 cm only partly significant changes of exchangeable Mn and NO₃-N were found. The total porosity and bulk density at 10–15 and 40–45 cm depths were not significantly different from those in the control plot, but the rate of infiltration increased significantly.     

Regeneration strategies in a temperate hardwood floodplain forest of the Upper Rhine: sexual versus vegetative reproduction of woody species

The regeneration mechanisms of woody species in the hardwood floodplain forest of the Upper Rhine are not well known, although they are of primary importance for future forest development. To gain a better understanding of the balance between sexual and asexual regeneration strategies and to assess the role of the seed bank in regeneration, the similarities in species composition and abundance of four fundamental compartments involved in regeneration (the seed rain, the seed bank, the recruits and the canopy) were compared in three hardwood forest stands with different flooding conditions. The results show that the floristic composition of the recruits is very similar to that of the canopy, whereas the composition of the seed bank is very dissimilar to the latter and comprises very few hardwood species. However, some species such as Fraxinus excelsior, Carpinus betulus and Acer pseudoplatanus which are very rare in the seed bank release a high number of diaspores, and seedlings of these species germinate abundantly in the field in the year following seed release. Moreover, the germinating seeds originate from the litter layer and not from the soil itself. This shows that most woody species regenerating by seed in the hardwood Rhine forest build transient seed banks and that the role of persistent seed banks for regeneration is very limited. Furthermore, it appears that many woody species have developed strategies favouring vegetative propagation for their regeneration, particularly understorey species, such as Cornus sanguinea and Prunus padus. As vegetatively grown individuals better withstand prolonged inundation in their early life stages than seedlings, species relying on vegetative regeneration strategies might be advantaged by regular and prolonged flooding of the Rhine forest over species regenerating only by seeds.     

Indices of soil nitrogen availability for an ecological site classification of British forests

An adequate supply of nitrogen (N) is essential for the successful establishment and sustainable productivity of forest stands. N deficits may necessitate the use of artificial fertilisers. Availability of N in the inorganic forms, and the relative abundance of the NO₃-N and NH₄-N components, influences the species composition of natural forest vegetation. Hence it is essential to use reliable measures of soil N supply that fully reflect its ecological significance. The new Ecological Site Classification (ESC) used in British forestry employs a multi-factorial definition of soil nutrient regime (SNR), including soil N. To develop this, a soil and vegetation study was made at 89 forest sites throughout Great Britain covering the major soil types used for forestry. “Total N” levels were compared with separate pre- and post-incubation measures of the two inorganic N components as potential indices of soil N supply. Multivariate statistical analysis showed that the major discriminant chemical variables for the sampled soils were pH, calcium and NO₃-N and that these were also the main variables influencing the species composition of the ground vegetation. Total N and NH₄-N were less effective discriminant variables for these sites. In some infertile soils the levels of NH₄-N or total N may be of greater importance, as NO₃-N is usually in very limited supply. A multivariate gradient of SNR, which incorporates the NO₃-N measures, has been adopted for use within the ESC system. The position of a site on this gradient can be estimated quantitatively from soil type, ground vegetation species composition and humus type. This enables soil N supply and overall SNR to be assessed in a simple but effective way that guides the operational management of British forest soils for sustainable productivity. It will also be possible to use these techniques to monitor the nutritional status of forest sites over time.     

Changes in microclimate after stand conversion in two northern hardwood stands

Changes in air temperature, soil temperature, and soil moisture were monitored for 5 years in two northern hardwood stands after whole-tree harvesting and conversion to red pine (Pinus resinosa Ait.) plantations. Soil temperatures at a depth of 5 cm and maximum air temperatures 2 m above the soil surface were increased 5–25% after stand conversion. Soil moisture content at a depth of 5 cm was increased by 10–20% in one stand but not in the other. Differences in stand, soil, and topographical characteristics between the two stands did not have any apparent effect on the magnitude of air or soil temperature changes after stand conversion. However, higher initial stand density and soil water holding capacity appeared to be related to increased soil moisture content at one of the sites. The increased soil temperatures after conversion were not only a result of the removal of the northern hardwood canopy but also the removal and redistribution of the forest floor caused by whole-tree harvesting. Five years after stand conversion air temperature, soil temperature, and soil moisture showed no evidence of recovering from initial post-harvest levels.     

Long-term response of planted conifers, natural regeneration, and vegetation to harvesting, scalping, and weeding on a boreal mixedwood site

This study reports 14th-year response of a boreal mixedwood stand to different harvest intensities (uncut, 50% partial cut with and without removal of residuals after 3 years, and clearcut), spot site preparation treatments (none and scalped), and chemical weeding frequencies (none, single, and multiple) in northeastern Ontario. The response variables include the survival and growth of planted white spruce (Picea glauca [Moench] Voss) and jack pine (Pinus banksiana Lamb.), height and density of natural regeneration and shrubs, and cover of shrubs and non-woody vegetation. Harvesting and weeding generally improved survival and growth of planted trees, although white spruce survival did not significantly differ among the three weeding frequencies. Harvesting tended to increase heights of hardwood (mostly trembling aspen (Populus tremuloides Michx.)) and conifer (largely balsam fir (Abies balsamea (L.) Mill.).) natural regeneration, cover and density of shrubs, and cover of herbs, lichens, and ferns. Chemical weeding reduced height, density and cover of shrubs, height and density of hardwood regeneration, and fern cover, but increased moss and lichen cover. Spot scalping did not significantly affect planted seedling, natural regeneration, or the vegetation.Maximum survival and growth of planted white spruce and jack pine were achieved using a combination of clearcutting and multiple weeding. However, partial cutting followed by a single weeding produced acceptable survival and reasonable growth of planted trees, particularly for white spruce. Partial canopy removal alone substantially reduced the amount of hardwood regeneration, relative to clearcutting, but did not adequately suppress understory shrubs. Significant improvement in seedling growth following multiple weedings was evident primarily in the complete canopy removal treatments: 50% partial cut with removal of residuals after 3 years and clearcut. While the effects of harvesting and weeding on planted crop trees found in the 5th-year assessments generally persisted at year 14, survival decreased, likely due to light competition from developing hardwood and shrubs.     

Influences of harvesting on functions of floodplain forests associated with low-order, blackwater streams

The influence of both aerial and ground-based harvesting on functions of forested floodplains of low-order streams was studied during a two-year period. The study sites were associated with low-order, blackwater streams with infertile and primarily organic soils. Responses to harvesting were assessed in relation to water quality, denitrification, hydrology, regeneration, and decomposition. Water quality indices included nitrate, phosphate, total and dissolved solids, and biological oxygen demand (BOD). All were unaffected by harvesting with the exception of BOD which rose once to undesirable levels. Denitrification was highly variable and also showed no significant harvest effect. Hydrologic parameters included groundwater table depths which, unexpectedly, indicated lowered water tables within harvested areas during the first growing season. Early regeneration responses were strongly linked to harvest system with primarily seed-origin species favored by ground-based activity whereas sprout-origin species dominated on aerial system plots. Owing to inherent soil wetness, decomposition responded slowly to harvest disturbance. However, after one year, decomposition was more rapid on harvested plots than in undisturbed areas.     

Factors affecting runoff and soil erosion: plot-level soil loss monitoring for assessing sustainability of forest management

The assessment on key ecological factors affecting runoff and soil erosion and the usefulness of plot-level monitoring of soil erosion was conducted by collecting runoff and soil loss records from 14 runoff plots. The runoff plots were set up in two catchments in Central Kalimantan, Indonesia, where conventional logging and Reduced Impact Logging (RIL) took place. Runoff plots were set up in forest areas with different levels of logging disturbances, i.e. harvesting areas (four plots), skid trails (six plots), and undisturbed/control areas (four plots). The magnitude of runoff and soil loss from skid trail plots were found to be the highest, followed by control plots and harvest plots. Canopy cover, sapling density, litter depth and woody debris appeared to be important ecological factors that determine the magnitude of soil loss. Tree canopy determines the size and erosive power of the raindrops. Sapling, litter layer, and woody debris protected soil surface, thus preventing soil detachment, and provided surface roughness that minimised soil particle movement down the slope. The roles of these ecological factors were less significant compared to rainfall in determining the magnitude of runoff.

Canopy cover, sapling density, litter depth and woody debris can be measured quantitatively or qualitatively without complicated equipment and methods. Furthermore, they are sensitive to logging disturbance which make them suitable verifiers of soil erosion. Forest managers need to limit disturbance to these factors in order to minimise soil erosion in their logging operation areas. Monitoring of soil loss using runoff plots was cost-effective and provided valuable information about soil erosion risks caused by logging operations. Runoff plots clearly demonstrated site disturbances where the plots are located. Monitoring allowed more direct linkages to be made between management practices and their impacts on runoff and soil erosion, thereby enabling forest managers to identify problems and take appropriate preventive measures to improve their management practices.     

Effects of experimental forest harvesting on oribatid mite biodiversity

Ecosystem-based management (e.g. partial cut harvesting) attempts to mimic natural forest dynamics and maintain structural complexity, and this less intense harvesting may minimize the impact on forest floor fauna and help maintain soil system biodiversity. We tested how different experimental harvesting regimes affect the diversity, abundance and composition of Oribatida at the sylviculture et aménagement forestiers écosystémique (SAFE) research forest located in the Abitibi region in NW Québec. Litter and soil were sampled in June 2006 in the mixedwood boreal forest at SAFE where the following treatments were applied and replicated three times: clear cut harvest, 1/3 partial cut harvest, 2/3 partial cut harvest, prescribed burn (after clear cut harvest) and uncut control. Eight years after harvest, partial cuts had more similar species composition to the uncut control within their respective blocks; however, burned habitat showed a shift in species dominance patterns and harboured a relatively distinct composition and species richness compared to treatments. With the exception of samples from clear cuts, species composition of the harvesting treatments was more similar within blocks than among blocks, suggesting that for less intense harvesting practices, spatial scale (i.e. regional factors) could have a greater influence in structuring oribatid assemblages than harvesting regime, but in more severe disturbances such as burn-after-clear cut harvest, habitat is altered enough to affect oribatid biodiversity.     

Effects of white-tailed deer on vegetation structure and woody seedling composition in three forest types on the Piedmont Plateau

White-tailed deer (Ododcoileus virginiana) can substantially affect the structure and species composition of a forest. The tolerance of a forest community to browsing may vary by type as a result of varying biotic and abiotic factors of the environment. To date, no studies have compared the effects of browsing among forest communities within a physiographic region. We investigated the effects of browsing on vegetation structure and woody seedling composition in three forest types (oak–hickory, Virginia pine–eastern red cedar, bottomland hardwood) in Manassas National Battlefield Park (MNBP), Virginia, USA. We compared forb cover, vertical plant cover (0–1.5 m tall), and survival of tagged seedlings in 10 exclosed (2 m × 6 m) and 10 unexclosed plots in each forest type during a 5-year period. No differential effects of browsing were found among forest types. In all forest types, deer (67 deer/km²) suppressed forb and vertical plant cover to levels less than would be expected in the absence of deer. Seedling survival rates of most species were significantly reduced by browsing. By the 4th year of the study, box elder (Acer negundo), hickory (Carya spp.), and red maple (Acer rubrum) had been eliminated from unexclosed plots, and red and white oaks (Quercus spp.) dramatically reduced. Ash (Fraxinus spp.), black cherry (Prunus serotina), and hackberry (Celtis occidentalis), although significantly impacted, remained the most abundant species throughout the study. These findings suggest that white-tailed deer may be modifying the structure of the forest interior to the extent that it adversely affects wildlife species dependent on a dense understory to thrive. We predict that the future composition of forests in MNBP will shift towards stands with fewer species and a greater dominance of ash, black cherry, and hackberry, particularly in the oak–hickory and bottomland hardwood forests, where the majority of current dominants are most affected.     

不同强度采伐5年后杉阔混交人工林土壤呼吸速率差异

【目的】比较不同采伐强度下闽北杉阔混交人工林土壤及其各组分的呼吸速率差异,揭示土壤总呼吸速率季节变化的主要影响因子,以期为区域森林采伐对土壤呼吸速率的影响研究提供科学依据。【方法】以闽北杉阔混交人工林为研究对象,2011年8月实施了不同蓄积量采伐强度(中度择伐34.6%、强度择伐48.6%、极强度择伐67.6%、皆伐)作业试验,并与未采伐对照;2016年7月—2017年7月运用Li-8100 A土壤碳通量自动测量系统,对土壤及其各组分的呼吸速率、土壤5 cm深处的温度和湿度开展了为期1年的定位观测。【结果】未采伐和各种强度择伐5年后,土壤总呼吸速率最大值都出现在7月份,最小值出现在1—3月份;皆伐5年后,土壤总呼吸速率最大值出现在6月份,最小值出现在11月份;各种强度采伐林地的矿质土壤呼吸速率与未采伐林地无显著差异( P >0.05);各种强度择伐林地的凋落物和根系呼吸速率都与未采伐林地无显著差异( P >0.05),而皆伐林地的凋落物和根系呼吸速率都显著低于未采伐林地( P <0.05),分别比未采伐林地(1.45和1.11 μmol ·m^-2 s^-1 )减少了0.93和0.53 μmol ·m^-2 s^-1;各种强度择伐林地的土壤总呼吸速率与未采伐林地无显著差异( P >0.05),而皆伐林地的土壤总呼吸速率显著低于未采伐林地( P <0.05),比未采伐林地(4.39 μmol ·m^-2 s^-1 )减少了1.64 μmol ·m^-2 s^-1;中度、强度和极强度择伐林地5 cm深处的土壤温度与未采伐林地没有显著差异( P >0.05),而皆伐使林地土壤温度显著升高( P <0.05),比未采伐林地(18.52 ℃)增加了4.7 ℃;中度、强度择伐林地的5 cm深处土壤湿度与未采伐没有显著差异( P >0.05),而极强度择伐和皆伐使林地土壤湿度显著降低( P <0.05),分别比未采伐林地(30.67%)减少了2.17%和3.98%;土壤总呼吸速率的土壤温度指数模型拟合效果最优,能解释未采伐和各种强度择伐林地土壤总呼吸变化的77.8%~83.3%以及皆伐林地土壤呼吸变化的35.5%;未采伐、中度、强度和极强度择伐林地土壤总呼吸的温度敏感性参数Q 10 为1.77~2.72,皆伐林地的 Q 10 为1.49。【结论】不同强度采伐5年后,各种强度择伐林地土壤及其各组分的呼吸速率与未采伐林地没有显著差异;皆伐使凋落物呼吸速率、根系呼吸速率和土壤总呼吸速率都显著降低;各种强度择伐没有改变土壤总呼吸速率的季节变化规律,但皆伐使土壤总呼吸速率最大和最小出现时间有所提前;研究区土壤温度是土壤总呼吸速率季节变化的主要影响因子。     

Effects of vegetation control on nutrient removal and Fagus orientalis, Lipsky regeneration in the western Black Sea Region of Turkey

Dense Rhododendron ponticum (L.) understories of eastern beech (Fagus orientalis Lipsky) stands in the Black Sea Region (BSR) of Turkey create challenging forest vegetation management problems relative to beech regeneration. Rhododendron traditionally is controlled in Turkish forests with grubbing and bulldozing. The effects of these practices on nutrient removal and natural beech regeneration have not been quantitatively studied. Two woody vegetation control treatments (bulldozing and hand-grubbing) were installed during late summer, 2002 in three mature beech stands with dense rhododendron understories in the Düzce Forest Management Directorate, in the Turkish western BSR. Aboveground biomass of each vegetation component, total aboveground vegetation biomass, nutrient concentrations, organic matter (OM) removal, and total amount of OM nutrients were determined for each woody vegetation control treatment. Soil bulk density and nutrient content, and beech seedling biomass, nutrient content, and natural regeneration also were studied. One year after treatment, the machine site preparation by bulldozing (MSP) that removed understory vegetation and attached roots, reduced mean forest floor OM content by about 84%, when compared to hand-grubbing. Mean soil C, N, K and Mg concentrations on the bulldozed sites were 36, 27, 50 and 55% less, respectively, than those on the grubbed sites. Total C and Mg amounts at the 5–10 cm soil depth were 24 and 47% lower, respectively, for mechanical site preparation (MSP) sites, when compared to grubbed sites. Overall, soil bulk density did not differ significantly between the grubbing and MSP treatments. Frequent passes on designated transects on MSP sites resulted in a significant (P < 0.01) one-third increase in bulk density of the first 20 cm soil depth, when compared to grubbed sites. No natural beech regeneration occurred on untreated control sites. Mineral soil exposure on the MSP sites increased beech seedling germination substantially. One year after treatment, the mean number of naturally regenerated seedlings for MSP sites was 2.5 times greater than for grubbed sites. Woody control treatments had significant effects upon beech seedling chemistry, with N, P, K, Ca and Mg concentrations averaging 35, 47, 12, 33 and 25% lower, respectively, for MSP sites, when compared to grubbed sites. However, mean seedling biomass and nutrient content were significantly greater (P < 0.05) on MSP sites. Long-term effects from windrowing on MSP site residues, associated topsoil and nutrients need to be evaluated.     

Biomass and carbon pools of disturbed riparian forests

Quantification of carbon pools as affected by forest age/development can facilitate riparian restoration and increase awareness of the potential for forests to sequester global carbon. Riparian forest biomass and carbon pools were quantified for four riparian forests representing different seral stages in the South Carolina Upper Coastal Plain. Three of the riparian forests were recovering from disturbance (thermal pollution), whereas the fourth represents a mature, relatively undisturbed riparian forest. Above and belowground carbon pools were determined from linear transects established perpendicular to the main stream channels and spanning the width of the riparian area. The objective of this study was to quantify the biomass and carbon pools in severely disturbed, early successional bottomland hardwood riparian forests and to compare these values to those of a less disturbed, mature riparian forest.

Aboveground biomass in all four riparian forests increased during the 2.5-year investigation period. The total carbon pool in these South Carolina Coastal Plain riparian forests increased with forest age/development due to greater tree and soil carbon pools. The mature riparian forest stored approximately four times more carbon than the younger stands. The importance of the herbaceous biomass layer and carbon pool declined relative to total aboveground biomass with increasing forest age. As stands grew older fine root biomass increased, but an inverse relationship existed between percentages of fine root biomass to total biomass. The root carbon pool increased with forest age/development due to a combination of greater fine root biomass and higher root percent carbon.

Aboveground net primary production (NPP) in young riparian forests rapidly approached and exceeded NPP of the more mature riparian forest. As a woody overstory became established (after 8–10 years) annual litterfall rate as a function of NPP was independent of forest age and litterfall amount in the young riparian forests was comparable to mature riparian forests. Biomass in the riparian forest floor and carbon pool declined with increasing riparian forest development. Woody debris in these riparian forests comprised a relatively small carbon pool. An understanding of bottomland hardwood riparian forest carbon pools at different stages of succession allows us to assess how time since disturbance influences these pools, leading to a better understanding of the recovery processes.     

Thinning, burning, and thin-burn fuel treatment effects on soil properties in a Sierra Nevada mixed-conifer forest

More than a century of fire exclusion and past timber management practices in many Sierra Nevada mixed-conifer forests have led to increased stand densities and fuel accumulation, with a corresponding risk of large, high severity wildfires. To reduce hazardous fuel accumulations and restore the health and natural processes of forest ecosystems, fuel management programs often employ thinning and prescribed fire treatments, both alone and in combination. We evaluated forest floor and mineral soil chemical and physical characteristics following these treatments in a managed Sierra Nevada mixed-conifer forest using a fully replicated study design with four separate treatments: THIN, BURN, THIN + BURN, and an untreated CONTROL. Compared to the CONTROL, the BURN and THIN + BURN treatments consumed a large amount of the forest floor, reducing the mass and depth by more than 80%. These treatments reduced the forest floor C and N pools by more than 85%, resulting in reductions of 25 Mg C ha⁻¹ and more than 700 kg N ha⁻¹ from the forest floor. Despite these large losses from the organic horizons, no significant differences in mineral soil total C and N pools were detected among treatments. Compared with the CONTROL and THIN treatments, the BURN and THIN + BURN significantly increased the mineral soil NO₃-N concentration, pool of inorganic N, pH, and exposed bare soil. The THIN + BURN treatment significantly increased the concentrations of NH₄-N and exchangeable Ca relative to the CONTROL. No significant differences in the net rates of nitrification, N mineralization, or bulk density were detected among the four treatments. The BURN treatment reduced mineral soil C concentration and CEC, while the THIN + BURN treatment had the greatest increase in inorganic N. Fire effects on soil pH and inorganic N were moderated in skid trails due to reduced fuel continuity and consumption. In light of the current management emphasis on hazardous fuels reduction, we recommend that researchers investigating fire effects in harvested stands include skid trail influences in their study design.     

Sedimentation associated with forest road surfacing in a bottomland hardwood ecosystem

Access systems are a necessary element of resource production in bottomland hardwood sites. However, road building may have a detrimental effect on hydrologic function of the site. This report describes initial results of a study designed to examine the effect of different road surfacing treatments on water quality.

Four surfacing treatments installed on two test roads included native soil, native soil with vegetative stabilization, 6 cm of gravel, and 15 cm of gravel over geotextile. During the first flooding season periodic sampling measured floodwater suspended sediments and location of erosion and sediment deposition within the road prism. Initial results suggest that sediment movement was confined to the road right-of-way, with no statistically significant sedimentation effects detected beyond the clearing limits of the road. The study is continuing for another field season.