闽西马尾松实生种子园高产家系选择研究

对闽西马尾松实生种子园143个选优家系子代测定林调查分析表明,家系间结实量存在极显著差异,经过多重极差检验分析,我们筛选出与对照漳平五一林场种子园种子有极显著差异以上的最优高产家系42个,进一步比较结果表明,既高产又速生的优良家系26个,其结实量平均遗传增益为40.41%,材积平均遗传增益为26.36%,这26个优良家系可供建下一代马尾松种子园的最好建园材料。     

马尾松实生种子园种子增产效果分析

通过对马尾松实生种子园种子增产效果的分析,结果表明：5年生林分的平均树高、胸径、材积比当地一般林分种苗造林分别增长13.71%、19.66%、47.68%;11年生林分分别增长8.81%、10.19%、24.14%。马尾松改良采用营建实生种子园的改良技术可行,增产效益明显。     

Mechanizsed microsite preparation and direct seeding of Pinus sylvestris in boreal forests — a way to create desired spacing at low cost

The main objective of this study was to examine the cost, flexibility, and appropriate scale of mechanized microsite preparation (MP), in combination with mechanical direct seeding of Pinus sylvestris L. with orchard seed. This technique was tested at four boreal forest sites in Northern Sweden. Orchard and stand seeds were sown with and without MP. The use of orchard seed increased seedling establishment by 41% and the use of MP increased seedling establishment by 47%, respectively, after two years. The best substrates for sowing when using MP were OA_h-, E- and BC-horizon, in ranked order. The use of orchard seed compared to stand seed increased mean seedling height by 25% after four years. These trials suggest that to obtain a density of 5,000 stems ha^-1 four years after seeding, 61,000 viable stand seeds or 41,000 orchard seeds ha^-1 should be sown if MP is not used. If MP is used, seeding rate could be reduced by about 32%. By using MP, and by further improving scarification technique so that all scarified area is thin OA_h-horizon, we predict that only 32,000 stand seeds or 22,000 orchard seeds ha^-1, i.e., half the dosage, should be needed. Under these optimal conditions, it would be necessary to sow about six and four germinable stand and orchard seeds, respectively, to ensure one seedling after four years. Furthermore, regeneration cost would be less than a third that of planting.     

Mating System and Pollen Contamination in a Pinus brutia Seed Orchard

Mating system parameters and pollen contamination were estimated in an 11-year-old Pinus brutia Ten. clonal seed orchard, located in Antalya, Turkey, with the aid of isozymes. Isozyme analysis was performed on both maternal (megagametophyte) and embryo tissues of seeds collected from the seed orchard and the surrounding natural stand. Fourteen loci encoding nine enzyme systems were assayed. Based on a multilocus estimator, the proportion of viable seeds originating from outcrossing (tˆ_m) in the seed orchard was found to be 0.947; the remainder (0.053) was due to selfing. We estimated that 85.7% of the orchard seed resulted from pollen contamination by trees in surrounding stands. This may be related to the young age and subsequent low pollen production of the orchard. Only 9% [i.e., 1.0−0.857 (contamination)−0.053 (selfing) = 0.090] of the orchard offspring was due to cross-pollination among clones within the orchard. Due to the high level of pollen contamination, it is estimated that genetic gain in this seed crop is less than 57% of that expected if all offspring resulted from random mating among orchard clones (i.e., no contamination).     

STUDY ON THE HALF-SIB FAMILY SELECTION FOR THE SUPERIOR TREE OF SCOTS PINE

Scotspine(PinussylvesIrisvar.mongoIica)isoneofthemaintreespeciesofsilviculture,shelterbeltsandgreeningbeltsinthenortheastofP.R.China.Asitisresistanttocoldanddry,notovercriticalaboutsoilmoisture,stronginrootsystemetc.,itwasafforestedinmanyregionsinChinarecently.Inordertoimprovescotspinefurther,makefulluseofthesuperiorgeneresources,wemustdotheworkofitsprovenanceandprogenytesttodistinguishindividualsandpopulationaccordingtothephenotypeofitsprogenies,andtose-lectfineprovenanceandfamilyandindi-vi…     

Effect of thinning intensity on tree growth and temporal variation of seed and cone production in a Pinus koraiensis plantation

Thinning of Korean pine (Pinus koraiensis Sieb. et Zucc.) is used to facilitate timber and cone production. The present study in Northeast China investigated the effects of thinning intensity on individual tree growth, temporal variation in cone yield, and seed quality in Korean pine plantation. In 2005, five thinning intensity levels (none, extreme, heavy, moderate and light) were set in 15 permanent plots in a 32-year-old Korean pine plantation at Mengjiagang Forest Farm, Jiamusi City, Heilongjiang Province. We recorded tree growth and seed cone production from 2013 to 2016, i.e., from 8 to 11 years after thinning. Except for height growth, thinning increased tree growth (diameter at breast height and crown size) and improved cone yield. The extreme thinning treatment (to 300 trees per hectare) resulted in the largest tree diameter, tree volume, crown size and 4-year cone production per tree. The highest cone yield per tree in the mast year (2014) was observed when stands were thinned to 500 trees per hectare (heavy thinning). Although the best cone and seed quality and the largest cone and seed mass per tree were recorded in the heavily thinned stand, no significant differences were found between heavy and moderate thinning stands (750 trees per hectare). At the stand level, the moderately thinned stand had the highest basal area, stock volume and seed cone production per stand. Our results suggest that thinning to 750 trees per hectare will improve timber and cone productivity in 40-year-old P. koraiensis stands.     

马尾松2个世代种子园6年生家系生长的遗传变异与增益比较

【目的】研究与评价马尾松一代、二代无性系种子园的遗传增益及改良效果,指导马尾松良种选育、生产及应用。【方法】利用营建在浙江省淳安县燕山林场和福建省邵武市卫闽林场的6年生马尾松一代和二代自由授粉家系遗传测定林,分析不同世代种子园子代生长性状的遗传变异,并进行家系育种值估算及生长评价。【结果】马尾松不同世代的种子园子代生长差异很大,其树高、胸径和单株材积等生长量显著大于马尾松优良天然林分的混系子代,其中一代种子园子代各家系单株材积表型值和育种值分别大于优良天然林分混系55.78%~84.62%和40.66%~71.39%,二代其值分别大于优良天然林分混系58.96%~108.97%和42.97%~91.89%,二代种子园子代的平均生长量和遗传稳定性总体上高于一代种子园子代。树高、胸径和单株材积等主要生长性状在世代内家系间的生长差异均达极显著水平,在一代种子园子代中最优家系的树高、胸径和单株材积分别大于最差家系21.94%、30.72%和97.75%,在二代种子园子代中最优家系的树高、胸径和单株材积分别大于最差家系23.42%、29.60%和90.09%,说明种子园中对亲本材料的再选择非常重要。依据子代的单株材积育种值在两测定点的位次及其变幅大小,将38个家系分成速生型、较速生稳定型、较速生不稳定型和慢生型4类,各类所占家系数分别有23.68%、26.32%、21.05%和28.95%,其中速生型和较速生稳定型家系所对应的亲本无性系可作为新建马尾松二代无性系种子园的优良建园材料。种子园子代主要生长性状的地点×家系、区组×家系的互作效应极显著,说明各家系在不同立地环境中生长表现不一致。【结论】马尾松无性系种子园良种效益显著,二代增益大于一代。通过选择速生型和较速生稳定型家系将有助于种子园建园亲本的再选择、现有种子园的留优去劣疏伐和重建新的种子园。鉴于子代主要生长性状的基因型与环境互作使各家系在不同立地环境中生长表现不同,建议对子代生长表现突出、结实量较多的无性系进行单系采种利用。     

杉木种子园球果病虫害监测研究

经调查统计,杉木种子园健康果、发育不良果、病害果、虫害果、病虫兼有果比例分别为55.1%、3.4%、8.4%、9.0%、24.1%,各无性系之间球果产量、不健康果比例,百果重、千粒重、出粒率、发芽率都达到极显著差异水平。球果病虫危害率与球果产量、球果出籽率、种子发芽率之间存在极显著负相关,病虫危害是种子园种子产量下降的主要原因之一。用1%波尔多液或500倍敌克松溶液喷洒种子园接株,效果十分理想,病虫危害可以控制在10.0%以内。     

台山红岭种子园湿地松基因资源引入效果分析

１９８９年至１９９６年间,台山红岭种子园从美国、澳大利亚、津巴布韦引入湿地松基因资源种子共４０７份,包括有６个种源、２１３个全同胞家系和１８８个半同胞家系。利用其中３７０份材料营建种源林１片、基因资源林６片,面积共１６．８５ｈｍ＾２。根据这７片测定林的调查数据分析了新引材料的遗传品质以及选择潜力,造林保存率平均达９０％以上;在生长量方面,有６６．６７％的种源、８２．１４％的家系优于台山湿地松初级种     

马尾松一代无性系种子园营建与丰产技术研究

本研究历经“六五”选优,“七五”建园与“八五”投产,长达工5年之久的科学研究与生产实践,比较系统地阐述了马尾松一代无性系种子园的营建、管理与种子丰产技术。“八五”期间种子园开始生产种子,产量逐年上升,1995年大面积平均每公顷产量为9.375kg,最高小区平均每公顷产量达27.825kg,属于马尾松投产初期高产种子园之列。种子园混系种子质量优异,5年生子代林材积增益比对照大41.4％。种子园的建成与投产,可为速丰林建设提供优质良种。     

Effects of Seed Weight and Seed Type on Early Seedling Growth of Pinus sylvestris under Harsh and Optimal Conditions

The effects of seed weight and seed type on seedling growth of Pinus sylvestris (L.) were studied by seeding individually weighed orchard and stand seed in different mixtures under harsh (direct seeding in field) and optimal (seeding in nursery) conditions. In the nursery experiment an increase in the seed weight from 3 to 7 mg increased the seedling height by 10-27% and total weight by 27-113%, and decreased the height/diameter ratio by 5-6% after 2 yrs. With elimination of seed weight effects, orchard seedlings were 2% taller than stand seedlings in year 2. Without elimination of seed weight effects, orchard seedlings were 7-13% taller. In the field experiment an increase in the seed weight from 3 to 7 mg increased seedling height by 18-65%, stem volume by 81-274% and the number of top-buds by 23-34% in year 5. After elimination of seed weight effects, orchard seedlings were 7-13% taller than stand seedlings and without elimination of seed weight effects 20-21% taller after 5 yrs. Even after elimination of both seed weight and genetic effects orchard seedlings were 3-9% larger than stand seedlings in the field experiment. In conclusion, the influence of seed weight and seed type on growth traits and slenderness is highly significant and the influence seems to be greater in harsh conditions.     

Autumn frost hardiness of Pinus sylvestris offspring from seed orchard grafts of different ages

Autumn frost hardiness of offspring from clones (originating from 66.0–68.8° N) in six southern (62° N) and one northern (66.5° N) Pinus sylvestris seed orchards in Finland, was evaluated. One‐year‐old seedlings derived from open pollinated grafts of young, intermediate, and fairly mature age, and natural stand seedlings, were artificially freeze tested. The northerly located orchard produced the least injured offspring. There were no significant systematic differences between age classes. Neither were there any indications of decreased freezing injury with later age classes. The family variance was significant in four orchards. The estimated injury levels of orchard offspring were, in general, closer to those of natural stands in the neighbourhood of the orchard than natural stands at the place of clonal origin. It is suggested that not only differences in mother clone hardiness but also clonal differences in pollen contamination rate caused the observed family variation.     

Natural Variation in Seed Characteristics of Eastern White Pine (Pinus strobus L.)

Natural variation in seed characteristics of eastern white pine (Pinus strobus L.) trees and its relationship with the size, form, and crown class of parent trees was examined in second-growth, white pine mixedwood stands in Ontario. Cones were collected prior to and following partial harvesting of two stands during 2 years of above-average seed production. Fully enlarged seed extracted from mature cones was X-rayed to determine seed yield per cone, the percentage of sound, partially filled, and empty seed, and the sound seed mass for each tree. Variation in seed yield per cone, the proportion of sound, partially filled, and empty seed, and seed mass was much greater among individuals than between stands or seed years. The strength and significance of the relationship of seed mass with sound seed yield differed with stand and seed year. In uncut stands, seed mass was higher for trees with dominant crown class, larger live crown ratios, and smaller height to diameter ratios, suggesting tree vigour and microsite quality influence seed mass. Proportion of partially filled seeds was significantly reduced in both stands in the second year likely due to increased growing degree days during female gametophyte development. The implications of these results to natural and artificial regeneration of white pine are discussed.     

Flowering and Airborne Pollen Occurrence in a Pinus sylvestris Seed Orchard Consisting of Northern Clones

To explain the high background pollination rate detected in earlier studies, this study investigated the flowering phenology of a Scots pine seed orchard with respect to airborne pollen occurrence during 1992 and 1993. The receptive period of the female strobili and the shedding period of the male strobili were observed visually. Airborne pollen was monitored with Rotorod traps located inside and outside the seed orchard. There was no time gap in the onset of flowering between the earliest female and male strobili in 1992. Female flowering began 3 days earlier than male flowering in 1993, and the earliest strobili were probably pollinated by pollen transported long distance from more southerly areas. The almost simultaneous peak in pollen occurrence inside the seed orchard and in surrounding pine stands indicates that the seed orchard clones are not temporally isolated from sources of contamination. However, pollen concentrations inside the seed orchard were higher than those in the surrounding stands. Therefore, flowering phenology cannot sufficiently explain the observed pollen contamination rates in this seed orchard.     

Phenotypic changes in progenies of northern clones of Picea abies (L.) Karst. grown in a southern seed orchard

Results are presented from three different field trials comparing controlled crossed materials of Norway spruce from the southern seed orchard at Lyngdal (58° N) with open‐pollinated progenies from the same mother trees standing in the northern natural forest (63–66° N). The seed orchard progenies flushed consistently later in the spring, terminated leader shoot growth later in the summer, had higher frequencies of lammas shoots, were delayed in lignification during autumn, and were 15% taller at age seven years from seed. The difference between the orchard‐ and the natural stand material appeared to be permanent from age four to age seven. It is suspected that the non‐native environment in the seed orchard could affect the genotypic performance of the seed orchard progenies.     

初论杉木种子园的园址选择

通过6个省内28个种子园的数据分析,发现位于杉木不同产区(北部产区、中心产区、南部产区)的种子园,其开花结实年龄、结实量、种子播种品质等均存在较有规律的差异。中心产区无性系一般开花结实迟于北部产区的无性系;产量和种子品质的年变幅大;历年平均产量、球果出籽率和种子发芽率显著低于北部产区。就上述趋势对影响杉木种子生长发育的主要环境因子进行了理论上的讨论分析,并提出杉木种子园园址选择的初步意见。     

Ground seed density patterns under conditions of strongly overlapping seed shadows in <Emphasis Type="Italic">Abies alba</Emphasis> Mill. stands

In three subsequent years, seeds were collected from 66 to 75 seed traps in three Abies alba stands that differed in vertical structure and the number of potentially reproducing trees. The objective was to compare empirical patterns of seed density with a random (Poisson) model, which assumes that seed density would be the same in every location, and a binomial negative model that is appropriate for over-dispersed data. The seed density patterns were tested for spatio-temporal independence. The effect of some dispersal variables (number of mother trees and seed fall seasons, dispersal distance, fecundity rate) on seed density pattern was studied in a simple simulation experiment. Several local stand density measures (stem density, basal area, distance-weighted indices, canopy openness) were also tested for their ability to predict local seed density. The Poisson model was rejected in all the stands studied, and a much better fit achieved using the binomial negative model. The simulation study revealed that random seed distribution may occur only at considerable dispersal distances, concave seed shadows, and at very low fecundity rates. The seed density patterns identified exhibited both spatial and temporal correlation, which can probably be linked to the arrangement of mother trees and variation in their seed output. To a minor degree, this seed density pattern also correlated with local stand density.     

普文试验林场思茅松无性系种子园营建技术

１９８８～１９９５年在云南省林业科学院普文试验林场建成了全国第一个思茅松无性系种子园,面积３３３ｈｍ２。本文应用思茅松天然优良林分选择的研究成果,在中选的优良林分中,以５株优势木法、指数法为评选优树的方法,详细介绍了思茅松无性系种子园建园材料的选择标准和方法。以普文思茅松无性系种子园建设为例,从园址选择、种子园区划、苗木培育、优树收集区和种子生产区的建立方面系统地介绍了思茅松无性系种子园的建园技术及方法。并就如何通过半同胞子代测定评估优良家系的遗传增益,以及建立展示林、试验林等作了论述。普文思茅松无性系种子园的建设经验,可供今后继续扩大建园参考。     

Optimal management of Korean pine plantations in multifunctional forestry

Korean pine is one of the most important plantation species in northeast China.Besides timber,it produces edible nuts and plantations sequester carbon dioxide from the atmosphere.This study optimized the management of Korean pine plantations for timber production,seed production,carbon sequestration and for the joint production of multiple benefits.As the first step,models were developed for stand dynamics and seed production.These models were used in a simulation–optimization system to find optimal timing and type of thinning treatments and optimal rotation lengths.It was found that three thinnings during the rotation period were optimal.When the amount or profitability of timber production is maximized,suitable rotation lengths are 65–70 years and wood production is 5.5–6.0 m~3 ha~(-1) a~(-1).The optimal thinning regime is thinning from above.In seed production,optimal rotation lengths are over 100 years.When carbon sequestration in living biomass is maximized,stands should not be clear-cut until trees start to die due to senescence.In the joint production of multiple benefits,the optimal rotation length is 86 years if all benefits(wood,economic profits,seed,carbon sequestration) are equally important.In this management schedule,mean annual wood production is 5.5 m~2 ha~(-1) and mean annual seed yield 141 kg ha~(-1).It was concluded that it is better to produce timber and seeds in the same stands rather than assign stands to either timber production or seed production.