The vertical distribution of fine roots of five tree species and maize in Morogoro,Tanzania

In order to assess the possibility of root competition in agroforestry, the vertical distribution of fine roots (< 2 mm in diameter) of five tree species in pure two-year-old stands was compared to that of mature maize.Cassia siamea, Eucalyptus tereticornis, Leucaena leucocephala andProsopis chilensis had a rooting pattern similar to that of maize, i.e. a slow decline in fine root mass from 0–100 cm soil depth.Eucalyptus camaldulensis had its roots evenly distributed down to 100 cm. On an average, the fine root biomass of the tree species was roughly twice as that of the maize. We conclude that the studied tree species are likely to compete with maize and other crops with a similar rooting pattern for nutrients and water.     

The vertical distribution of fine roots of five tree species and maize in Morogoro,Tanzania

In order to assess the possibility of root competition in agroforestry, the vertical distribution of fine roots (< 2 mm in diameter) of five tree species in pure two-year-old stands was compared to that of mature maize.Cassia siamea, Eucalyptus tereticornis, Leucaena leucocephala andProsopis chilensis had a rooting pattern similar to that of maize, i.e. a slow decline in fine root mass from 0–100 cm soil depth.Eucalyptus camaldulensis had its roots evenly distributed down to 100 cm. On an average, the fine root biomass of the tree species was roughly twice as that of the maize. We conclude that the studied tree species are likely to compete with maize and other crops with a similar rooting pattern for nutrients and water.     

Observations on the rooting patterns of some agroforestry trees in an arid region of north-western India

The present study deals with root architecture of 6-year-old trees of 9 indigenous and 3 exotic species growing in arid climate of north-western India. Observations, made on excavated root systems (3 tree replicates of each species) showed large variation in horizontal and vertical spread of roots. In Morus alba, Melia azedarach and Populus deltoides, the roots were confined to 80 cm, while in Prosopis cineraria, Acacia nilotica and Eucalyptus tereticornis, roots penetrated more deeply to 233 cm. The number of total roots ranged from 103 in Acacia catechu to 1932 in Eucalyptus tereticornis, and 62 to 80% of the roots were less than 2 mm in diameter. The primary roots were more horizontal than the secondary roots. The total root biomass varied from 2.2 kg in Acacia catechu to 30.6 kg^–1 tree in Populus deltoides, and top 30 cm soil contained 42 to 78% of the total biomass. The implications of the results are discussed in the context of the ecological niche of the species, and its usefulness in agroforestry systems.     

Nitrogen fixation in L. leucocephala L. as affected by rooting volume and competition with E. camaldulensis

Fast growing trees, in particular nitrogen fixing trees, are essential for the establishment of sustainable agroforestry practices by resource-poor and/or environmentally conscious aarmers. An important selection factor for trees is therefore high N₂ fixing capability. This study was conducted to examine biological nitrogen fixation (BNF) in Leucaena leucocephala L., as affected by growth in pots containing 4, 10, 15 and 30 kg soil and when grown together with Eucalyptus camaldulensis. BNF was measured by the ¹⁵N isotope dilution technique. Leucaena and Eucalyptus were grown either sole (SL, SE) or in two mixtures (1:1 - L1:E1 and 1:2 - L1:E2) for a period of 6 months. Planting pattern influenced (P < 0.05) dry matter and N content of shoot and roots of Eucalyptus, as the sole planted trees gave the highest yield. Dry matter, N content and total amount of N fixed by Leucaena significantly decreased with decreasing rooting volume. Also, the smaller the rooting volume (or soil weight), the higher was the proportion (%) of N derived from the atmosphere (%Ndfa) by leucaena. For each rooting volume, the sole or mixed planting pattern did not significantly influence either the proportion (%) or amount of N fixed by leucaena. For these trees, therefore, rooting volume rather than cropping pattern influenced the proportion and amount of nitrogen fixed.     

Soil andLeucaena leucocephala L. growth variability underFaidherbia albida Del. andZiziphus spina-christi L. Desf.

In order to obtain data which will facilitate the scientific development of multiple intercropping of trees in agroforestry systems, the effects ofFaidherbia albida Del. orZiziphus spina-christi L. Desf. on soil quality and on yield of interplantedLeucaena leucocephala L. were investigated. The distance to which bothFaidherbia albida andZiziphus spinachristi trees improved soil quality as well asLeucaena leucocephala andSorghum vulgare L. production around them was about two times the radius of the tree crown. Higher yields of plants underZiziphus spina-christi thanFaidherbia albida were explained by similarly higher concentrations of available phosphorus (P). Other soil parameters, including total nitrogen (N), were similar under both species; being higher under the tree canopies than away from them. The study identified optimum tree spacing for agroforestry to be twice the tree crown radius. It also demonstrated the advantages of multiple tree mixtures over pure stands in improving soil quality and productivity.     

Biomass production of trees and grasses in a silvopasture system on marginal lands of Doon Valley of north-west India 2. Performance of grass species

In a long-term study (1980 to 1993), the survival, growth performance and biomass production of two grass species viz. Chrysopogon fulvus (Spreng.) Chiov. and Eulaliopsis binata (Retz.) C.E. Hubb. were assessed when intercropped with four tree species viz. Albizia lebbek (L.) Benth., Grewia optiva Drumm., Bauhinia purpurea L. and Leucaena leucocephala (Lamk.) de Wit. on the bouldery riverbed lands of Doon Valley of north-west India. The survival of grass clumps was higher under the canopy of B. purpurea (48.3% in 1993 of the initial planting density of 20,000 clumps ha⁻¹ in 1980) and G. optiva (47.4%) than of L. leucocephala (31.1%) and A. lebbek (29.4%), and at 75% intensity of tree lopping (40.9%) than at 50% (37.1%). The effect of tree species on different growth parameters of grasses was not uniform, with G. optiva and B. purpurea causing comparatively more synergistic effect on clump height and clump diameter respectively. Although the performance of grasses was good under A. lebbek and L. leucocephala in the early years (1980--1987), these tree species resulted in the lowest grass dry weight in the later years up to 1993. The beneficial effect of higher lopping intensity of 75% was observed on the growth and biomass production of grasses over 50% lopping, presumably due to increased light penetration into the under-storey. The biomass production of grasses reached a maximum at four years of growth in 1983, after which, it decreased gradually to less than half in 1993. Eulaliopsis binata showed higher survival, growth and biomass production than C. fulvus throughout the period of study. The total biomass production was highest in association with B. purpurea followed by G. optiva, which appeared to be the most suitable tree species along with E. binata for sustainable silvopasture development on the marginal lands of Doon Valley of north-west India. This revised version was published online in June 2006 with corrections to the Cover Date.     

Moisture extraction and crop yield as a function of distance from a row of Eucalyptus tereticornis

A study was made on the effect of an east-west tree line of years old Eucalyptus tereticornis on soil physical parameters and adjoining mustard and wheat crops as a function of distance from the tree-crop interface in north- and south-facing sides. Maximum soil temperature, pan evaporation, moisture extraction, number of plants/m², and yields of mustard and wheat did not differ significantly on the two sides of plantation. Plant growth parameters decreased linearly with distance from the plantation. Mustard yield, Y (kg/ha) was linearly related to moisture extraction, ME (mm) as Y = 1914–120.3 ME. Eucalyptus tereticornis was found to extract 5 times more water than mustard from 0–150cm soil profile. Mustard and wheat yield reductions in adjoining 10 meter strips on both sides of the plantation were 47 and 34 per cent, respectively. Eucalyptus tereticornis is, therefore, not a suitable species for row plantations in agro-forestry systems in deep water table conditions of semi-arid regions.     

Benefit-cost analysis of selected agroforestry systems in Northeastern Thailand

Potential productivity and financial returns from selected agroforestry systems and traditional monocrops located in the Phu Wiang watershed were estimated from limited trials of cropping alternatives using cost-benefit analysis. The agroforestry systems studied consisted of combinations of Eucalyptus (Eucalyptus camaldulensis), Leucaena (Leucaena leucocephala), or Acacia (Acacia auriculiformis) inter-cropped with cassava (Manihot esculenta) or mungbean (Vigna radiata). Evidence from trials at short, 3-year rotations, demonstrate that early supplementary and complementary relationships between some system components can imply synergistic financial gains. Although these biological interactions turn competitive over time, in this case, the gains should be sufficient to make early adopters consider the agroforestry systems financially preferable to traditional monocrops.     

Effect of trees on the yield of wheat crop

A study was conducted at the National Agricultural Research Center (NARC) of Pakistan Agricultural Research Council (PARC) by planting trees of four different species: Eucalyptus camaldulensis, Albizia procera, Morus alba and Leucaena leucocephala along the boundary of wheat fields in a randomized complete block design. Data on crop yield for each tree species and control (no trees) at different distances viz 2, 4, 6, 8, 10 and 12 meter (m) from the tree bases and control were collected and analyzed. The statistical analysis did not show any significant difference in the wheat yield among different tree species. However, the wheat yield was numerically lowest at 2 m distance in case of all the four tree species and control. In case of mulberry, it was lowest statistically also from other distances. Numerically higher wheat yield values were noted at later distances (8, 10 and 12 m) in case of all tree species including control except for siris where numerically highest value was found at 6 m distance. Therefore, it can be generalized that tree's impact on wheat yield can be experienced up to 2 m distance, there is little, if any, impact up to 6 m distance and almost no impact at 8, 10 and 12 m distances.     

Woody legumes as live support systems in yam cultivation

This paper reports on the use of three leguminous perennials (Leucaena leucocephala, Flemingia macrophylla and Gliricidia sepium) as live support systems in yam cultivation. In a planting arrangement in which yam rows alternated with rows of the woody species the tuber yields per ha were 3.4 (leucaena), 5.3 (flemingia) and 10.1 (gliricidia) ton fresh weight. TUKEY's L.S.D. value equalled 2.9 ton. Leucaena leucocephala is unsuitable as live support since the species shows a strong competitive power expressed in terms of leaf productivity and relative density of the root mass in the upper soil stratum, the zone explored by the yam crop. Flemingia macrophylla is unsuitable as support species mainly because of its structural weakness: Its branches do not sufficiently lignify to carry the yam leaf mass. The significantly higher tuber yield of the yam crop grown with Gliricidia sepium is a function of specific properties of the tree species: Low leaf productivity, a relatively weakly developed root system and an open architecture, which leaves sufficient space for a yam crop grown in association.     

Fine-root dynamics of coffee in association with two shade trees in Costa Rica

Fine-root dynamics (diameter < 2.0 mm) were studied on-farm in associations of Coffea arabica with Eucalyptus deglupta or Terminalia ivorensis and in a pseudo-chronosequence of C. arabica-E. deglupta associations (two, three, four and five years old). Coffee plants were submitted to two fertilisation types. Cores were taken in the 0–40 cm soil profile two years after out-planting and subsequently in the following year in depth layers 0–10 and 10–20 cm, during and at the end of the rainy season, and during the dry season. Fine root density of coffee and timber shade trees was greater in the coffee fertilisation strip as compared to unfertilised areas close to the plants or in the inter-rows. Coffee fine roots were more evenly distributed in the topsoil (0–20 cm) whereas tree fine roots were mostly found in the first 10 cm. Although the two tree species had approximately the same fine root length density, lower coffee / tree fine root length density ratios in T. ivorensis suggest that this shade tree is potentially a stronger competitor with coffee than E. deglupta. Coffee and tree fine root length density for 0–10 cm measured during the rainy season increased progressively from two to five-year-aged associations and coffee fine root length density increased relatively more than E. deglupta fine root length density in the four and five-year-aged plantations suggesting that contrary to expectations, coffee fine roots were displacing tree fine roots.     

Biomass production of trees and grasses in a silvopasture system on marginal lands of Doon Valley of north-west India. 1. Performance of tree species

Bouldery riverbed lands occupy about one-third of the geographical area in Doon Valley of north-west India. These marginal lands need rehabilitation through establishment of silvopasture system for meeting the biomass requirements of resource-poor farmers. A study was carried out for 14 years (1980 to 1993) at Dehradun, involving four multi-purpose tree species viz. Albizia lebbek (L.) Benth., Grewia optiva Drumm., Bauhinia purpurea L. and Leucaena leucocephala (Lamk.) de Wit.; and two grass species viz. Chrysopogon fulvus (Spreng.) Chiov. and Eulaliopsis binata (Retz.) C.E. Hubb. Fifty and 75% of the tree crown was lopped once a year after four years of age. Tree survival decreased with age, and was highest with L. leucocephala (86.9% in 1993 of initial density of 625 trees ha⁻¹) and lowest with B. purpurea (49.3%). Canopy parameters of trees progressed differently, presumably due to their varying growth habits. The trees of A. lebbek were taller with more crown spread, whereas G. optiva had more crown height with lesser crown spread. Stem diameter growth of A. lebbek was highest followed by B. purpurea. Lopping intensity did not affect tree survival and also canopy growth in most years, but collar diameter and diameter at breast height (dbh) decreased significantly at the higher intensity of lopping. Intercropped grass species E. binata caused a greater adverse effect on the tree survival and growth parameters than C. fulvus. The biomass production (leaf and branch wood weight of lopped material) was highest at eight years of age in 1987 but showed no definite trend over the study period. Bauhinia purpurea produced the highest biomass and L. leucocephala the lowest, despite opposite trends in their survival. The performance of A. lebbek was on par with B. purpurea in the later years (1990--1993), but G. optiva was inferior despite its superiority in the early years (1984--1985). The effects of lopping intensity were not appreciable, but intercropping with E. binata decreased leaf and branch wood production significantly compared with C. fulvus throughout. However, the adverse effect was comparatively less pronounced on B. purpurea and A. lebbek than on G. optiva and L. leucocephala. The fibre yield of G. optiva also decreased due to intercropped E. binata compared with C. fulvus. This revised version was published online in June 2006 with corrections to the Cover Date.     

Evaluation of coastal sandy soils and their saline ground waters for afforestation: A case study from India

A study was conducted in coastal Andhra Pradesh to evaluate current practices and suitability of coastal sandy soils for raising tree plantations. Poor soil moisture retention characteristics of these soils make irrigation an essential, but costly silvicultural practice for obtaining desirable establishment and growth. Saline ground water is the only available source of irrigation. Its use seems safe and successful in such soils. The farmers have adopted a specialized manual splashing technique (Doruvu) of seepage pits for irrigating tree plantations in their establishment phase. This technique was observed to be well suited to the sandy soils. Casuarina equisetifolia and Eucalyptus tereticornis were the two most popular species being afforested. E. camaldulensis is also in an introduction phase. Comparison of growth performance and biomass accumulation by C. equisetifolia and E. tereticornis plantations at different growth stages showed the former to be significantly better than the latter. Build up of the soil organic carbon was also found to be markedly more in comparable C. equisetifolia plantations than those of E. tereticornis. Similarly, the blanketing influence of C. equisetifolia plantations was observed to modify soil temperature extremes considerably more than in the case of E. tereticornis. The findings, indicated a vast scope for utilization of coastal sandy soils for afforestation with the successful use of saline ground waters.     

A comparison of cold tolerance and biomass production in Leucaena leucocephala L. pulverulenta and L. retusa

Expansion of leucaena into temperate regions of the world will require additional tolerance to freezing stress. Leucaena leucocephala may be the most widely tested woody legume in the last 15 years. Leucaena pulverulenta and Leucaena retusa are two species with more cold tolerance than L. leucocephala. A screening trial with 38 accessions of L. pulverulenta, 26 of L. leucocephala, and 1 of L. retusa was begun in 1983 at Kingsville, Texas where freezing temperatures may occur every year. A biomass production trial using L. leucocephala (K-8) was planted in 1982 to evaluate the biomass production of this widely used strain under semi-arid conditions with minimum fertilizer inputs and no irrigation. Biomass estimations were made with regression equations that predicted weight from log₁₀ basal stem diameter measurements. The trees in both trials were harvested after a − 12°C freeze in December 1983 killed all trees except L. retusa to ground level. Less severe freezes in 1984–1985 of − 7°C and − 9°C killed all L. leucocephala to ground level whereas less than 2% of the L. pulverulenta were killed to ground level, thus demonstrating the difference in cold tolerance between these two species. Dry wood production in the screening trial of the L. leucocephala after 10 months' coppice growth ranged from 654 to 3602 g dry biomass per tree and that of the L. pulverulenta during the same period ranged from 437 to 2505 g dry biomass per tree. Dry wood productivity of L. leucocephala (K-8) 1-year coppice growth ranged from 1.4 t ha⁻¹ on a poorly drained soil to 10 t ha⁻¹ on a moderately drained soil.     

Root biomass and distribution of five agroforestry tree species

Knowledge of the quantitative assessment and structural development of root systems is essential to improve and optimize productivity of agroforestry systems. Studies on root biomass recovery by sieves of different mesh sizes (2.0, 1.0, 0.5 and 0.25 mm) and root distribution for four-year-old individuals of five agroforestry tree species viz.; Acacia auriculiformis A. Cunn. ex Benth, Azadirachta indica A. Juss, Bauhinia variegata L., Bombax ceiba L. and Wendlandia exserta Roxb. were conducted at the research farm of Rajendra Agricultural University, Pusa, Bihar, India. The results indicated that the 0.5 mm sieve was adequate for recovery of the majority of roots. All the tree species exhibited a large variation in root depth and horizontal root spread four years after planting. The maximum root depth was recorded in W. exserta (2.10 m) and minimum in B. variegata (1.00 m). Horizontal root spread was 2.05 m in B. ceiba and 8.05 m in A. auriculiformis. Root spread exceeded crown cover for all species. The primary roots were more horizontal than the secondary roots. The length and diameter of the main root were highest in A. indica (108.3 cm) and B. ceiba (23.2 cm), respectively. Highest length and diameter of lateral roots were recorded in B. variegata (201.6 cm) and A. indica (1.8 cm), respectively. Total root biomass among different species accounted for 18.2–37.9% of the total tree biomass. Results of this study infer that although all the species have potential to conserve moisture and improve fertility status of the soil, A. auriculiformis is the most effective for promoting soil fertility. The deep rooted W. exserta and A. auriculiformis will be preferred for cultivation under agroforestry systems and could reduce competition for nutrients and moisture with crops by pumping from deeper layers of soil.     

Establishment and growth potential of fuelwood species in northeastern Mexico

Fifteen multipurpose tree species native and introduced to the matorral (low dry shrubland and forest) of the Gulf Coastal Plain in north-eastern Mexico were planted in monoculture in four randomized blocks. Measurements of various growth parameters and volume of trees over the first five years were evaluated. Eucalyptus camaldulensis, E. microtheca, Leucaena leucocephala (exotic species) and Parkinsonia aculeata (native species) performed best while Prosopis glandulosa, Helietta parvifolia and Acacia rigidula (native species) did not establish well due to biotic problems which arose under plantation conditions. Acacia farnesiana, A. wrightii and two Pithecellobium spp. (native species) showed high yields which is of great interest since their multipurpose potential is the best of all fifteen species. These species could be of great importance in rehabilitating badly degraded parts of the matorral before such areas deteriorate to an irreversible desertified condition.

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Resumen Para cuantificar el crecimiento y la productividad de las plantas leñosas, quince especies nativas e introducidas con caracteristicas multipropósito fueron plantadas en monocultivo en una zona de matorral de la Planicie Costera del Golfo Norte de México a partir de 1984. Durante cinco años de medición continua sobre diferentes parámetros de crecimiento, fueron evaluados los resultados sobre el comportamiento y productividad (volumen de la madera) de las especies. Eucalyptus camaldulensis, E. microtheca, Leucaena leucocephala (especies exóticas) y Parkinsonia aculeata (especie nativa) presentaron los mejores crecimiento en altura y en volumen en comparación con Prosopis glandulosa, Helietta parvifolia y Acacia rigidula (especies nativas). El desarrollo de Acacia farnesiana, A. wrightii y ambos Pithecellobium spp. (especies nativas) se puede considerar bastante bueno. Estas últimas especies son de gran interés como potencial forestal y forrajero. Todas las especies ensayadas son de suma importancia en la rehabilitación de matorral degradado y en el manejo silvicola de estos vastos recursos.

     

Influence of fly ash mixtures on early tree growth and physicochemical properties of soil in semi-arid tropical Alfisols

The beneficial effects of planting mixtures comprising of fly ash at the rate of 66%, 33% and 17% by soil volume were assessed for the early growth of two economic tree species, teak (Tectona grandis) and leucaena (Leucaena leucocephala) in rainfed Semi-Arid Tropical (SAT) Alfisols in India. Measurements of tree growth were made at six monthly intervals over a 3-year period. Fly ash at 66% by soil volume of the planting pit significantly increased the diameter of teak at breast height (dbh). In leucaena, application of fly ash at 17% by soil volume was found to be effective in increasing dbh during most of the study period. Changes in soil physicochemical properties (bulk density, water retention at 0.033 and 1.5 MPa, profile moisture content, pH, soluble salt content and organic carbon) were simultaneously studied. Among these soil properties, plant available water (PAW) and organic carbon (OC) contents explained variations in diameter at breast height (dbh) in teak during most of the sampling period, while none of the soil properties significantly explained dbh in leucaena. The highest dose of fly ash applied (66% by volume) decreased bulk density (BD) and increased PAW in teak and to an extent in leucaena during the first 2 years of tree growth. Higher profile moisture content was noted in pits treated with 66% fly ash during the dry period of November (after the end of seasonal rainfall which occurred from June to October) during the first and second year after application. An erratum to this article can be found at     

银合欢生态适应性研究

１９９０年引入１１个种源的银合欢，在南亚热带云南景东育苗栽培试验，结果表明银合欢有较强的生态适应性，其中勐腊、元谋、元法种源生长较好，东扑硅Ｋ×１、Ｋ×３Ｃ显示了抗逆性强的杂交优势，为我国热带、南亚热带地区推广栽培银合欢提供了科学依据。     

Effect of soil fertility on early growth of Leucaena and Gliricidia in alley farms

Young leucaena (Leucaena leucocephala (Lam) de Wit) and gliricidia (Gliricidia sepium (Jacq) Steud) alley-cropped with food crops on farmers' fields in southwestern Nigeria showed marked variability in growth and foliage coloration. A field study was undertaken to determine whether variability in soil fertility was responsible for the differential growth of the two tree species. Plant height of leucaena and gliricidia at 6 and 9 months after planting (MAP) were significantly correlated with soil organic C and total N. Dry matter yield at 12 MAP was also significantly correlated with plant height, soil organic C and total N for leucaena but not for gliricidia. Soil and plant N concentrations were significantly higher in farms with normal than those with chlorotic plants.     

Predicting field performance of five irrigated tree species using seedling quality assessment in Burkina Faso,West Africa

Five exotic tree species (Acacia angustissima (Mil.) Kuntze, A. mangium Wild, Gliricidia sepium (Jacq.) Alp., Leucaena hybrid (L×L), and Leucaena leucocephala (Lam.) de Wit) were investigated to determine whether parameters of nursery seedling stock quality could be used to predict their field performance in a plantation irrigated with treated waste-water to produce fodder and wood. Plants were grown in the nursery in two contrasting rooting substrates (ordinary nursery soil and sand), predicted to have different effects on resource allocation. Three categories of morphological indicators were measured, i.e., plant dimensions (height, diameter, root length), plant weights (shoot, root and whole plant weights) and indices (sturdiness quotient ‘SQ’, shoot:root dry weight ratio ‘SRR’ and Dickson’s quality index ‘DQI’). In the nursery, all species performed better in the ordinary nursery soil for all growth parameters except root length. Thus ordinary nursery substrate appeared superior to sand in terms of plant quality. However, a follow up at plantation phase revealed that only some morphological attributes or ratios were suitable to predict field performance for the five tested species in irrigated plantation. In addition, the effect of the substrate observed at the nursery stage had disappeared 12 months after out planting due to the availability of water and nutrients provided by the treated waste water used for the irrigation. The results showed that root collar diameter and DQI appeared to be the most appropriate indicators to predict the outplanting performance of the five tested species in a short-rotation irrigated plantation in semi-arid Burkina Faso. The former measure is simpler and non-destructive.