An evaluation of the use of short-term weight changes in grazing sheep for estimating herbage intake

A technique for estimating herbage intake by weighing sheep before and after grazing, with an allowance for insensible weight loss (IWL) was evaluated.
Live weights were estimated by weighing each animal 200 times using an electronic balance and a microcomputer. This enabled each sheep to be weighed to ± 10 g and overcame the problem of fluctuation in weight due to movement. Mean rates of IWL in housed sheep were 2·3, 1·4, and 0·8 g min⁻¹ for walking, standing eating or idling and when lying ruminating respectively. Intake rate (IR) estimated by weighing gave a small but significant bias of −0·8 g min⁻¹ when compared with measurements of IR made using housed sheep. Estimates of IWL and IR were made with ewes grazing continuously stocked ryegrass swards maintained at surface heights of 3,6,9 and 12 cm. IWL was higher outdoors than indoors and tended to be higher on the longer swards. IR estimated by weighing gave similar results to those obtained using the chromic oxide technique. At a sward surface height of 3 cm, IR and total intake were lower than for taller swards and total time spent grazing decreased with increasing sward surface height.     

Voluntary intake and in vivo digestibility by sheep of five lilies of Cenchrus ciliaris selected on the basis of preference rating

A study is reported on the voluntary intake and digestibility by sheep of five lines of Cenchrus ciliaris when grown as swards and cut at six different stages of growth. Previous studies with C. ciliaris grown in small rows had shown large differences between these lines in grazing preference by cattle. The ranking order for preference wasline31>10 = 23>30 = 36.
When offered singly to sheep in pens the five lines could be divided into three groups according to their voluntary intakes. Lines 10 and 30 had the highest mean daily intakes of 56·1 and 57·6 g (kg W)-^0·75 compared with lines 23 and 36 with low mean intakes of 48·5 and 48·6 g (kg W)^-0·75. Line 31 was intermediate with a mean intake of 543 g (kg W)^-0·.
Mean dry matter digestibility of the five lines ranged from 0·577 to 0·597%. Lines 10 and 30 had the lowest digestibility. The difference in intake between the five lines was positively correlated with their proportion of leaf ( r = 0·97). Low-intake lines (23 and 36) also had the highest proportion of seed head.
It was concluded that selection of C. ciliaris on the basis of preference by cattle in a cafeteria study provided no useful guide to voluntary intake when the different lines were grown, cut, dried, chopped and fed separately to sheep.     

The long-term effects of a range of pasture treatments applied to three semi-natural hill grassland communities.

Five pasture treatments were applied to three semi-natural hill grassland communities. The pasture treatments were: (1) controlled grazing, (2) controlled grazing + lime, (3) controlled grazing + lime + phosphate, (4) controlled grazing + lime + phosphate + oversown white clover and (5) controlled grazing + lime + phosphate + oversown white clover + oversown perennial ryegrass. The communities were dominated by Agrostis/Festuca (site 1), Molinia (site 2) and Nardus (site 3). The Nardus at site 3 was substantially reduced by herbicide before treatments were applied. All treatments were grazed simultaneously by mature wether sheep on three occasions each year to a residual herbage mass of 560 kg DM ha^?1. There were two grazing periods, each of 4 weeks duration, beginning in mid-May and mid-July respectively and a further grazing period of 3 weeks duration beginning in mid-October. The number of sheep grazing days and liveweight gain of sheep were recorded during each grazing period for 13 years at each site. Organic matter intake (OMI) and digestibility (OMD) of ingested herbage were measured in years 2, 5 and 13. Mean daily OMI per head ranged from 1290 g for treatment 1 to over 1400 g for treatments 4 and 5. OMI values were higher for the Agrostis/Festuca site (mean OMI 1450 g) than for the Molinia and Nardus sites (1310 g and 1370 g respectively) largely owing to differences in the values for treatments 1, 2 and 3. OMI values for each treatment decreased by around 300 g between May and October. Mean OMD values from treatments 1, 2 and 3 were higher for the Agrostis/Festuca site (0·66) than for the Molinia and Nardus sites (0·63 and 0·64 respectively). The OMD values for treatments 1, 2 and 3 at the Molinia and Nardus sites declined by between 0·05 and 0·02 between years 2 and 13 and also showed the greatest decrease between May and October (0·08). OMD values for treatments 4 and 5 (0·67 and 0·69 respectively) were higher than for treatments 1, 2 and 3 at all sites and levels were maintained over 13 years. Individual liveweight gains of sheep increased from treatment 1 (?18 g d^?1) to treatment 5 (82 g d^?1), but they decreased over time on all treatments. The Agrostis/Festuca site gave 17% more annual sheep grazing days than the Molinia site and 33% more than the Nardus site. The mean number of annual sheep grazing days for each treatment ranged from 2250 for treatment 1 to 3640 for treatment 5. Annual sheep grazing days increased over 13 years by between 35 and 45% for treatment 1 and 55 and 70% for treatment 5. The results are described in relation to the changes in pasture composition and herbage accumulation.     

Comparative studies of the influence of sward conditions on the ingestive behaviour of cows and sheep

Separate groups of non-lactating cows and wether sheep grazed at similar herbage allowances for two successive 5-d periods on swards that had previously been grazed frequently or infrequently with the intention of creating differences in canopy structure. Measurements were made of sward structure and composition, ingestive behaviour and diet composition. The preliminary treatments had little effect upon either sward conditions or animal behaviour.
Herbage mass was reduced from 4020 kg dry matter (DM) ha⁻¹ to 3290 kg DM ha⁻¹ on average over a 5-d grazing period as a consequence of the relatively low grazing pressure imposed. This resulted in a mean decline in intake per bite of 28%, and the changes for cattle and sheep did not differ significantly. However, there was a marked difference in the other behavioural responses of the two species; in the sheep biting rate fell and grazing time increased with declining herbage mass, particularly in the sward previously grazed infrequently, whereas the changes in the cattle were small.
Differences in the botanical composition of the herbage eaten by cattle and sheep were minor, but there was a small but consistent advantage to the sheep in the digestibility of the herbage eaten.
In the first of the two periods the variation in surface height after grazing was substantially greater for sheep-grazed than for cattle-grazed swards, indicating more patchy grazing by the sheep.
Estimates of daily herbage organic matter (OM) intake calculated from ingestive behaviour variables were high (means 38 and 32 g (kg LW) ⁻¹ for cattle and sheep respectively) and usually declined substantially over a grazing period.     

Bitterness and Toxicity in Wild Yam (<Emphasis Type="Italic">Dioscorea</Emphasis> spp.) Tubers of Nepal

Wild yams make a significant contribution to diets of tribal people in Nepal. However, these wild tubers are unpalatable, taste bitter, produce inflammation and show occasional toxicity. Four wild yam species, which are eaten after primary treatment by Nepali aborigines, were analyzed for bitter and toxic principles. Bitter components were identified as furanoid norditerpenes (diosbulbins A and B). Diosbulbins A and B were found in the range of 0.023–0.046 and 0.151–0.442 g kg⁻¹, respectively. Results demonstrated that diosbulbin B, with an average value of 0.314 g kg⁻¹, was the principal bitter compound as compared to diosbulbin A (0.037 g kg⁻¹). The toxic alkaloid, dioscorine and histamine (an allergen) were not detected in these tubers, whereas cyanogens (as HCN equivalent) content were found ranging from 3.2 to 6.0 ppm. Our results revealed that Nepali wild yam tubers are not toxic varieties, as they do not contain either toxic dioscorine or histamine and cyanogens contents were satisfactorily below the safety limits. The inflammation and occasional toxicity observed could possibly be due to the presence of high level of oxalate in these tubers. Domestic cooking methods were found to be very efficient in removing bitterness, thus making the bitter yams palatable.     

THE EFFECT OF STOCKING DENSITY ON THE PRODUCTIVITY OF YEARLING FEMALE SHEEP

The effect of stocking density on the productivity of yearling female sheep was studied over the 1960 grazing season. The cumulative effect on pasture productivity of previous differences in grazing pressure was eliminated by re-randomization of the pasture after each grazing cycle. Stocking densities from 12 to 30 sheep/acre/unit of time were imposed from March to September, followed by 7 months on a common treatment to measure subsequent effects on animal fertility. Correlations were found between stocking density and (a) liveweight gains, (b) loss of weight in fasting, (c) quality and quantity of herbage residues at the end of the last grazing cycle. Subsequent animal fertility was not measurably affected.     

Utilization of silage nitrogen in sheep and cows: amino acid composition of duodenal digesta and rumen microbes

Two experiments are reported. In the first experiment the amino acid compositions of rumen bacteria and protozoa isolated from sheep given a diet of grass silage were compared with those taken from sheep given a diet of hay and barley; four sheep were assigned to each diet. There were no significant differences (P<0·05) between diets in the amino acid compositions of the microbial fractions and bacterial contents of α-ɛ-diaminopimelic acid were also similar for both diets.
In the second experiment the digestion of a grass silage and barley diet (65:35; 135 g CP (kg DM)^-1) was studied using four non-lactating Ayrshire cows fitted with cannulae in the rumen and in the proximal duodenum. The mean N intake was 154 g d^-1 and the corresponding duodenal flow was 126 ± 10 g d^-1 indicating that 0·21±0·07 of the N intake was absorbed between the mouth and the duodenum. The mean rumen NH₃-N concentration was high, 297 g litre^-1, and the mean rate of microbial protein synthesis in the rumen was low, 17 g microbial N (kg OM apparently digested in the rumen)^-1. There was a low content of bacterial N in the non-ammonia N at the duodenum (mean proportion 0·52±005) and low concentrations of methionine and lysine in the duodenal digesta protein. The results are discussed in relation to previously published data on the digestion of silage diets in sheep and to the utilization of silage diets for milk production in the cow.
It is concluded that with silage diets the supply of methionine and lysine to the duodenum is likely to be low because of the low rates of microbial protein synthesis in the rumen and that in cows those two amino acids may be limiting for milk production.     

A PORTABLE-CORRAL TECHNIQUE FOR MEASURING THE EFFECT OF GRAZING INTENSITY ON YIELD, QUALITY AND INTAKE OF HERBAGE

A grazing experiment using four stocking rates of sheep, equivalent to 34, 45, 57 and 68 sheep/ha on an assumed 200-day grazing season, was conducted using a portable grazing corral technique. Plots were subjected to fotir grazing periods between mid May and early September. The effect of stocking rate on herbage yield and quality and the influence of these factors and feed intake on liveweights of the sheep were recorded. The mean daily herbage organic matter available over the trial was 9.6, 5.8, 3.3 and 23% of the total liveweight of the sheep at each of the four stocking rates. The highest grazing pressure led to the production of high-quality herbage, but also led to reduced productivity, low feed intake and liveweight losses. The most lenient grazing pressure failed to provide adequate herbage utilization. Even at the most intensive stocking pressure, only 66% of the herbage available to ground level was utilized in grazings after July. Intake results suggested that sheep of 45 kg liveweight required 1000–1200 g digestible organic matter per day to maintain body weight. Despite the higher in vitro digestibility of herbage on offer at the higher stocking rates, intake was limited through lack of herbage; a high degree of correlation existed between herbage availability and herbage intake over the four stocking rates and at all grazings. It is concluded that the portable corral technique is well suited for grazing studies and the assessment of sward response to varying stocking rates and is particularly useful where facilities for more extensive studies are limited.     

Effect of different grazing management systems on the herbage mass and pasture height of a Nardus stricta grassland in western Scotland, United Kingdom

The aims of this study were to examine the effect of three grazing treatments (year‐round stocking rates of 0·8 ewes ha^?1, 0·5 ewes ha^?1 and 0·5 ewes ha^?1 plus grazing cattle in summer), imposed for 4 years, on the herbage mass and surface height of a Nardus stricta‐dominated grassland in western Scotland and to obtain estimates of annual productivity of this grassland. Nardus stricta‐dominated grassland comprised proportionately 0·20 of the grazing area. Stocking rate of sheep had no significant effect on the herbage mass of the grassland in the first 2 years of the experiment, although mean summer pasture heights were significantly higher under the lower stocking rate of sheep. The pasture on the treatment with cattle grazing in summer had a significantly lower herbage mass and lower surface height than the two sheep‐only grazing treatments. Year‐to‐year variation in the herbage mass and surface height of herbage in summer was greater than the effect of treatments. Despite changes in surface height, the structural diversity of the grasslands was not increased by the treatments. The annual production of vascular plant material ranged from 417 g DM m^?2 in 1994 to 628 g DM m^?2 in 1996.     

Selection by sheep and goats for perennial ryegrass and white clover offered over a range of sward height contrasts

The selection by sheep (six Coopworth ewe hoggets, 44·3 ± 4·6 kg live weight) and goats (six Saanen/Anglo‐Nubian yearling males, 38·1 ± 3·8 kg live weight) for perennial ryegrass (Lolium perenne) and white clover (Trifolium repens) and for sward height was measured in two experiments involving paired turves. Pairs of turves with herbage of differing height and of either the same or different plant species were offered. One sward (fixed height species, FHS) was always offered at 130 mm and the other (variable height species, VHS) at 130, 90 or 50 mm. Turves (450 mm × 220 mm) were cut to a soil depth of 100–150 mm from areas of perennial ryegrass and white clover regrown to the desired height after previously being cut to 30 mm. Each turf in a pair was weighed (±1 g) before and after grazing by penned animals maintained on a barley‐based pelleted diet. The number of prehending bites taken from each turf was recorded over a grazing period (128 ± 12 s). Bite mass, bite rate and intake rate were calculated. As the sward height of the VHS turf declined, an increasing proportion of the diet was selected from the 130 mm turf. When averaged over all height contrasts, both animal species selected a higher proportion (0·776 ± 0·026) of their diet from 130‐mm white clover than from 130‐mm perennial ryegrass (0·591 ± 0·018) turves. On average, goats selected a higher proportion (0·721 ± 0·022) of their dry‐matter (DM) intake from the 130‐mm turf than sheep (0·646 ± 0·019), but the effect was not consistent. In contrasts with perennial ryegrass as the VHS (and both perennial ryegrass and white clover as FHS), the proportion of the diet selected from the 130‐mm turf was very similar for both animal species. However, with white clover as the VHS (and both perennial ryegrass and white clover as FHS), goats selected a higher proportion of their intake from the 130‐mm turf to the extent that in the 130‐mm perennial ryegrass/50‐mm white clover contrast sheep showed as strong selection for 50‐mm white clover as goats did for 130‐mm perennial ryegrass. This lesser selection of goats for white clover as its height in a sward declines is likely to contribute to the higher white clover content observed in swards grazed by goats. Bite mass was greater on white clover (246 ± 5 mg DM bite^–1) than on perennial ryegrass (173 ± 5 mg DM bite^–1) and was greater for goats (255 ± 6 mg DM bite^–1) than for sheep (195 ± 5 mg DM bite^–1). Bite rate was greater on perennial ryegrass (45·9 ± 1·0 bites min^–1) than on white clover (39·9 ± 1·0 bites min^–1) and was greater for sheep (45·5 ± 1·1 bites min^–1) than for goats (42·5 ± 1·1 bites min^–1). Apparent intake rate by both sheep and goats was lower (mean, 5·0 ± 0·29 g DM min^–1) on 130 mm perennial ryegrass/white clover than on 130 mm perennial ryegrass/perennial ryegrass (7·0 ± 0·27 g DM min^–1), but was higher (9·62 ± 0·29 g DM min^–1) on 130‐mm white clover/perennial ryegrass than on 130‐mm white clover/white clover (8·2 ± 0·29 g DM min^–1) combinations.     

THE EFFECT OF INTRAVENOUS INFUSIONS OF METHIONINE ON THE VOLUNTARY INTAKE OF GRASS SILAGE BY SHEEP

The effect of intravenous supplements of L-methionine on the voluntary intake of a ryegrass silage, prepared using formic acid as an additive, was determined in a 4×4 Latin Square experiment with 4 sheep. Animals receiving no methionine consumed 1061 g DM/day; those receiving 0.8, 1.2 and 2.0 g methionine/day consumed 1015, 1012 and 1078 g DM/day. The concentration of methionine in blood plasma (Y; (_imoles/I) increased with the dose rate of methionine infused (X; g/day) such that Y=36.7 + 38.3X (r=0.80; P<0.001). It is concluded that with a diet of this type methionine may be limiting for protein synthesis in the tissues but is not involved in the control of food intake.     

Social network analysis of sheep grazing different plant functional groups

Social Network Analysis (SNA) is used for the first time to investigate the relations between plant functional groups and social structure of grazing sheep. The research was conducted on a grassland in Crete's island Lefka Ori, Greece during 2016. A flock of 20 sheep of Sfakion race was analysed as a network and the sheep as nodes. The focal sampling technique was applied during four experimental periods of four consecutive days × eight hours/day. The plant species selected by sheep during grazing were categorized into four functional groups (grasses, broad‐leaved forbs, shrubs and trees), and the proximity and conflict relations that were developed among the flock members were recorded. The tools of SNA were defined and interpreted in the sheep flock, and the Visone software is used to calculate the network variables of proximity and conflicts relations of nodes. Correlations between the plant functional groups and network variables were examined by Spearman's bivariate correlation test. Sheep collectivity was enhanced while grazing grasses and simultaneously, a tendency for hierarchization within the flock was identified. Similarly, the proximity of sheep increased during forbs’ grazing while their individualism was strengthened. When grazing shrubs, the proximity of sheep was only in part maintained while the grazing of trees seems as the most deconstructive forage option of sheep proximity relations. In general, the grazing of all plant functional groups decreases conflicts among sheep except that of trees, which was insignificant for developing such relations.     

Bovine faeces as a source of micro-organisms for the in vitro digestibility assay of forages

The aim of this study was to investigate whether bovine faeces could replace rumen liquor collected from fistulated sheep in the determination of organic matter digestibility (OMD) of forages in vitro . The first experiment compared two inocula that had been prepared by mixing 83 and 333 g of cattle faeces with 1 litre of artificial saliva according to the first stage of the in vitro technique of Tilley and Terry ( Journal of the British Grassland Society, 18, 104–111, 1963). Inocula made from 333 g of faeces gave higher values of OMD than that prepared from 83 g of faeces and was used in subsequent studies. When a 48-h acid pepsin digestion, the second stage of the Tilley and Terry (1963 ) technique, was included, the OMD values and the ease of filtering of undigested residues were increased. The second experiment compared digestibility determined with inocula produced from faeces with that produced from rumen liquor. The OMD of a ryegrass hay determined with an inoculum prepared from faeces was 0·468 compared with 0·528 when using an inoculum prepared from rumen liquor. The third experiment investigated the accuracy of estimating the OMD of eight contrasting forages determined using rumen liquor from three sheep ( y ) and faeces from two cows ( x ). All regressions were significant ( P  < 0·001) and had residual standard deviations (r.s.d.) of between ±0·019 and ±0·022. The final study compared faeces collected from cattle fed with either hay or concentrates. The OMD of forage was higher from the hay-fed animals, but the r.s.d. values of the regressions were similar. It was concluded that cattle faeces shows potential as an alternative to rumen liquor collected from rumen-fistulated sheep for use in the in vitro digestibility assay of forages.     

Evaluation of propionic acid-treated hay as a feed for growing sheep

Two experiments were carried out to evaluate propionic acid-treated hay as a feed for sheep in which eighteen ewes (experiment 1) and eighteen ewe lambs (experiment 2) were fed ad libitum on hay only. At feeding, the propionic acid-treated hays had higher D-values and water-soluble carbohydrate concentrations, but a lower dry matter concentration, than the untreated hays. The voluntary feed intake and liveweight gain of the sheep fed on two acid-treated hays and on an untreated hay in experiment 1 were similar. In the second experiment an acid-treated moist hay was eaten in greater amounts by the sheep and liveweight gains were greater than on the corresponding untreated hay, but were not significantly different from those of sheep fed on field-cured hay.     

The effects of regrowth and maintenance height on a grass sward with a high density of tillers

A perennial ryegrass sward was grazed by sheep in April 1993 to a target sward surface height (SSH) of 3 cm to create a high density of grass tillers. From 3 May, the sheep were removed and small plots were established on the sward, when the average tiller density (± s.e.) was 35 900 ± 420 live tillers m⁻². Different regrowth treatments were then imposed by allowing plots to regrow to target SSHs of 6 cm (18 g dry matter (DM) M⁻²), 9 cm (78 g DM m⁻²), 12 cm (132 g DM m⁻²) or 15 cm (197 g DM m⁻²). The plots were then maintained by cutting at either 6 or 9 cm SSH until the end of the experiment on 30 September. Live tiller density was reduced by regrowth beyond 8.9 cm (78 g DM m⁻², P <0.001) and leaf-stem ratio and in vitro organic matter digestibility were reduced by regrowth beyond 6.1 cm (18 g DM m⁻², P < 0.05). The effect on live tiller density was sustained through the remainder of the season. From the beginning of June to the end of September, maintenance of SSH at an average of 9.1 cm compared with 6.4 cm also resulted in lower live tiller density, live-dead tiller ratio and leaf-stem ratio and higher herbage mass (at least P < 0. 05). There were significant interactions between regrowth SSH and maintenance SSH, so that leaf-stem ratio, live-dead tiller ratio and live tiller density were reduced by regrowth to a SSH of 16–4 cm followed by maintenance at 9.1 cm, compared with regrowth to 6.1 cm and maintenance at 6.4 cm.     

THE INFLUENCE OF SEVERAL FACTORS IN DETERMINING THE GRAZING BEHAVIOUR OF BORDER LEICESTER × MERINO SHEEP

Over a period of 20 months twenty continuous, 24-hour, observations were made on the grazing behaviour of Border Leicester × Merino sheep. Distinct seasonal changes were found in the pattern of grazing. Grazing usually began during the dawn period and decreased at dusk. In spring and summer there were two peaks of grazing, but there was only one peak in autumn and winter. The percentage of grazing taking place between 6 a.m. and 6 p.m. varied with season, and was highest in spring and early summer. These seasonal changes in the pattern of grazing appeared to be independent of grazing pressure, or of the nutritional requirements of the sheep.
The mean grazing time, for all observations, was 8·2 hours with a standard deviation of 0·97. Decreasing pasture availability increased grazing time, as shown by an increased intensity of grazing.
Ruminating times were found to be considerably lower than have been reported for cattle. There was a significant negative correlation between ruminating time and the nitrogen content of available pasture.
There was no clear evidence that the number of lambs carried before parturition, or suckled after parturition, had any effect on grazing time.
There was no evidence that the behaviour of one sheep influenced that of another, even of a twin, grazing nearby.
Consistent differences between sheep were found for ruminating time. The variation in grazing time between sheep, at any one time, was considerably greater than previously reported and differences between sheep were not consistent.
Variation in grazing time appeared to be less in flocks of less than 6 sheep than in flocks of 6 to 12 sheep.     

Evaluation of white clover varieties under grazing and their role in farm systems

Four experiments were established in 1981–84 to investigate the effect of defoliation treatments on white clover varieties when grown with S23 perennial ryegrass. Treatments included a cutting only regime, as used in National List trials, and grazing systems simulating as near as possible those used on farms.
Differential effects of cutting and continuous sheep stocking on white clover varieties, together with significant variety × defoliation interactions, illustrated the importance of the grazing animal in the evaluation of white clover. Differential effects of cattle and sheep grazing were also evident. Cattle grazing was less detrimental to white clover than was sheep grazing, i.e. cattle were not selective, and less stolon was removed. In general, with both cattle and sheep grazing the larger the clover leaf size the greater the loss in stolons, which in turn decreased persistency. The results illustrate how alternating cutting, sheep grazing and cattle grazing managements can be used to maintain optimum clover/grass balance.
Successful clover/grass swards depend on the retention of clover, yet avoiding clover dominance. The difference in N transfer between clover varieties, especially those within the same leaf category, and the extra grass produced without fertilizer N, emphasized the importance of varietal choice. Breeding programmes have been concerned with the selection of larger-leaved, long-petioled varieties for growing in competition with grass in the presence of fertilizer N. However, the present results showed that, under continuous sheep stocking, increase in leaf size does not increase clover yield or persistency.
The results presented emphasize the importance of the grazing animal in the evaluation of white clover varieties and indicate that yield of clover dry matter should not be the major criterion for selection of varieties for farm systems.     

FURTHER STUDIES ON SHEEP GRAZING IN THE NORTHERN PENNINES

The grazing of 3 relatively productive grasslands ( Festucaf Agrostis ) by hill sheep was examined in relation to herbage consumption, grazing pressure, dung deposition and the number of sheep in the area. Herbage intake ranged from 400 Ib/ac/annum (dry weight) on the highest (2450 ft O. D.) and most exposed grassland to 800 Ib/ac/annum on a grassland (1700 ft O. D.) adjacent to extensive heather moorland on blanket bog. The highest pressure (5·5 sheep/ac) was also found on grassland surrounded by blanket bog (1 sheep/14 ac).     

Diet selection by sheep and goats and sward composition changes in a ryegrass/white clover sward previously grazed by cattle, sheep or goats

An experiment was conducted to examine how variation in the composition and structure of mixed grass/white clover swards affected diet selection by sheep and goats. Sward composition in a mixed perennial ryegrass/white clover sward was manipulated by continuous grazing from 28 May to 28 July (pre-experimental phase) with cattle, sheep or goats, and then from 29 July to 2 September (experimental phase) with sheep or goats in a factorial design replicated twice. Sward surface height was maintained at 6 cm by regular adjustment of stocking density. Grazing by different sequences of animal species resulted in significant differences in the proportions of white clover in the sward, and especially in the proportion of clover lamina and petiole. Grazing by goats in the pre-experimental phase led to greater proportions of clover lamina and petiole in the whole sward and the sward surface. The proportion of white clover in the diet selected by sheep in the experimental phase was consistently higher than that in the sward as a whole, but was closely related to that near to the sward surface (approximately the top 2 cm). For goats there was no significant relationship between the proportion of clover in the diet and in the whole sward, and they generally selected a diet with a lower proportion of white clover than was present in approximately the top 2 cm of the sward. It is concluded that on mixed grass/white clover swards goats do not graze as deeply into the sward as sheep and that this results in a lower proportion of white clover in their diet and therefore allows higher proportions of white clover to develop under grazing by goats than by sheep.