Winter cover crop: the effects of grass–clover mixture proportion and biomass management on maize and the apparent residual N in the soil

The advantages and disadvantages of varying mixture proportion of crimson clover (Trifolium incarnatum L.) and Italian ryegrass (Lolium multiflorum Lam.), used as winter cover crops, and cover crop biomass management before maize sowing (Zea mays L.) were studied in a series of field experiments in Eastern Slovenia. Pure stands and mixtures of cover crops on the main plots were split into different cover crop biomass management subplots: whole cover crop biomass ploughed down before maize sowing, aboveground cover crop biomass removed before ploughing and sowing, or aboveground cover crop biomass removed before sowing directly into chemically killed residues.Cover crop and cover crop biomass management affected the N content of the whole aboveground and of grain maize yields, and the differences between actual and critical N concentrations in the whole aboveground maize yield. The whole aboveground and grain maize dry matter yields, and the apparent remaining N in the soil after maize harvesting, showed significant interaction responses to cover crop × management, indicating positive and negative effects. Crimson clover in pure stand provided high, and pure Italian ryegrass provided low maize dry matter yields and N content in the yields in all the observed methods of biomass management. However, within individual management, mixtures containing high proportions of crimson clover sustained maize yields and N contents similar to those produced by pure crimson clover. Considering the expected ecological advantages of the mixtures, the results thereby support their use.     

Cover crops and interrow tillage for weed control in short season maize (Zea mays)

Weed competition can cause substantial maize (Zea mays L.) yield reductions. Interseeding maize with cover crops or a combination of interrow cultivation and interseeded cover crops are possible alternative methods of weed control. This study was conducted to examine the potential of interrow cultivation plus cover crops to reduce weed density in maize without reducing the grain yield. Field experiments were conducted in 1993 and 1994 at two sites in Québec to determine the effects of planting 12 cover crops with maize on weed control. Fall rye (Secale cereal L.), hairy vetch (Vicia villosa Roth), a mixture of red clover (Trifolium pratense L.) and ryegrass (Lolium multiflorum Lam), a mixture of white clover (Trifolium repens L.) and ryegrass, subterranean clover (Trifolium subterraneum L.), yellow sweet clover (Meliotus officinalis Lam), black medic (Medicago lupulina L.), Persian clover (Trifolium resupinatum L.), strawberry clover (Trifolium fragiferum L.), crimson clover (Trifolium incarnatum L.), alfalfa (Medicago sativa L.), and berseem clover (Trifolium alexandrinum L.) were seeded at two planting dates, 10 and 20 days after maize emergence. Interrow cultivation was carried out weekly until forage seeding, with a final cultivation being conducted just prior to cover crop seeding. Cover crop planting date did not affect maize yields or the ability of interrow tillage plus cover crops to suppress the development of weed populations. Maize yield was less affected by the interseeded cover crops under conditions of adequate rainfall. Corn planted in fields heavily infested with weeds resulted in substantial yield reductions even when rainfall was adequate. Except for 1993 at l'Assomption interrow tillage plus cover crop treatments had consistently lower weed biomass when compared to the weedy control. Most of the weed control was due to the interrow cultivation performed prior to seeding of the cover crops. The lowest weed density occurred in the herbicide treated plots. The ability of interrow tillage plus cover crops to suppress the development of weeds was affected by the level of weed infestation, the growing conditions and location. The cover crops provide additional weed control but the interrrow tillage or some herbicide application may still be necessary.     

Nitrogen use efficiency and fertiliser fate in a long-term experiment with winter cover crops

The use of winter cover crops enhances environmental benefits and, if properly managed, may supply economic and agronomic advantages. Nitrogen retained in the cover crop biomass left over the soil reduces soil N availability, which might enhance the N fertiliser use efficiency of the subsequent cash crop and the risk of depressive yield and pre-emptive competition. The main goal of this study was to determine the cover crop effect on crop yield, N use efficiency and fertiliser recovery in a 2-year study included in a long-term (10 years) maize/cover crop production system. Barley (Hordeum vulgare L.) and vetch (Vicia sativa L.), as cover crops, were compared with a fallow treatment during the maize intercropping period. All treatments were cropped following the same procedure, including 130 kg N ha⁻¹ with ¹⁵N fertiliser. The N rate was reduced from the recommended N rate based on previous results, to enhance the cover crop effect. Crop yield and N uptake, soil N mineral and ¹⁵N fertiliser recovered in plants and the soil were determined at different times. The cover crops behaved differently: the barley covered the ground faster, while the vetch attained a larger coverage and N content before being killed. Maize yield and biomass were not affected by the treatments. Maize N uptake was larger after vetch than after barley, while fallow treatment provided intermediate results. This result can be ascribed to N mineralization of vetch residues, which results in an increased N use efficiency of maize. All treatments showed low soil N availability after the maize harvest; however, barley also reduced the N in the upper layers before maize planting, increasing the risk of pre-emptive competition. In addition to the year-long effect of residue decomposition, there was a cumulative effect on the soil’s capacity to supply N after 7 years of cover cropping, larger for the vetch than for the barley.     

Vorfruchtwirkurigen unterschiedlicher Rotationsbrachen auf Winterweizen

Effects of rotational fallows (‘set-aside land’) on subsequent winter wheat. The aim of the present study was to investigate the effects of different fallow treatment on subsequent winter wheat. The field trials included rotational fallows planted with Trifolium repens, Festuca rubra and Lolium perenne sown under winter barley compared to complete fallow and natural fallow without seed application and fallows planted with Trifolium pratense, Festuca rubra and Dactylis glomerata sown under winter wheat. After ploughing up the fallow vegetation, winter wheat was planted for 2 succeeding years at two levels of N-fertilization. Herbicides and fungicides were not applied. The following criteria were investigated: biomass-production, N-uptake, yield, weed infestation, nitrate and water content of the soil. In autumn, after ploughing up the fallow vegetation, the nitrate content of the soil (0—150 cm) increased by up to 210 kg NO₃-N/ha after complete fallow, by up to 60 kg NO₃-N/ha after natural fallow and by up to 75 and 130 kg NO₃-N/ha after fallows cropped with Trifolium repens and Trifolium pratense, respectively. Low nitrate levels of 20—27 kg NO₃-N/ha were observed after fallows planted with grass. N-immobilization caused by ploughing up grass fallows continued until the first harvest of the subsequent winter wheat. In the second year of winter-wheat, no differences of N-mineralization dependent on the previous fallow crop occurred, except in the case complete fallow which showed lower N-mineralization. It can be concluded that fallows cropped with grass lead to a higher nitrogen fertilizer requirement m the succeeding crops. Festuca rubra was able to form dense swards in strong competition with weeds and to decrease the abundance of Alopecurus myosuroides and Apera spíca-venti in subsequent winter wheat, while natural fallow and fallow planted with Trifolium repens and Lolium perenne caused epidemical increases in grass-weed density. Preceding crop effects on grain yield of the winter wheat showed a close relation to N-supply and were compensated by mineral N-fertilization. After natural fallow and fallow covered with Trifolinm repens, yield reductions due to grass-weed competition occurred. Undersown Festuca rubra seems to possess a special suitabihty for cultivation in routional fallows. It establishes itself strongly under different cover crops and is able to form dense swards in strong competition with weeds. Grass-weed density in the succeeding crops will be reduced and nitrate leaching will still be prevented after ploughing up the fallow vegetation. N-fertilization of the subsequent crops must be carried out under considerations of higher N-requirements which is probably not entirely due to a stronger N-immobilization.     

Replacing bare fallow with cover crops in a maize cropping system: Yield,N uptake and fertiliser fate

Cover crops in dry regions have been often limited by low nutrient and water-use efficiency. This study was conducted during 3.5 years to determine the effect of replacing bare fallow by a cover crop on yield, N uptake, and fate of labeled fertiliser in an intensive maize production system. Three treatments were studied: barley (Hordeum vulgare L.), vetch (Vicia villosa L.) and bare fallow during the intercropping period of maize (Zea mays L.). All treatments were irrigated and fertilised following the same procedure, and a microplot in each plot was established with 210 kg N ha^?1 of double labeled ammonium nitrate. Crop yield and N uptake, soil mineral N (N_min), and recovery of ¹⁵N in plant and soil were determined after maize harvest and killing the cover crop. Replacing bare fallow with cover crops did not affect subsequent maize yield but affected N uptake. Vetch increased N supply by legume residues after the second year, and the N content in grain by the third. Nitrogen recover from fertiliser was not affected by treatment and averaged 46%. Barley recovered more ¹⁵N during the autumn–winter period than vetch or fallow. Under representative conditions, average barley N content was 47, vetch 51, and spontaneous vegetation content 0.8 kg N ha^?1. Recovery of ¹⁵N in barley comprised 19% of total N content in aerial biomass, while only 4% in vetch. Vetch enhaced soil ¹⁵N recovery more than other treatments, suggesting its presence in a fairly stable organic fraction unavailable for maize uptake or lost. Replacing bare fallow by a cover crop only reduced fertiliser losses in a year with abundant precipitation. Nevertheless, reduction in soil N_min in vetch and bare fallow treatments was similar, showing that N losses can be reduced in this cropping system, either by replace bare fallow with barley or smaller N fertiliser applicationto maize.     

Legume cover crops as living mulches for winter wheat: Components of biomass and the control of weeds

To gain information about the possible use of legume cover crops as an alternative and sustainable weed-control strategy for winter wheat (Triticum aestivum L.), an experiment was conducted at two sites in the Swiss Midlands in 2001/2002. Under organic farming conditions winter wheat was direct-drilled into living mulches established with four different legume genotypes or into control plots without cover crops. Compared to NAT (control plots without cover crops but with a naturally establishing weed community), white clover (Trifolium repens L.), subclover (Trifolium subterraneum L.), and birdsfoot trefoil (Lotus corniculatus L.) reduced the density of monocotyledonous, dicotyledonous, spring-germinating, and annual weeds by the time of wheat anthesis. Strong-spined medick (Medicago truncatula Gaertner) was less efficient in this regard. While the grain yield was reduced by 60% or more for all legumes when compared to NOWEED (control plots kept weed-free), a significant negative correlation between the dry matter of the cover crop and weeds as well as between the cover crop and the winter wheat was observed by the time of wheat anthesis. The effect of manuring (60 m³ ha⁻¹ liquid farmyard manure) was marginal for weeds and cover crops but the additional nutrients significantly increased total winter wheat dry matter and grain yields. The suppression achieved by some legumes clearly demonstrates their potential for the control of weeds in such cropping systems. However, before living legume cover crops can be considered a viable alternative for integrated weed management under organic farming conditions, management strategies need to be identified which maximise the positive effect in terms of weed control at the same time as they minimise the negative impact on growth and yield of winter wheat.     

Contribution of White Clover Varieties in High-Productivity Systems under Grazing and Cutting

Two medium leaf size varieties of white clover (Trifolium repens L.) were grown in mixture with a range of grass species and varieties in a field experiment conducted over 4 years. The clovers were grown singly or as a blend in swards with a tetraploid perennial ryegrass (Lolium perenne L.), a diploid Italian ryegrass (Lolium multiflorum L.) or tetraploid hybrids between these two species (Lolium x boucheanum Kunth). These grasses were also grown in monoculture. Swards were managed in a regime involving two periods of sheep grazing, two periods of cattle grazing and a silage cut in the same year for 4 years. The objectives of this experiment were 3‐fold: (1) to examine the yield and persistency of white clover in highly productive systems with modern perennial, hybrid and Italian ryegrasses; (2) to compare the performance of white clover varieties grown singly in mixture with grasses with their performance together as a blend, and (3) to examine the persistency of new Italian ryegrass varieties, exemplified by AberComo. The clover content of the sward remained high throughout the experiment, with a mean of 31 % overall. Differences in clover yield between plots were largely attributable to the effects of companion grasses. Total yields were very similar in plots containing different clover varieties and in plots where these varieties had been grown singly or as a blend. Grass yields in mixture were greater than those in monoculture but showed similar relative rankings. The diploid Italian ryegrass variety AberComo showed an unexpectedly high yield in the 4th year of the experiment. The implications of these results for strategies for productive, sustainable agriculture using modern white clover and ryegrass varieties are discussed.     

Establishment, Development and Yield of Forage Legumes and Grasses as Cover Crops in Grain Corn in Eastern Canada

Interseeded cover crops can minimize soil erosion, increase organic matter and nutrient levels and the yield of subsequent crops. However, their performance is very sensitive to local conditions. Field experiments were conducted at two Quebec locations in 1993 and 1994 in order to evaluate the potential use of forage legumes and grasses as interseeds in corn in eastern Canada. Twelve forage species were evaluated. Fall rye ( Secale cereal L.), hairy vetch ( Vicia villosa Roth), a mixture of red clover ( Trifolium pratense L.) and ryegrass ( Lolium multiflorum Lam), a mixture of white clover ( Trifolium repens L.) and ryegrass, subterranean clover ( Trifolium subterraneum L.), yellow sweet clover ( Melilotus officinalis Lam.), black medic ( Medicago lupulina L.), Persian clover ( Trifolium resupinatum L.), strawberry clover ( Trifolium fragiferum L.), crimson clover ( Trifolium incarnatum L.), annual alfalfa ( Medicago saliva L.) and berseem clover ( Trifolium alexandrinum L.) were seeded at two planting dates (10 and 20 days after corn emergence). The control treatments were: hand weeding, chemical weeding and non-weeded. Early seeded forages established better and had higher biomass accumulation than the late seeded ones. In the presence of larger weed populations, the interseeded forages did not develop well due to competition with the weeds. At Macdonald crimson clover provided good soil cover while Persian clover, fall rye and alfalfa provided relatively little cover. Strawberry clover and hairy vetch did not provide early ground cover due to their late development. Forage mixtures of red or white clover and rye grass established well and achieved high populations at the end of the growing season. Fall rye provided good early ground cover but senesced by the middle of the season. The better establishment and early germination of crimson clover caused a 19% reduction in corn grain yield in 1993. In 1994 none of the cover crops caused a reduction in corn yield.     

Intensive cropping of maize in the Southeastern United States

The long growing season of the southeastern Coastal Plains allows planting of a second crop after spring-planted maize (Zea mays L.). Second crops have been shown to reduce erosion and prevent leaching of nutrients and pesticides. Maize grown with a second annual crop might also have a yield advantage over mono-cultured maize. Seven tillage/cropping systems were compared. They included disking for weed control, disking for seedbed preparation, or no disking. Double-cropped treatments included sunflower (Helianthus annuus L.), soybean (Glycine max. L.), a cover crop [crimson clover (Trifolium incarnatum L.)] or no double crop. Double-cropped soybean yields did not respond to irrigation. They averaged 0.63 Mg/ha over 4 years. This is less than half of the local non-double-cropped yields. Sunflower yields averaged 0.89 Mg/ha, also less than non-double-cropped yields (1.0–2.5 Mg/ha). The best continous maize yields (7–8 Mg/ha) were from treatments with disking in some phase of the operation. Treatments with lower maize yields generally had higher plant nutrient contents. Double-cropped maize yields significantly (P<0.10) outyielded mono-cropped maize yields in two of the three years. In 1984, a dry year, the minimum tillage treatment had lower tensiometer readings than the conventionally tilled treatment.     

Effect of biogas digestate,animal manure and mineral fertilizer application on nitrogen flows in biogas feedstock production

The expansion of biogas feedstock cultivation may affect a number of ecosystem processes and ecosystem services, and temporal and spatial dimensions of its environmental impact are subject to a critical debate. However, there are hardly any comprehensive studies available on the impact of biogas feedstock production on the different components of nitrogen (N) balance. The objectives of the current study were (i) to investigate the short-term effects of crop substrate cultivation on the N flows in terms of a N balance and its components (N fertilization, N deposition, N leaching, NH₃ emission, N₂O emission, N recovery in harvested product) for different cropping systems, N fertilizer types and a wide range of N rate, and (ii) to quantify the N footprint of feedstock production in terms of potential N loss per unit of methane produced. In 2007/08 and 2008/09, two field experiments were conducted at two sites in Northern Germany differing in soil quality, where continuous maize (R1), maize–whole crop wheat followed by Italian ryegrass as a double crop (R2), and maize–grain wheat followed by mustard as a catch crop (R3) were grown on Site 1 (sandy loam), and R1 and a perennial ryegrass ley (R4) at Site 2 (sandy soil rich in organic matter). Crops were supplied with varying amounts of N (0–360 kg N ha⁻¹, ryegrass: 0–480 kg N ha⁻¹) supplied as biogas digestate, cattle slurry, pig slurry or calcium-ammonium nitrate (CAN).Mineral-N fertilization of maize-based rotations resulted in negative N balances at N input for maximum yield (Nopt), with R2 having slightly less negative balances than R1 and R3. In contrast, N balances were close to zero for cattle slurry or digestate treatments. Thus, trade-offs between substrate feedstock production and changes of soil organic matter stocks have to be taken into consideration when evaluating biogas production systems. Nitrogen losses were generally dominated by N leaching, whereas for the organically fertilized perennial ryegrass ley the ammonia emission accounted for the largest proportion. Nitrogen balance of the ryegrass ley at Nopt was close to zero (CAN) or highly positive (cattle slurry, digestate). Nitrogen footprint (NFP) was applied as an eco-efficiency measure of N-loss potential (difference of N input and N recovery) related to the unit methane produced. NFP ranged between −11 and +6 kg N per 1000 m³ methane at Nopt for maize-based rotations, without a significant impact of cropping system or N fertilizer type. However, for perennial ryegrass ley, NFP increased up to 65 kg N per 1000 m³. The loose relation between NFP and observed N losses suggests only limited suitability for NFP.     

Residual nitrogen effect of clover-ryegrass swards on yield and N uptake of a subsequent winter wheat crop as studied by use of N methodology and mathematical modelling

The residual effect of 2-year-old swards of clover-ryegrass mixture and ryegrass in monoculture on yield and N uptake in a subsequent winter wheat crop was investigated by use of the ¹⁵N dilution method and by mathematical modelling. The amount of N in the wheat crop, derived from clover-ryegrass residues was 25–43% greater than that derived from residues of ryegrass which had been growing in monoculture. Expressed in absolute values, the N uptake in the subsequent winter wheat crop was 23–28 kg N ha ⁻¹ greater after clover-ryegrass mixture than after ryegrass in monoculture. Up to about 54 kg N ha⁻¹ of the N mineralised from the clover-ryegrass crop was calculated to be leached, whereas only 11 kg N ha⁻¹ was leached following ryegrass in monoculture.     

Yield and quality components of silage maize in killed and live cover crop sods

In sloping areas with high precipitation, planting maize into live winter cover crop sods may help to alleviate the environmental problems associated with clean-tillage production systems of maize. The present study evaluates the performance of silage maize (Zea mays L.) under several cultivation methods: CC (conventional cropping system, i.e., maize was sown into the bare, autumn-ploughed soil); LGS/CK (maize was planted into a living Italian ryegrass (Lolium multiflorum Lam.) sod which was subsequently herbicidally killed); and LGS/MR (similar to LGS/CK, but the ryegrass was mechanically regulated). The research was conducted in the midlands of Switzerland on a fertile sandy loam under humid conditions during three cropping seasons. With 110 kg N ha⁻¹ (fertilizer nitrogen plus mineral nitrogen of the soil at maize planting), the CC system was much more productive than were the LGS/CK and LGS/MR systems in terms of dry matter and nitrogen yields of maize. Increasing the nitrogen supply to 250 kg N ha⁻¹ considerably reduced the yield advantage of CC over the LGS/CK and LGS/MR systems, indicating that nitrogen was the most limiting factor for maize yield in the mulch seeding systems. With 250 kg N ha⁻¹, the LGS/CK and LGS/MR systems produced greater total yields of digestible organic matter (maize plus ryegrass) than did the CC system, whereas the total nitrogen yield was similar for all cropping systems. The whole-shoot concentrations of nitrogen were highest under CC, irrespective of the level of nitrogen supply. With 110 kg N ha⁻¹, concentrations of phosphorus and magnesium were clearly higher for the mulch seeding systems. There were only minor differences among the cropping methods in the concentrations of potassium and calcium in the whole shoot. When 250 kg N ha⁻¹ were applied, there were no significant variations among the cropping systems in the concentrations of minerals. Changes in the botanical composition of the cover crop sod and in the time and method of cover crop control may help to reduce the competition for nitrogen between maize and the living mulch.     

Straw management, crop rotation, and nitrogen source effect on wheat grain yield and nitrogen use efficiency

Nitrogen fertilizer management from different sources and annual crop rotations are important components of wheat (Triticum aestivum L.) production systems, especially where air and soil quality issues have prompted a search for alternatives to wheat straw burning. This study examined the effects of two different wheat straw management options (burning and incorporation by tillage), three crop rotations [wheat-sesbania (Sesbania spp.), wheat–maize (Zea mays L.), and wheat-clean fallow] and three N sources (urea, chicken manure, and urea plus chicken manure) on wheat grain yield and N use efficiency. The experiment was conducted as split–split plot treatment arrangement with three replications for eight wheat cropping seasons in the state of Sonora, Mexico. Results indicated that both wheat grain yield and N use efficiency were higher with burning than incorporation of wheat straw and with fertilization with urea or urea plus chicken manure than chicken manure alone. As shown by the crop rotation-by-straw management interaction, planting sesbania following incorporation of the straw by tillage produced comparable grain yields to straw burning treatment. In contrast, wheat in annual rotation with maize produced the lowest wheat grain yield and N use efficiency irrespective of the wheat straw management and N source applied.     

Nitrogen Uptake by Italian Ryegrass after Destruction of Non-fertilized Set-aside Covers at Different Times in Autumn and Winter

The introduction of the European set-aside into the rotation can modify the nitrogen availability for subsequent crops. With a view to evaluating this effect, three non-fertilized covers were sown in May 1995. These covers were ploughed in at three different times: October 1995, December 1995 and March 1996. The ploughed-in nitrogen quantities did not vary greatly with destruction time. The greatest quantities were incorporated into red clover plots and the smallest quantities into perennial ryegrass plots. The C/N of perennial ryegrass was on average 24 and that of red clover 14.
The soil cover by clover just before ploughing is significantly related to the ploughed-in nitrogen quantities.
An Italian ryegrass was sown as the subsequent crop to the set-aside. Overall, during the whole year 1996 and 1996–1997, there was no difference between the yields for different destruction times except for the first cut of 1996.
The highest production was obtained after red clover and the lowest after perennial ryegrass. The relationship between the nitrogen amount or C/N of the ploughed-in material and the nitrogen export for the annual harvest of Italian ryegrass showed highly significant correlations in 1996 as well as for the 2-year harvest. Two multiple regressions were calculated in order to take into consideration the characteristics of the ploughed-in material of the set-aside.
The measured percentage of soil cover by clover before ploughing and the annual nitrogen export of Italian ryegrass were also significantly related in 1996 and for both years.     

Nitrogen mineralization in sandy loam soils under an intensive double-cropping forage system with dairy-cattle slurry applications

Nitrogen (N) mineralization and soil mineral N contents were measured at 2-week intervals over a 2-year period (June 1994–May 1996) on two different sites in the North West region of Portugal. The experiment was established in fields, which had for many years been under a double-cropping forage system with maize from May to September and a winter crop (mixture of cereals and Italian ryegrass) during the rest of the year. In addition to N fertilizers, dairy-cattle slurry was applied regularly at the sowing of each crop. On this intensive forage system, quantification of N released from slurry, crop residues and soil organic matter becomes important when better N use efficiency and reduced environmental impact from agricultural practices are required. Net N mineralization rates of the 0–10 cm soil layer fluctuated considerably between consecutive incubation periods and ranged from −0.88 to 1.87 mg N kg⁻¹ day⁻¹ with annual average rates of between 0.41 and 0.65 mg N kg⁻¹ day⁻¹. The total N mineralized in the 10 cm depth soil layer reached values between 122 and 224 kg N ha⁻¹ year⁻¹, showing that mineralization was a very important N source for the crops. The amounts of N released during the cold season (November–February) were equivalent to 27–48% of the yearly total. Regression analysis indicated that seasonal variation in N mineralization was only poorly explained by soil moisture and temperature. The changing balance during the year between soil moisture and temperature will contribute to the relatively constant N mineralization rates. Soil mineral N contents during the maize crop were high and exceeded the nutrient requirements for the optimum yield of this crop. Under the climatic conditions of the region and due to the poor development of the winter crop plants at the time, the mineral N left in the soil after the maize crop and released by mineralization during the cold season is particularly vulnerable to nitrate leaching losses.     

The use of hedges to reduce winter wheat losses caused by Italian ryegrass (Lolium multiflorum)

We hypothesized that short duration plant interference aiming to change mainly light intensity and red:far red ratio during crop establishment in Italian ryegrass-infested winter wheat fields, could hinder the development of Italian ryegrass and thus reduce its effect on wheat yield. To test this hypothesis, wheat was planted between previously established live (low red:far red ratio) or dead (high red:far red ratio), barnyardgrass or maize, plant hedges. Experimental results demonstrated that the presence of live or dead plant hedges during the first 30 days of crop establishment and also hedge orientation were important factors regulating weed and crop biomass production and competitive relationships. In barnyardgrass hedge treatments wheat yield was improved up to 67% and Italian ryegrass production was reduced by more than 20%. In plots with maize live hedges oriented N–S the biomass production of wheat was independent of production of Italian ryegrass. Data presented here suggests that there is room for developing weed control technologies on the basis of understanding photosensory processes of weed and crop species.     

Italian ryegrass establishment by self-seeding in integrated crop–livestock systems: Effects of grazing management and crop rotation strategies

We evaluated the re-establishment of an Italian ryegrass pasture by self-seeding on a no-till integrated crop–livestock systems (ICLS) in the southern region of Brazil. This work is part of a long-term experimental protocol initiated in 2003. We tested the effects of various management practices, such as summer crop systems (soybean vs. maize–soybean rotation), stocking methods (continuous vs. rotational) and grazing intensities (low vs. moderate), on Italian ryegrass pasture establishment. In addition, we tested resilience of the system by testing pasture's ability to re-establish following a year without seed head production. The experiment consisted in the rotation, on the same area, of Italian ryegrass pasture grazed by sheep during the winter and up to the end of the grass production cycle, and soybean or soybean–maize grain crops rotation cultivated during the summer. The pasture established itself by self-seeding since 2005. Data were collected in 2011 and 2012 stocking season. The soybean summer crop, continuous stocking and low grazing intensity, all positively affected the production of reproductive tillers in 2011. Grazing intensity in 2011 strongly influenced early vegetative tiller densities (before crop harvest) in 2012. However, none of the grazing intensity or the stocking method treatments affected herbage mass at the end of pasture establishment in 2011 or 2012. On the other hand, the soybean summer crop positively affected pasture establishment, both in term of tiller densities and herbage mass at the end of pasture establishment. The removal of all seed heads in 2011 (preventing seed production) resulted in the total failure of pasture establishment in 2012. Overall, Italian ryegrass establishment by self-seeding relies on the annual replacement of the soil seed bank. This experiment demonstrated that under various stocking methods, moderate grazing intensity and maize or soybean summer crop, Italian ryegrass pasture establishment by self-seeding remains successful even when the stocking periods extended up to the end of the grass production cycle. Self-seeding with moderate grazing intensity ensures successful pasture establishment, reduces labour and costs and allows to increase the stocking period and so animal live weight gain over the grazing season.     

Effects of species diversity on seasonal variation in herbage yield and nutritive value of seven binary grass-legume mixtures and pure grass under cutting

Intensively managed sown temperate grasslands are generally of low species diversity, although swards based on grass-legume mixtures may have superior productivity and herbage quality than grass-only swards. We conducted a cutting experiment over two years to test the effect of species composition and diversity on herbage yield, contents of N, neutral detergent fibre (NDF) and in vitro organic matter digestibility (IVOMD). Perennial ryegrass (PR, Lolium perenne) was sown alone and with each of four forage legumes: red clover (RC, Trifolium pratense), lucerne (LU, Medicago sativa), birdsfoot trefoil (BT, Lotus corniculatus) and white clover (WC, Trifolium repens); WC was also sown with hybrid ryegrass (HR, Lolium × boucheanum), meadow fescue (MF, Festuca pratensis) and timothy (TI, Phleum pratense). Herbage productivity was lowest in pure PR followed by PR/BT, and highest in PR/RC; this mixture had the highest legume proportion, N content and N yield. There was less WC in swards with HR and MF than with PR and TI. These differences were reflected in N contents of herbage of the mixtures. Legumes had higher N and lignin and lower NDF contents and IVOMD than grasses. Among legumes, NDF content was highest and IVOMD lowest in LU, followed by BT and the clovers. The highest N content was in WC. Among grasses, PR and HR had lower NDF contents and a higher IVOMD than MF; the highest N content was in PR. The grass component of mixtures had less effect than the legume component on herbage yield and quality. Results are discussed in terms of their potential to contribute to forage resources in farming practice and enhance resource use efficiency and ecosystem services.     

Temperature Before Cold Acclimation Affects Cold Tolerance and Photoacclimation in Timothy (Phleum pratense L.), Perennial Ryegrass (Lolium perenne L.) and Red Clover (Trifolium pratense L.)

The expected temperature rise in late summer/early autumn can change the conditions for acclimation and affect the winter survival of perennial crops. This study examined the effect of the temperature just before the onset of cold acclimation (pre‐acclimation) on freezing tolerance of timothy (Phleum pratense L.), perennial ryegrass (Lolium perenne L.) and red clover (Trifolium pratense L.) populations (both cultivars and breeding populations) adapted to either northern or southern parts of Norway. Using phytotron experiments, we studied whether increasing pre‐acclimation temperature delays growth cessation, affects photoacclimation and reduces freezing tolerance. Furthermore, we assessed whether these effects were related to the latitudinal adaptation of the plant material. The results showed that a rise in pre‐acclimation temperature decreased both cold acclimation capacity and photoacclimation in these species. This affected the freezing tolerance, which was reduced significantly more in northern‐adapted population of timothy and perennial ryegrass compared with southern‐adapted populations. Red clover was less affected by temperature changes than the grasses.     

Effect of Legume Management on Forage Production and Residual Effects on Upland Rice

An experiment was conducted over the period 1995–96 in a warm-temperate environment in Nepal to investigate the effect of cutting frequency on forage yields of three temperate legume species, grown during the winter season, and the residual treatment effects on a subsequent upland rice crop. The three species, Persian clover ( Trifolium resupinatum ), white clover ( Trifolium repens ) and vetch ( Vicia benghalensis ), proved to be well adapted to the winter growing conditions and produced cumulative forage yields between 6.8 t DM ha⁻¹ (vetch) and 9.2 t DM ha⁻¹ (Persian clover). Vetch grew vigorously throughout the winter months and provided reasonable forage yields between December and February, whereas the clovers provided green fodder up to July. These species may therefore make a substantial contribution to alleviating the severe shortage of quality feed during the winter months. Grain yields of the subsequent rice crop ranged from 3.6 to 7.3 t ha⁻¹. Rice yields were greatly affected by the previous legume species and forage management practices. In general, the removal of legume forage greatly reduced the residual effect, and farmers will have to seek a compromise between maximizing green fodder production and the immediate beneficial residual effect of the legume crop on rice yield.