Weed seed rain interception by grass cover depends on seed traits

Weed management requires a better understanding of the dynamics of the weed seedbank, which is a primary source of weeds in a field. Seeds reaching the ground after seed rain replenish the seedbank and therefore contribute to future weed infestations. Our investigation is based on the hypothesis that a permanent vegetation cover, such as a grassland, can prevent weed seeds from reaching the ground. Therefore, we developed an innovative experimental device to simulate in controlled conditions the seed rain of 12 weed species (Capsella bursa‐pastoris, Conyza canadensis, Myosotis arvensis, Papaver rhoeas, Poa annua, Polygonum aviculare, Ranunculus sp., Rumex obtusifolius, Sonchus asper, Stellaria media, Taraxacum officinale and Veronica persicaria). We quantified the interception of weed seeds by a grass cover. Grass cover height, seed size and seed appendage (e.g. pappus, wing or awn) increased seed interception, in contrast to seed weight and shape index. From these results, we established a linear model to predict weed seed interception by a grass cover as a function of their seed trait values. The relationship between the predicted interception and weed community dynamics observed in grasslands was negative for some species, indicating that other processes may be involved depending on weed species. The weed seed interception model will be incorporated into an existing model of weed population dynamics to simulate the impact of grassland insertion into arable crop rotations.     

Weekly defoliation controls,but does not kill broad-leaved dock (Rumex obtusifolius)

Broad-leaved dock (Rumex obtusifolius L.) is a troublesome weed that predominantly grows in pastures and grassland. We hypothesised that frequent defoliation of Rumex will, over time, result in a reduction in root weight and leaf area, to the point where the impact on grass production is negligible. In order to investigate this hypothesis, we conducted three experiments. The objective of the first experiment was to perform a preliminary test of the hypothesis, using potted plants growing in the controlled conditions of a glasshouse. This experiment showed a rapid decline in leaf growth in plants that were defoliated weekly. The objective of the second experiment was to test the hypothesis in realistic outdoor conditions while still being able to collect detailed plant growth information. This experiment confirmed the findings of the glasshouse experiment and provided evidence that leaf growth ceased as a result of a dwindling supply of carbohydrate reserves in the root. Defoliated plants did not exhibit increased mortality. Finally, the objective of the third experiment was to test the hypothesis in a commercial pasture where normal field operations, specifically grass harvesting (three times) and slurry injection (twice), were performed. The results of this experiment were consistent with the results of the other two experiments. We conclude that weekly defoliation, maintained for three or more months, is an effective method to control (reduce the impact on grass production), but not kill, R. obtusifolius in pasture.     

Weed flora and soil seedbanks in fields dominated by Imperata cylindrica in the moist savannah of West Africa

Imperata cylindrica (L.) Raeuschel is a dominant and infamous grass weed in the savannah of West Africa. Research to reduce the weed to non-damaging levels is a priority activity at many agricultural institutions. The successful development and implementation of long-term I. cylindrica management strategies depend on the ability to predict changes in weed composition after I. cylindrica has been controlled effectively. The weed flora and soil seedbank were assessed from 329 fields dominated by this species in the fringes of the humid forest (HFF), coastal/derived savannah (CDS) and in the southern Guinea savannah (SGS) in 1996 and 1997. The objectives of the study were to correlate species composition of the weed flora with that of the soil seedbank and to determine the effect of management factors and soil properties on the composition of the weed flora. Species richness in the weed flora and in the weed seedbank was higher in the SGS than in the CDS and HFF. Mean weed density per field was generally higher in the HFF (156 ± 25.0 weeds m^–2) than in the CDS (108 ± 8.1 weeds m^–2) and in the SGS (92 ± 6.3 weeds m^–2). Weed composition varied with agroecological zone as well as with management factors and soil properties. Sørenson's index of similarity was low (mean=0.20) in all zones, indicating poor similarity between the weed flora above-ground and the soil seedbank.     

Soil potassium supply and Rumex obtusifolius and Rumex crispus abundance in silage and grazed grassland swards

Three experiments investigating factors influencing the abundance of Rumex spp. (docks) in silage and grazed grassland swards are presented. In Experiment 1, Rumex obtusifolius plants were sown with perennial ryegrass and white clover in pots in March and harvested at either 5- or 10-week intervals between June and October. The 10-week harvest interval increased root dry-matter production of R. obtusifolius compared with the 5-week interval; herbage (above-ground material) production was not significantly affected. In Experiment 2, R. obtusifolius and Rumex crispus population densities in grassland swards were correlated with soil P, K and Mg concentrations, and soil pH. In general, silage swards contained higher population densities than grazed swards. There were significant positive correlations between soil K concentrations and abundance of Rumex spp. in grazed swards and in silage swards. In Experiment 3, R. obtusifolius was sown with perennial ryegrass in pots in March. Treatments consisted of nine rates of K fertilization ranging between the equivalent of 0 and 600 kg K ha⁻¹ year⁻¹. Herbage was harvested at regular intervals (4–6 weeks except during the winter) until May of the following year. In general, perennial ryegrass dry-matter yields were not greatly affected by soil K, whereas limited soil K supply tended to reduce dry-matter production of R. obtusifolius . It is possible that maintenance of moderate soil K concentrations may play a role in limiting abundance of Rumex spp. in grassland.     

Mapping Ridolfia segetum patches in sunflower crop using remote sensing

Ridolfia segetum is a frequent umbelliferous weed in sunflower crops in the Mediterranean basin. Field and remote sensing research was conducted in 2003 and 2004 over two naturally infested fields to determine the potential of multispectral imagery for discrimination and mapping of R. segetum patches in sunflower crops. The efficiency of the four wavebands blue (B), green (G), red (R) and near‐infrared (NIR), selected vegetation indices and the spectral angle mapper (SAM) classification method were studied using aerial photographs taken in the late vegetative (mid‐May), flowering (mid‐June) and senescence (mid‐July) crop growth stages. Discrimination efficiency of R. segetum patches in sunflower crops is consistently affected by their phenological stages, in this order: flowering > senescence > vegetative. In both fields, R. segetum patches were efficiently discriminated in mid‐June, corresponding to the flowering phase, by using the waveband G, the ratio R/B or SAM with overall accuracies ranging from 85% to 98%. The application of the median‐filtering algorithm to any of the classified images improved the accuracy. Our results suggest that mapping R. segetum weed patches in sunflower to implement site‐specific weed management techniques is feasible with aerial photography when images are taken from 8 to 10 weeks before harvesting.     

Weed populations in winter wheat as affected by crop sequence, intensity of tillage and time of herbicide application in a cool and humid climate

Summary There is a lack of information on the combined effects of preceding crop, reduced tillage (especially no-tillage) and the time of herbicide application on the development of weed populations and the efficiency of weed control in winter wheat in humid temperate climates. An experiment was conducted with a crop rotation (winter wheat – oilseed rape – winter wheat – maize) on a sandy loam and a loamy silt soil in the Swiss midlands to investigate the impact of different preceding crops and pre- and post-emergence control of weeds in conventional tillage (CT; mouldboard plough), minimum tillage (MT; chisel plough) and no-tillage (NT; no soil disturbance systems). When winter wheat was grown after maize and winter wheat was grown after oilseed rape, the ranking order of weed density in treatments without herbicide application was NT < MT < CT and CT < MT < NT respectively. Analysis of variance and canonical discriminant analysis showed that Epilobium spp., Sonchus arvensis , Myosotis arvensis and volunteer crops were more abundant in NT than in MT and CT. The efficiency of post-emergence weed control was generally better than that of pre-emergence weed control, regardless of tillage intensity.     

Response of winter wheat (Triticum aestivum L.) and weeds to tank mixtures of 2,4-D plus MCPA with clodinafop propargyl

Field and greenhouse experiments were conducted in 2004 and 2005 to study weed control and the response of winter wheat to tank mixtures of 2,4-D plus MCPA with clodinafop propargyl. The field experiments were conducted at Yazd and Oroumieh, Iran, with factorial combinations of 2,4-D plus MCPA at 0, 975, and 1300 g ai ha⁻¹ and with clodinafop propargyl at 0, 64, 80, 96, and 112 g ai ha⁻¹ in four replications. The greenhouse experiments further evaluated the effect of these tank mixtures on weed control, where each herbicide mixture was considered as one treatment and the experiment was established in a randomized complete block design with four replications. In the field experiments, the herbicides were applied at wheat tillering, while in the greenhouse experiments they were applied at the beginning of the tillering stage and at the four-leaf stage of the grass and broadleaf weeds, respectively. The results indicated antagonistic effects between 2,4-D plus MCPA and clodinafop propargyl. The best tank mixture with regard to weed control efficacy was 2,4-D plus MCPA at 975 g ai ha⁻¹ with clodinafop propargyl at 96 g ai ha⁻¹. The wheat grain yield was also increased by the tank mixture of clodinafop propargyl with 2,4-D plus MCPA. Generally, to inhibit clodinafop propargyl efficacy reduction due to tank-mixing with 2,4-D plus MCPA, it is recommended that the application dose of 64 g ai ha⁻¹ should be increased to 96 g ai ha⁻¹.     

Effects of Azolla species on weed emergence in a rice paddy ecosystem

The effects of Azolla pinnata R. Br. on weed emergence were evaluated in terms of plot area coverage by an A. pinnata mat, its biomass production and the amount of weed emergence, using fresh and dry weights, in a rice paddy field experiment. The experiment was conducted following a randomized block design with five combinations of fertilizer and A. pinnata treatments (control, A. pinnata  + superphosphate, A. pinnata  + urea, A. pinnata  + compound fertilizer, A. pinnata  + cow manure). The results revealed that after 18 days of inoculation, all superphosphate (T1) and cow manure (T4)-treated plots were fully covered by the A. pinnata mat. However, coverage of the urea (T2) and compound fertilizer (T3)-treated plots were only 80% and 70%, respectively. The full plot area coverage by the A. pinnata mat and the highest biomass production with superphosphate and cow manure-treated plots were able to completely inhibit two weed species ( Scirpus juncoides Roxb. var. hotarui Ohwi and Monochoria vaginalis Burm. f. Presl var. plantaginea (Roxb.) Solms-Laub) and significantly suppressed four other weeds ( Cyperus serotinus Rottb, Echinochla oryzicola Vasing, and Eclipta prostrata L.). In all the treatments, the fresh weight of weeds significantly reduced to 13, 29, 34, and 9%, respectively, for treatments T1, T2, T3, and T4. The dry weights also were significantly reduced to 10.00, 16.00, 22.00, and 7.26%, respectively, for treatments T1, T2, T3, and T4 over the control. The results revealed that there was a significant correlation among plot area coverage by the A. pinnata mat, its biomass production and weed emergence in a rice paddy field over the control. Azolla pinnata did not have any detrimental effect on the growth of rice plants.     

Impact of cropping systems on the weed seed banks in the northern Great Plains, USA

Very little research has been done on weed seedbank communities in the northern Great Plains region of the USA. The objectives of this study were to evaluate the effects of management systems on the weed seed banks and to assess the relationship between the weed seed banks and the above-ground weed communities. The seed banks were sampled along a range of areas of above-ground weed diversity in organic and conventional spring wheat production fields near Big Sandy, Montana, over 2 years. Eight 1 m × 0.33 m areas were selected in each field to encompass a wide range of above-ground weed diversity and soil cores were taken in each area to a depth of 20 cm and split in half at 10 cm. The number of seeds recovered from the top 10 cm of soil and the 10–20 cm depth were significantly affected by the sampling year and an interaction between the cropping system and the sampling year. The year was the only significant predictor affecting the weed seedbank diversity at both depths. A significant interaction between the management system and the year observed at the 10–20 cm sampling depth reflected the higher rate of decrease in diversity occurring between 2005 and 2006 in the conventionally managed fields compared with the organic fields. A multivariate ordination indicated that, although the year played a significant role in determining the weed seedbank communities, the management system had a role only during 2006. We failed to detect strong correlations between the above-ground and underground weed communities. The results of this study suggest that, in the studied region, yearly fluctuations in environmental factors have significant impacts on the weed seed banks.     

Correlation between the soil seed bank and weed populations in maize fields

Annual weed populations establish every year from persistent seed banks in the soil. This 3 year study investigated the relationship between the number of weed seeds in the soil seed bank and the resultant populations of major broadleaf and grass weeds in 30 maize fields. After planting the crop, 1 m² areas were protected from the pre-emergence herbicide application. Soil samples were collected soon after spraying to a depth of 100 mm and the weed seeds therein were enumerated. The emerged weed seedlings in the field sampling areas were counted over the following 8 weeks. Up to 67 broadleaf species and five grass weeds were identified, although not all were found at every site and some were specific to a region or soil type. For the most abundant weeds in the field plots, on average 2.1–8.2% of the seeds of the broadleaf species and 6.2–11.9% of the seeds of the grass weeds in the soil seed bank emerged in any one year, depending on the species. Overall, the results showed a strong linear relationship between the seed numbers in the soil and the seedling numbers in the field for all the grasses and for most broadleaf weeds. For some species, like Trifolium repens , only a weak relationship was observed. In the case of Chenopodium album , which had the largest seed bank, there was evidence of asymptotic behavior, with seedling emergence leveling off at high seed numbers. An estimate of the soil seed bank combined with knowledge of the germination and behavior of specific weed species would thus have good potential for predicting future weed infestations in maize fields.     

小麦田不同杂草群落及防除时间对小麦产量的影响

为明确小麦田不同杂草群落及防除时间对小麦产量的影响,于2013—2015连续两年在山东省聊城市高唐县小麦田中设无草对照区、禾本科杂草区、阔叶杂草区以及混合杂草自然发生区4种不同杂草群落以及不同的杂草防除时间,测定在不同条件下小麦产量及各项产量构成指标的情况。结果表明,阔叶杂草对小麦产量的影响大于禾本科杂草,冬小麦田杂草的最佳防除时间为4月1日之前,在此期间除草对小麦产量影响均不大,但之后防除或不除草会造成小麦严重减产甚至绝产,2014年4月15日除草可造成在阔叶杂草区和杂草混合生长区的小麦减产30.5%和32.6%,不除草可造成在禾本科杂草区、阔叶杂草区和杂草混生区的小麦减产8.6%、91.4%和94.3%,2015年趋势和2014年一致。从构成小麦产量的3个指标来看,杂草危害主要影响小麦的总穗数,其次为穗粒数,对千粒重影响最小。     

Plant classification for weed detection using hyperspectral imaging with wavelet analysis

The goal of this study is to develop a new weed detection method that can be applied for automatic mechanical weed control. For successful weed detection, plants must be classified into crops and weeds according to their species. In this study, we employed a portable hyperspectral imaging system. The hyperspectral camera can capture landscape images that include crops, weeds, and the soil surface, and can provide more extensive information than conventional red, green, and blue (RGB) images. Although RGB images consist of red, green, and blue wavebands, the obtained hyperspectral images consist of 240 wavebands of spectral information. Hyperspectral imaging is expected to provide powerful technology for agricultural sensing. In the initial step of this study, the image pixels of the plants (crop or weeds) were segmented from the background soil surface using Euclidean distance as the discriminant function. In the next step, the image pixels of the crop (sugarbeet) and weeds (four species) were classified using the difference in the spectral characteristics of the plant species. In this process, classification variables were generated using wavelet transformation for data compression, noise reduction, and feature extraction, and then stepwise linear discriminant analysis was applied. The validation results indicate that the developed classification method has potential for practical use.     

Evaluation of the allelopathic potential of Kava (Piper methysticum L.) for weed control in rice

Kava is a perennial pepper plant from the Oceanic region, which is commonly used as a drink by natives and for pharmacological purposes. Results of this study concluded that Kava has allelopathic potential and suppressed germination and growth of lettuce ( Lactuca sativa L), barnyardgrass ( E. crus-galli Beauv var. formosensis Ohwi.), and duck-tongue weed ( M. vaginalis Presl var. plantaginea Solms-Laub.) Moreover, Kava inhibited emergence of weeds in paddy soils. The inhibition of Kava became stronger as the applied concentration increased. In a greenhouse experiment, Kava had the greatest inhibition on emergence and weed growth in paddy soils when a dose of 1 ton ha⁻¹ of Kava was divided and applied at 3, 16, and 23 days after watering. Inhibition by Kava varied among weed species. Results from this study suggest that Kava might be useful for weed control in rice or reduce dependence on herbicide.     

Weed infestation and tea growth under various weed management methods in a young tea (Camellia sinensis[L.] Kuntze) plantation

A field experiment was conducted in the low country of Sri Lanka, during the period 1994–1995 to investigate the severity of weed infestation and tea growth in relation to weed management methods in newly established tea ( Camellia sinensis [L.] Kuntze). Manual weeding (hand and slash weeding) at various intervals was compared with various herbicides, with or without mulching. Weed control with herbicides was superior to that of hand weeding at 6-week intervals or more. Weed control with oxyfluorfen at 0.29 kg ai ha⁻¹ + paraquat at 0.17 kg ai ha⁻¹ or glyphosate at 0.99 kg ai ha⁻¹ + kaolin at 3.42 kg ha⁻¹ were superior. Plots unweeded for 12 weeks or more produced significantly greater ( P  < 0.05) weed biomass than plots unweeded for 6 weeks. Although the least weed dry weight ( P  < 0.05) and the greatest number of weed species were recorded with hand weeding at 2 week intervals, there was no particular benefit on tea growth when compared with hand weeding at 6 and 12 week intervals. Inter row mulching in chemically treated plots was more favorable for tea growth than no mulching, while living weed cover in unmulched slash weeded plots suppressed tea growth. A combination of mulching and herbicides, particularly oxyfluorfen and paraquat, followed by hand weeding at least every 6–8 weeks was considered the most appropriate weed management system for young tea.     

Site-specific weed control technologies

Site-specific weed control technologies are defined as machinery or equipment embedded with technologies that detect weeds growing in a crop and, taking into account predefined factors such as economics, take action to maximise the chances of successfully controlling them. In this study, we describe the basic parts of site-specific weed control technologies, comprising weed sensing systems, weed management models and precision weed control implements. A review of state-of-the-art technologies shows that several weed sensing systems and precision implements have been developed over the last two decades, although barriers prevent their breakthrough. Most important among these is the lack of a truly robust weed recognition method, owing to mutual shading among plants and limitations in the capacity of highly accurate spraying and weeding apparatus. Another barrier is the lack of knowledge about the economic and environmental potential for increasing the resolution of weed control. The integration of site-specific information on weed distribution, weed species composition and density and the effect on crop yield, is decisive for successful site-specific weed management.     

Allelopathic potential of Trifolium resupinatum L. (Persian clover) and Trifolium alexandrium L. (Berseem clover)

This experiment was conducted in order to determine if Persian clover ( Trifolium resupinatum ) and Berseem clover ( Trifolium alexandrium ) contained water-soluble and methanol-soluble constituents that could affect the seedling growth of four test species; namely, Amaranthus retroflexus , Convolvulus arvensis (field bindweed), Secale cereale , and Sinapis arvensis (wild mustard) . The above-ground tissues of the Persian and Berseem clover plants were collected during the vegetative growth stage and oven-dried. Three concentrations of aqueous and methanolic extracts were used at full-strength (33.3 g L⁻¹), half-strength (16.7 g L⁻¹), and quarter-strength (8.3 g L⁻¹). Distilled water was used as the control. The weed seeds were placed in Petri dishes containing the legume extract or distilled water (control). The seedling growth of the weed species declined with the increasing concentration of the clover extracts. Wild mustard exhibited the highest sensitivity to both the legume extracts. Compared to the aqueous extract, the methanolic extract caused a greater decline in the seedling growth of the weeds. Berseem clover was the stronger inhibitor of the weed seedling growth, as compared to Persian clover. Therefore, the amounts of allelochemicals present might differ in these two clover species. Field bindweed showed the least sensitivity to both the legume extracts. Field bindweed showed more tolerance to the allelochemicals produced by the clover species.     

Phosphorus fertilizer effects on the competition between wheat and several weed species

Information on phosphorus (P) fertilizer affecting crop–weed competitive interactions might aid in developing improved weed management systems. A controlled environment study was conducted to examine the effect of three P doses on the competitive ability of four weed species that were grown with wheat. Two grass and two broad-leaved weed species were chosen to represent the species that varied in their growth responsiveness to P: wild oat (medium), Persian darnel (low), round-leaved mallow (high), and kochia (low). Wheat and each weed species were grown in a replacement series design at P doses of 5, 15, and 45 mg P kg⁻¹ soil. The competitive ability of the low P-responsive species, Persian darnel and kochia, decreased as the P dose increased, supporting our hypothesis that the competitiveness of species responding minimally to P would remain unchanged or decrease at higher P levels. As expected, the competitiveness of the high P-responsive species, round-leaved mallow, progressively improved as the P dose increased. However, wild oat's competitive ability with wheat was not affected by the P fertilizer. The results suggest that fertilizer management strategies that favor crops over weeds might deserve greater attention when weed infestations consist of species known to be highly responsive to higher soil P levels. The information gained in this study could be used to advise farmers of the importance of strategic fertilizer management in terms of both weed management and crop yield.     

Long‐term changes in weed occurrence,yield and use of herbicides in maize in south‐western Germany,with implications for the determination of economic thresholds

In 393 field experiments in Baden‐Württemberg region in south‐western Germany, herbicide efficacy, yield loss and crop tolerance of maize (Zea mays) were investigated between 1981 and 2011. The collected data served to determine changes in weed frequencies, in herbicide use, yield loss functions and economic thresholds (ETs). Over 60 weed species were reported. Chenopodium album and Galium aparine were the most frequent broad‐leaved weeds, the former becoming more frequent over time. Species of the genera Lamium, Polygonum, Veronica and Matricaria occurred in about every fifth trial. Alopecurus myosuroides and Echinochloa crus‐galli were the most frequent grass weeds; the former declining in frequency by 1.1% per year, the latter increasing by 1.5%. Results suggest a weed population shift towards thermophilic species. aceto‐lactate‐synthase and 4‐HPPD‐inhibitor herbicides became important in the 1990s. Pendimethalin and bromoxynil have been integral components of weed control since the 1980s. ETs, the point at which weed control operations provide economic returns over input costs, ranged between 3.7% and 5.8% relative weed coverage. Without weed control, no yield increase was found over 24 years. Yield increased by 0.2 t ha ^{? 1} year ^{? 1}, if weeds were controlled chemically. Despite intensive use of effective herbicides in maize, problematic weed species abundance and yield losses due to weed competition have increased in Baden‐Württemberg over a period of 30 years.     

Real-time weed detection, decision making and patch spraying in maize, sugarbeet, winter wheat and winter barley

Information on temporal and spatial variation in weed seedling populations within agricultural fields is very important for weed population assessment and management. Most of all, it allows a potential reduction in herbicide use, when post‐emergence herbicides are only applied to field sections with weed infestation levels higher than the economic weed threshold; a review of such work is provided. This paper presents a system for site‐specific weed control in sugarbeet (Beta vulgaris L.), maize (Zea mays L.), winter wheat (Triticum aestivum L.) and winter barley (Hordeum vulgare L.), including online weed detection using digital image analysis, computer‐based decision making and global positioning systems (GPS)‐controlled patch spraying. In a 4‐year study, herbicide use with this map‐based approach was reduced in winter cereals by 60% for herbicides against broad‐leaved weeds and 90% for grass weed herbicides. In sugarbeet and maize, average savings for grass weed herbicides were 78% in maize and 36% in sugarbeet. For herbicides against broad‐leaved weeds, 11% were saved in maize and 41% in sugarbeet.     

Effects of crop management practices on Echinochloa crus-galli and Chenopodium album seed production in a maize/soyabean rotation

Seed production of residual weed populations needs to be taken into account when estimating the long-term impact of low-input agronomic practices. The objective of this study was to measure the effects and interactions of crop, weed control, tillage practice and nutrient source on the seed production of the dominant residual weed species in a maize/soyabean rotation at two sites: Echinochloa crus-galli (L.) Beauv. on a Sainte-Rosalie clay and Chenopodium album L. on a Duravin clay loam. Seed production per unit area was estimated in each experimental unit. Weed seed production was greater under mechanical weed control compared with chemical weed control. In 1997, E. crus-galli seed production reached over 326 000 seeds m^–2 in mechanical weed control treatments, but averaged less than 500 seeds m^–2 in the chemical weed control treatments. Chenopodium album produced in the range of 766 000 and 73 000 seeds m^–2 in mechanical and chemical weed control treatments respectively. Very few or no weed seeds were produced in soyabean under chemical control. Tillage intensity and nutrient source did not affect seed production of either weed species, with the exception that E. crus-galli produced more seeds in chisel than in mouldboard plough tillage in soyabean. Weed control method had more impact on seed production than tillage intensity and nutrient source in a maize/soyabean rotation.