Measuring ecologically important quantities at relevant operational scales: An example using the dusky flycatcher in selectively logged forests in Idaho

Effective management of forest wildlife requires information about habitat features that can be altered by operations, if those habitat features promote reproductive success and survival, and if changes in organism density influence reproductive success and survival. Habitat selection studies often provide the first type of information, although relatively few studies examine how different selection decisions or densities of organisms influence fitness measures or measure habitat features at relevant operational scales. We examined patterns of habitat selection and estimated how habitat use, territory size, and conspecific density were associated with territory success (probability of a territory producing one or more fledglings) for the dusky flycatcher in 2002–2003 on eight study plots in managed grand fir (Abies concolor) forests in Idaho, USA. Flycatchers selected territories with greater cover of deciduous vegetation in two different height strata (1–2 and 3–4 m) and reduced cover of conifers less than 4 cm dbh compared to what was available. While flycatchers selected habitat characteristics at the territory scale, model selection results did not support any associations between vegetation features and territory success. Territory success had weak positive associations, for a given territory size, with conspecific density, although the relationship was highly variable. An interaction between territory size and density was not supported. These results are consistent with the view that avian breeding habitat selection occurs at multiple scales, and that the nest site scale may explain more of the variance in reproductive success than larger scales. Our results suggest that operational management of vegetation features, which often occurs at the stand scale, may not influence variation in critical demographic metrics. Increasing total shrub cover of commonly used plant species should benefit the dusky flycatcher, as well as other shrub-nesting forest birds, but stand-level habitat alterations alone cannot be relied upon to increase reproductive success, an important component of population change.     

湖南黄桑国家级自然保护区植被景观格局探讨

①斑块特征指数分析表明:各景观类型的密度、分维数指数都较小,植被景观的深展指数都不高,扩散比较均匀,没有极端复杂的形状产生,植被景观在演替系列中还没有普遍达到演替的顶级,斑块之间自相似程度较高。②多样性指数分析表明:总体多样性程度高。景观多样性指数、优势度指数与均匀度指数的结果所包含的意义一致,这种一致性充分说明了各植被景观类型分布均匀,各景观类型地位相差较小。③空间构型指数分析表明:保护区的斑块密度为1.001,即1 km2上有1.001个斑块。这说明斑块数目较少。各景观类型分布也较均匀。保护区景观破碎度指数为0.007,这表明该区植被景观在研究期内所表现的生境破碎化程度是很低的。每一植被景观类型破碎度指数最高的是灌丛和灌草丛景观,其次为栲林景观,其余的破碎度都处于较低的水平。内部生境破碎化指数以灌丛和灌草丛景观、栲林景观为最低,它们的单独斑块面积较大,故破碎化指数较低。其它景观类型的破碎化指数相差不大,这与植被顺向演替是相关的。景观分离度指数的排序结果与斑块密度的排序基本一致。     

西双版纳自然保护区鼷鹿生境选择的初步研究

对中国仅见于西双版纳勐腊县鼷鹿（ＴｒａｇｕｌｕｓｊａｖａｎｉｃｕｓｗｉｌｉａｍｓｏｎｉＫｌｏｓ）栖息地生境因子的调查分析，初步得出鼷鹿栖息环境与植被类型、下木、地被、坡度、坡向、海拔、土质的相关程度。对各生境因子的优先选择顺序分别为：下木郁闭度，地表覆盖物，坡度，植被类型，土质，海拔，坡向。并初步划分出鼷鹿的４个生境等级为：（１）最佳生境地；（２）良好生境地；（３）一般生境地；（４）劣等生境地。并在此基础上提出相应的保护对策。     

三江自然保护区丹顶鹤春季觅食生境初步研究

采用样方因子测定法对三江国家级自然保护区丹顶鹤春季觅食生境进行调查研究的结果表明:丹顶鹤春季主要在以苔草、小叶章为主的沼泽湿地中觅食;觅食地植被高度小于40 cm,植被密度低于400株/m2,植被盖度为0～25%,觅食地水深多为5～15 cm,人类干扰距离大于1 500m;人为活动严重影响着丹顶鹤的觅食;春季丹项鹤有在...     

A study of habitat fragmentation in Southeastern Brazil using remote sensing and geographic information systems (GIS)

The purpose of this work was to study fragmentation of forest formations (mesophytic forest, riparian woodland and savannah vegetation (cerrado)) in a 15,774-ha study area located in the Municipal District of Botucatu in Southeastern Brazil (São Paulo State). A land use and land cover map was made from a color composition of a Landsat-5 thematic mapper (TM) image. The edge effect caused by habitat fragmentation was assessed by overlaying, on a geographic information system (GIS), the land use and land cover data with the spectral ratio. The degree of habitat fragmentation was analyzed by deriving: 1. mean patch area and perimeter; 2. patch number and density; 3. perimeter-area ratio, fractal dimension (D), and shape diversity index (SI); and 4. distance between patches and dispersion index (R). In addition, the folowing relationships were modeled: 1. distribution of natural vegetation patch sizes; 2. perimeter-area relationship and the number and area of natural vegetation patches; 3. edge effect caused by habitat fragmentation. The values of R indicated that savannah patches (R = 0.86) were aggregated while patches of natural vegetation as a whole (R = 1.02) were randomly dispersed in the landscape. There was a high frequency of small patches in the landscape whereas large patches were rare. In the perimeter-area relationship, there was no sign of scale distinction in the patch shapes. In the patch number-landscape area relationship, D, though apparently scale-dependent, tends to be constant as area increases. This phenomenon was correlated with the tendency to reach a constant density as the working scale was increased. On the edge effect analysis, the edge-center distance was properly estimated by a model in which the edge-center distance was considered a function of the total patch area and the SI.     

Importance of habitat type classifications for predicting ruffed grouse use of areas for drumming

To incorporate ruffed grouse (Bonasa umbellus) habitat planning in forest management, it is necessary for managers to understand factors contributing to grouse habitat use. Previous studies examining ruffed grouse drumming habitat documented relationships between drumming grouse and broad vegetation categories (e.g., northern hardwoods, young aspen [Populus spp.], oak [Quercus spp.]), but few studies have documented how drumming grouse respond to ecological variations in site conditions of aspen or other vegetation types that might be used. Our objectives were to determine the utility of habitat type classifications in predicting the occurrence of ruffed grouse drumming habitat in the western Upper Peninsula of Michigan, and demonstrate how classifications may be used to understand how forest management may affect ruffed grouse habitat. We used survey routes on state land and conducted drumming surveys during mid-April and early May at 78 points in 2005 and 2006. We recorded the number of drumming males heard at each point, the azimuth to where the grouse was heard, and a qualitative measure of distance to determine in which forest stands grouse were drumming. Using GIS, we determined the specific vegetation type, age class, and habitat type, evaluated habitat suitability, and determined a suitability score for areas in which grouse were drumming. We constructed a logistic regression model that calculated the probability of grouse use of areas for drumming based on vegetation characteristics at used and random locations. Our results indicated that the probability of grouse use of an area for drumming is based on inherent site characteristics (i.e., habitat type) and habitat suitability. The model is useful for planning forest management activities and understanding how grouse may respond to spatial or temporal changes in vegetation through succession or manipulation.     

Vegetation community and soil characteristics of abandoned agricultural land and pine plantation in the Qinling Mountains,China

In order to determine the changes in the characteristics of the vegetation and soil following agricultural abandonment and compare the effects of different restoration approaches on ecosystem recovery, we studied the vegetation community and soil characteristics (nutrients, bulk density, water content and pH) of Pinus tabulaeformis plantations and abandoned croplands in different successional stages in the Foping National Nature Reserve, located in the Qinling Mountains, northwest China. The results indicated that natural vegetation and habitat could be restored via natural regeneration. These spontaneous restoration forests were characterized by high diversity, high soil fertility and rich unique species. The soil organic matter, total carbon, total nitrogen, available potassium, community cover, depth of litter, depth of humus and soil water content increased significantly with years after abandonment, while the total potassium, total phosphorus, available phosphorus, soil pH did not seem to change significantly with abandonment time. The soil mineral nitrogen (NH₄-N + NO₃-N) tended to increase during the first 50 years after abandonment and then decreased. The pine plantations tended to show a low level of biodiversity in tree and herb layer, but the shrub layer (including sapling) composition and diversity were similar to secondary forests. Although the pine plantation showed lower soil fertility, they did not seem to result in the habitat-degradation.     

Do the size and landscape context of forest openings influence the abundance and breeding success of shrubland songbirds in southern New England?

Early successional birds have declined in the northeastern United States due to the regeneration of forest on abandoned farm fields and the suppression of natural disturbances that once provided appropriate habitat. These species have become increasingly dependent on early successional habitats generated by such activities as timber harvesting. Recent approaches of timber harvesting, which range from single-tree harvesting to clearcutting, create forest openings of different sizes and configurations embedded in landscapes with different land use patterns. To assess the importance of forest openings created by timber harvesting for shrubland birds, we surveyed birds on 50 m radius plots in 34 harvest sites (0.5–21 ha). We collected data on multi-scaled habitat variables ranging from plot-level vegetation characteristics to land use patterns within 1 km of each study site. We also monitored mating and nesting success of Blue-winged Warblers (Vermivora pinus) in 10 forest openings.

The abundance of most shrubland species was influenced by plot-level habitat variables, such as tree density and vegetation height, rather than shrubland area or the composition of land uses in the surrounding landscape. Only Eastern Towhees (Pipilo erythrophthalmus) were more frequent in survey plots in larger forest openings. In contrast, neither abundance nor reproductive activity of Blue-winged Warblers was correlated with the size of the forest opening. Their abundance was negatively related to vegetation height, however. Only 54% of the territorial male Blue-winged Warblers in forest openings were mated. We documented relatively low nest success rates of 21.1% during the egg laying and incubation nest stages, but increased success rates during the later stages of nesting.

Our results indicate that even small forest openings with low vegetation provide habitat for Blue-winged Warblers and other shrubland birds. The overall reproductive rate of territorial male Blue-winged Warblers in forests openings was low during the 2 years of the study, however. Further studies are needed to assess the long-term value of this type of habitat for sustaining shrubland bird populations.     

Habitat assessment for forest dwelling species using LiDAR remote sensing: Capercaillie in the Alps

Large-scale information on habitat suitability is indispensable for planning management actions to further endangered species with large-spatial requirements. So far, remote sensing based habitat variables mostly included environmental and land cover data derived from passive sensors, but lacked information on vegetation structure. This is a serious constraint for the management of endangered species with specific structural requirements. Light detection and ranging (LiDAR), in contrast to passive remote sensing techniques, may bridge this gap in structural information at the landscape scale. We investigated the potential of LiDAR data to quantify habitat suitability for capercaillie (Tetrao urogallus), an endangered forest grouse in Central Europe, in a forest reserve of 17.7 km². We used continuous variables of horizontal and vertical stand structure from first and last pulse LiDAR data and presence–absence information from field work to model habitat suitability with generalized linear models (GLM). The two final habitat suitability models explained the observed presence–absence pattern moderately well (AUC of 0.71 and 0.77) with horizontal structure explaining better than vertical structure. Relative tree canopy cover was the most important variable with intermediate values indicating highest habitat suitability. As such, LiDAR allowed us to translate the results from habitat modeling at the landscape scale to effective management recommendations at the local scale at a level of detail that hitherto was unavailable for large areas. LiDAR thus enabled us to integrate individual habitat preferences at the scale of entire populations and thus offers great potential for effective habitat monitoring and management of endangered species.     

内蒙古图牧吉自然保护区丹顶鹤春季觅食生境初步研究

于2008年5月1~7日,对内蒙古图牧吉国家级自然保护区丹顶鹤觅食地进行了生境因子调查.分析认为,丹顶鹤觅食地主要属沼泽地,其植物主要为芦苇、苔草、水葱、香蒲,植被高度多为100~150 cm,水深约10 cm,水质良好,透明度约为10 cm.     

Mulching effects on vegetation recovery following high severity wildfire in north-central Washington State,USA

Straw mulch application after high severity wildfire has gained favor in recent years due to its efficacy in reducing soil erosion hazards. However, possible collateral effects of mulching on post-fire vegetation recovery have received relatively little study. We assessed mulching effects on plant cover and species richness, tree seedling establishment, and exotic species densities in the second year following the 2006 Tripod Wildfire in north-central Washington State, USA, by observing vegetation responses to spatial variability in mulch cover and depth. Mulch cover averaged about 35%, with a median depth of 0.5 cm. Vegetation recovery was generally slow, with median plant cover of only 10%. Tree seedling densities were low and spatially variable. Vegetative cover, species richness, and seedling densities all declined with increasing elevation. Mulch cover was positively associated with plant cover, plant species richness, and conifer seedling densities when second year mulch cover did not exceed 40%. Only when mulch cover exceeded 70% did mulching begin to negatively affect vegetation recovery relative to areas with no mulch. Vegetation responses to mulch depth were minimal at depths under 3 cm, but quite strong when mulch depth exceeded 5 cm. Exotic plant frequency and density were positively associated with mulch cover, but exotic plant cover was low on average (<1%). In this study, mulch added significant cover to sites with slow natural recovery of vegetation, thereby likely reducing erosion hazard. Mulching also appears to have facilitated native plant recovery and conifer seedling establishment except at very high application levels, easing management concerns about longer-term impacts of mulching treatments on post-fire vegetation recovery.     

基于MAXENT生态位模型对内蒙古根河地区驼鹿生境适宜性评价

2013～2015年冬季,综合运用动物生态调查、种群生态学和行为生态学等技术,在内蒙古根河林业局共布设71条长约2～4 km样线、调查313个10 m×10 m样方、1 489个2 m×2 m样方、共收集117份粪便,373个驼鹿出现点数据。利用驼鹿出现数据作为分布点数据,选取地形、植被类型等22种因子作为生境变量,利用MAXENT生态位模型分析了深冬时期根河地区驼鹿冬季生境适宜性分布特征和主要生境因子对驼鹿分布的影响。结果表明:模型预测结果准确性较高,平均AUC(area under the curve,受试工作者曲线下面值)值为0.92,Jackknife检验结果显示:驼鹿生境选择的主要影响因子为距道路距离、距火烧迹地距离、距农田距离、海拔、柳、距沼泽距离、距人为活动点距离、距灌丛距离。驼鹿的适宜生境在根河地区呈现连续性分布,主要分布在东部和东北部,从植被类型上看,适宜生境大部分分布于针叶林,且适宜生境的面积为1 155.8 km^2,约占根河研究区总面积的25.6%。     

基于MODIS卫星影像的湖南省景观格局

景观格局是由自然或人为形成的一系列大小、形状各异,排列不同的景观要素共同作用的结果.利用2005年MODIS卫星数据,经过一系列处理后得到湖南省景观图.利用决策树分类法将研究区分为5种景观类型:水、自然植被、耕地、城镇用地、其它用地.选取斑块数、斑块密度、破碎度、斑块分维数、多样性指数、优势度、均匀度、聚集度、分离度等景观指数,从斑块、斑块类型和景观3个层次上对研究区景观格局进行定量分析.结果表明:湖南省景观斑块共39357块,总面积为21.1984万km2;在5种景观类型中,植被和耕地是主要景观类型,占研究区总面积的88.38%;湖南省景观多样性程度低,且斑块空间形状简单.     

Impacts of different forest management scenarios on forestry and reindeer husbandry

In northern Sweden, the forests are used simultaneously for both timber production and reindeer husbandry. During the winter months, lichen is the most important fodder for reindeer. Forest management operations are generally considered having a negative impact on reindeer husbandry as harvesting and dense stands remove or obscure the ground lichen cover. In this study, we simulate three different scenarios for forest management, differing in the intensity and types of harvest operations. The resulting 100-year scenarios are analyzed with respect to their estimated suitability for providing reindeer pasture areas. Suitability is determined by vegetation type, stand density and stand height. The results indicate that the current trend of a decrease in lichen area will continue if existing forestry practice prevails. Implementing continuous cover forestry as a management alternative and carrying out precommercial thinning could halt the decrease in reindeer pasture area and even lead to a future increase in pasture area, with losses of approximately 5% in the net present value of forestry.     

广西大青山森林植物分布立地适宜性与土壤关系的研究

本文将大青山范围植被、土壤垂直带内的主要森林植物,分草、灌、乔及不同科、属,着重个体生态的共性与特性,探讨了它们分布立地的适宜性及化学成分与土壤肥力的关系,并简要说明其在该处土壤、母质母岩上的生产量。     

Characterizing woodland caribou habitat in sub-boreal and boreal forests

Woodland caribou (Rangifer tarandus caribou) are sensitive to changes in understory vegetation resulting from forest harvesting and are, therefore, of special concern for foresters and habitat biologists. Effective management of this species requires reliable habitat inventories which, because of the large heterogeneous areas over which caribou range, can be costly. We used Landsat Thematic Mapper (TM) imagery and digital elevation data to identify 23 vegetative cover types across the 5100 km² range of the Wolverine caribou herd of northcentral British Columbia, Canada. The classification was augmented with available geographical information system (GIS) data for a total of 27 cover types. We achieved an overall accuracy of 76.7% based on known ground samples; however, accuracy varied according to cover type. Considering the size of the study area, the procedure we employed was relatively cost effective and efficient. We discuss the advantages of such an approach for wildlife-habitat studies reliant on large-scale vegetation maps.     

A weeding-duration model for Abies sachalinensis plantations in Hokkaido, northern Japan

I developed a weeding-duration model for Sakhalin fir (Abies sachalinensis (Fr. Schmidt) Masters) plantations that employs a generalized linear model. The number of years following planting that weeding is necessary is the response variable, and elevation, slope steepness, maximum snow depth, annual precipitation, geology, soil, site index, slope aspect, and vegetation type are explanatory variables. Among the explanatory variables, geology, soil, slope aspect, and vegetation type are categorical data. A Poisson distribution is assumed for the response variable, with a log-link function. Elevation, slope steepness, maximum snow depth, annual precipitation, site index, and vegetation type had a significant effect on weeding duration. Among the eight models with the smallest Akaike information criterion (AIC), I chose the model with no multicollinearity among the explanatory variables. The weeding-duration model includes site index, maximum snow depth, slope steepness (angle) and vegetation type as explanatory variables; elevation and annual precipitation were not included in the selected model because of multicol- linearity with maximum snow depth. This model is useful for cost-benefit analyses of afforestation or reforestation with Abies sachalinensis.     

Ruffed grouse-habitat preference in the central and southern Appalachians

Ruffed grouse (Bonasa umbellus; hereafter grouse) populations in the central and southern Appalachians (CSA) are declining due to widespread maturation of forest cover. Effective management of this species requires a sex- and age-specific understanding of habitat preferences at multiple temporal and spatial scales. We used multivariate logistic regression models to compare habitat within 1440 grouse home ranges and 1400 equally sized buffered random points across 7 CSA study areas. On most sites, grouse home ranges were positively associated with roads and young forest (<20 years old). Sex and age status affected habitat preference. In general, males used younger forest than females, likely because of differences in habitat use during reproductive periods. Juveniles had fewer vegetation types preferred by adult grouse and more of the avoided vegetation types within their home ranges, indicative of competitive exclusion. Adult females had the greatest specificity and selectivity of habitat conditions within their home ranges. Habitat selection varied among seasons and years on most sites. Winter habitat use reflected behavior that maximized energy conservation, with open vegetation types avoided in the winter on the northernmost study areas, and topography important on all areas. Summer habitat selection reflected vegetation types associated with reproductive activities. Scale influenced habitat preference as well. Although roads and forest age predominantly influenced grouse home range location within the landscape, mesic forest types were most important in determining core area use within the home range. This was likely a result of increased food availability and favorable microclimate. Habitat management efforts should attempt to maintain ∼3–4% of the landscape in young forest cover (<20 years old), evenly distributed across management areas. Roads into these areas should be seeded as appropriate to enhance brood habitat and provide travel corridors connecting suitable forest stands.     

Effectiveness of nature reserves for bird conservation in urban parks in Tokyo

In some urban parks in Tokyo,semi-natural habitat patches are maintained as nature reserves for birds,called bird sanctuaries.Bird censuses and vegetation surveys were conducted in eight urban parks in Tokyo from December 2015 to July 2016 to determine the effectiveness of bird sanctuaries on avian species composition.The relationship between avian species composition and environmental conditions was analyzed by partial redundancy analysis(partial RDA) using vegetation variables,number of visitors,presence or absence of reserves within the same park,area of wooded parts,and normalized difference vegetation index(NDVI)in the surrounding area.The results of the partial RDA are as follows:(1) Area,lower vegetation cover,higher vegetation cover,tree species composition obtained from detrended correspondence analysis(DCA) on vegetation survey data,and NDVI in the surrounding area were considered as covariates and explained 17.4-33.6% of the total variation in bird species composition,(2) The presence or absence of sanctuary significantly explained bird species compositional variation regardless of season,indicating that the sanctuaries were beneficial for urban avoider species not only in the sanctuaries but also in the surrounding wooded area,and(3)Tree coverage within a 100 m range and leaf litter coverage also influenced avian species composition.We concluded that bird sanctuaries and other nature reserves can be beneficial to bird conservation,even within Tokyo.The positive effect of sanctuaries could be due to the limited influence of human presence and the developed vegetation within them.These reserves may play the role of core habitats within the studied urban parks.     

The relationship between cavity-nesting birds and snags on clearcuts in western Oregon

Relationships between cavity-nesting birds (CNB) and density and characteristics of snags were investigated on 13 clearcuts in central coastal Oregon. Species richness and density of CNB were positively (P<0.05) related to snag density and were still increasing at the maximum snag density evaluated. Cavity-nesting birds selected (P<0.05) snags taller than 6.4 m and greater than 78–102 cm in diameter, and avoided (P<0.05) snags less than 28 cm in diameter. Snags of intermediate decay stages were used for nesting more (P<0.05) than snags of early and advanced stages of decay. Cavity-nesting birds selected snags with more (P<0.05) bark cover (greater than 11%) than the average cover found on available snags. Individual CNB species exhibited significantly different (P<0.05) selections for snag height, diameter, hardness and bark cover. To optimize density and richness of CNB, forest managers should provide ≥ 14 snags ha⁻¹ between 28 and 128 cm diameter at breast height (dbh), between 6.4 and 25 m tall, with at least 10% bark cover, and with a majority in hardness stages 3 and 4.