Persistence and colonization of gaps in sown swards of grass and clover under different sward managements

The effects of extensive sward management and patch size on the persistence and colonization of gaps in sown swards was examined by creating gaps of five different sizes (2·3, 7, 10, 14 and 19 cm in diameter) in four different sward treatments: a fertilized sward grazed to 4 cm, i.e. relatively intensive management, and three extensively managed unfertilized swards, which were not grazed or grazed to 4 cm or 8 cm. The swards were originally sown with ryegrass (Lolium perenne L.) and white clover (Trifolium repens L.), but had developed differences in species composition as a result of the management treatments imposed 2 years before and during the experiment. Light quality measurements, i.e. red-far red (R/FR) ratio, were used to determine when the light environment in the gaps no longer differed from that in control, uncut patches and this was used as an estimate of gap persistence. Persistence of gaps depended on both sward management and gap size. Gaps disappeared most rapidly in the ungrazed sward and fertilized 4-cm sward, and most slowly in the unfertilized 8-cm sward. Small gaps persisted for up to 2 weeks in all but 8-cm swards, whereas larger gaps were estimated to persist for up to 20–25 weeks in unfertilized, grazed swards. There was no evidence that the number of grass or dicotyledonous species increased in the gaps compared with the control areas. There were significant positive linear relationships between the vegetation that developed in gaps and that in the control, uncut patches, reflecting the different species composition of the established sward of the grazed (grass-dominant) and ungrazed (Ranunculus repens-dominant) treatments. For total grass dry matter and tiller numbers, as well as L. perenne tiller numbers, there was a small, but significant, effect of both patch size and sward management on the slopes of the regressions between the controls and gaps. The results are discussed in relation to the potential for species composition of sown swards to change as a result of gap creation.     

Relationship between vegetation state and bite dimensions of sheep grazing contrasting plant species and its implications for intake rate and diet selection

This study used both experimental evidence and a mathematical model to address some differences in interpretation in the literature on the relationship between sward height and the bite dimensions (bite depth, bite area and bite mass) of sheep grazing contrasting vegetation types. Individual non-fasted sheep were released onto small areas (10 × 10m) of white clover (Trifolium repens) and perennial ryegrass (Lolium perenne) and bite dimensions were measured as they grazed across patches (0·7 × 0·7 m) of predetermined sward surface height (SSH). Sward heights were 4, 7, 10, 13 and 16cm for white clover patches and 4, 6, 5, 9, 11·5 and 14cm for ryegrass patches. Four sheep were assigned to each plant species and each sheep grazed one patch of each height (five patches/sheep). Bite depth, bite area and bite mass increased linearly with SSH in both white clover and ryegrass. At a given SSH, bite depth was similar in white clover and ryegrass, but bite area and bite mass were greater in white clover than in ryegrass. The linear relationships observed between bite mass and SSH contrasted with the asymptotic relationships observed in some other studies, but it is suggested that different relationships may arise because of methodological differences between studies. Furthermore, when linear relationships for bite mass were compared with asymptotic relationships for bite mass in a mechanistic model of animals grazing from ryegrass-white clover pastures it was demonstrated that the nature of the relationship had relatively little effect on the relationship between intake rate and SSH. This was because intake rate depended on the fundamental mechanistic relationship between bite mass and prehension bite rate. This relationship meant that the greater bite masses found when linear relationship were assumed were associated with reduced prehension bite rates and thus the effect on intake rate was relatively small. In addition, the predictions of the model regarding the bases of diet selection by animals grazing ryegrass-white clover mixtures were simplified, and stabilized, when linear relationships were assumed.     

Effects of grazing by sheep or goats on sward composition in ryegrass/white clover pasture and on subsequent performance of weaned lambs

An experiment was conducted to assess the effects of grazing a perennial ryegrass (Lolium perenne) / white clover (Trifolium repens) sward by sheep or goats on sward composition and structure and on subsequent diet selection, herbage intake and liveweight gain by weaned lambs. From mid-May to late July (phase 1), ewes with twin lambs or yearling Scottish Cashmere goats grazed continuously swards maintained at 4- or 8-cm sward surface height. From mid-August to the end of September (phase 2), weaned lambs continuously grazed the same swards maintained at 4 cm (treatment 4–4) or at 8 cm (treatment 8–8) or which had been allowed to increase from 4 cm to 8 cm (treatment 4–8). By the end of phase 1, swards grazed by goats had higher proportions of white clover in the whole sward (0.377 vs. 0.181; s.e.d 0.0382; P < 0.001) than those grazed by sheep, irrespective of sward height treatment. This resulted in phase 2 in a higher proportion of white clover selected ( P <0.001), higher herbage intakes ( P < 0.001) and higher liveweight gains ( P < 0.001) by weaned lambs grazing swards previously grazed by goats compared with those previously grazed by sheep. There were higher proportions of clover present in the swards from treatment 4–8 at the beginning of phase 2 compared with the other sward height treatments and consequently weaned lambs had, on this treatment, a higher proportion of clover in their diet ( P <0.001), higher herbage intakes ( P <0001) and higher liveweight gains ( P <0.001). It is concluded that goats can be integrated into sequential grazing systems with sheep on grass/clover swards and this can result in an increase in the proportion of clover in swards and increased sheep performance.     

Effects of grazing by sheep or cattle on sward structure and subsequent performance of weaned lambs

The study was designed to test the hypothesis that grazing management in early season could alter sward structure to facilitate greater animal performance during critical periods. The effects of grazing a mixed perennial ryegrass/white clover sward at different sward surface heights, by cattle or sheep, in early season on sward composition and structure, and on the performance of weaned lambs when they subsequently grazed these swards in late season were determined. In two consecutive years, from mid‐May until mid‐July, replicate plots (three plots per treatment) were grazed by either suckler cows and calves or ewes and lambs at 4 or 8 cm sward surface heights (Phase 1). From mid‐August (Year 1) or early August (Year 2), weaned lambs continuously grazed, for a period of 36 d (Year 1) or 43 d (Year 2) (Phase 2), the same swards maintained at 4 cm (treatment 4–4), 8 cm (treatment 8–8) or swards which had been allowed to increase from 4 to 8 cm (treatment 4–8). Grazing by both cattle and sheep at a sward surface height of 4 cm compared with 8 cm in Phase 1 resulted in a higher (P < 0·001) number of vegetative grass tillers per m² in Phase 2, although the effect was more pronounced after grazing by sheep. Sheep grazing at 8 cm in Phase 1 produced a higher number of reproductive tillers per m² and a greater mass of reproductive stem (P < 0·001) than the other treatment combinations. The mass of white clover lamina was higher under cattle grazing (P < 0·05), especially on the 8‐cm treatment, and white clover accounted for a greater proportion of the herbage mass. These effects had mainly disappeared by the end of Phase 2. On the 4–4 and 8–8 sward height treatments the liveweight gain of the weaned lambs was higher (P < 0·05) on the swards previously grazed by cattle than those grazed by sheep. The proportion of white clover in the diet and the herbage intake also tended to be higher when the weaned lambs followed cattle. However, there was no difference in liveweight gain, proportion of white clover in the diet or herbage intake between swards previously grazed by cattle or sheep on the 4–8 sward height treatment. It is concluded that grazing grass/white clover swards by cattle compared with sheep for the first half of the grazing season resulted in less reproductive grass stem and a slightly higher white clover content in the sward, but these effects are transient and disappear from the sward by the end of the grazing season. They can also be eliminated by a short period of rest from grazing in mid‐season. Nevertheless these changes in sward structure can increase the performance of weaned lambs when they graze these swards in late season.     

A comparison of diploid and tetraploid perennial ryegrass and tetraploid ryegrass/white clover swards under continuous sheep stocking at controlled sward heights. 3. Sward characteristics and animal output, years 4–8

Results for years 4–8 of a long-term grazing experiment on swards of a diploid perennial ryegrass (Lolium perenne), var. Contender (D swards), a tetraploid ryegrass, var. Condesa (T swards) and Condesa with S184 white clover (Trifolium repens) (TC swards), direct sown in May 1987, are presented. The swards were continuously stocked with sheep from 1988 to 1990, as previously reported, and for a further 5 years, 1991–95, at a target sward surface height (SSH) of 4–6 cm. Control of sward height was successfully achieved by variable stocking, except in 1993 when paddocks were set stocked and the resulting mean SSH was 9·3 cm. Grass swards received on average 160 kg N ha^?1 year^?1; grass/clover swards were mainly not fertilized with N with the exception that they were given 30 kg N ha^?1 as a remedial mid-summer application during a period of low herbage mass on offer in 1994 and 1995. Mean white clover content of the swards fell from 18·2% of herbage dry-matter (DM) in 1992 to 8·5% in 1993, whereas stolon lengths fell from 120 to 58 m m^?2. A return to lower sward heights in 1994–95 resulted in an increase in white clover content to 12·8% by the final sampling in August 1995. Perennial ryegrass content of the grass swards remained high throughout (mean 96·7% in 1995). Perennial ryegrass tiller densities recorded in August 1991, 1993 and 1994 showed consistently significant (P < 0·001) sward differences (3-year mean 16 600, 13 700 and 10 100 perennial ryegrass tillers m^?2 for the D, T and TC swards). In 1994, the year after lax grazing, a low perennial ryegrass tiller density (9100 m^?2) and low white clover content (mean 4·3%) in the TC swards resulted in a much lower herbage bulk density than in the grass swards (April–July means 72, 94 and 44 kg OM ha^?1 cm^?1 for the D, T and TC swards). There was a consistent 40 g d^?1 increase in lamb liveweight gain on the TC swards over the T swards, except in 1994. In that year there was a reduction in lamb liveweight gain of 33 g d^?1 on the TC swards and a significant increase in ewe liveweight loss (117 g d^?1) associated with low herbage bulk density despite optimal sward height. Lamb output (kg liveweight ha^?1) on TC swards reflected white clover content, falling from a similar output to that produced from grass given 160 kg N ha^?1, at 18% white clover DM content, down to 60% of grass + N swards with around 5% clover. A 6% greater output from the T than the D swards was achieved mainly through higher stocking rate. The experiment demonstrated a rapid, loss in white clover under lax grazing, and showed that the relationship between performance and sward height is also dependent on herbage density. High lamb output from a grass/clover sward was only achieved when the clover content was maintained at 15–20% of the herbage DM.     

Defoliation and productivity of a phalaris-subterranean clover sward, and the influence of grazing experience on sheep intake

Swards of Phalaris aquatica-Trifolium subterraneum were subjected to four defoliation treatments—zero, low (11 sheep ha⁻¹) and high (22 sheep ha⁻¹) stocking rates, and weekly cutting. At high stocking rate the annual grass Hordeum leporinum dominated while clover was dominant at low and zero stocking rates. Weekly cutting suppressed species other than clover and so failed to simulate grazing.
There were similarities in net herbage production between zero and lightly grazed swards and between heavily grazed and repeatedly cut swards. Net herbage production decreased in the order undisturbed sward < lightly grazed sward < heavily grazed sward < repeatedly cut sward.
When sheep grazed swards where herbage mass was low their daily consumption of herbage, and therefore liveweight change, depended on their recent grazing experience. Sheep accustomed to swards where herbage mass was low ate more because they grazed for much longer each day than unaccustomed sheep, although they selected a diet of similar digestibility.     

INTERACTIONS OF GRASS, CLOVER AND NURSE CROP IN THE SEEDING YEAR

Tall fescue S170 and Italian ryegrass S22, slow- and rapid-establishing grasses, respectively, were sown either with white clover or with a mixture of red and white clovers, with or without a companion- or cover-crop, and with or without nitrogen. Swards established without a cover-crop were grazed frequently or cut infrequently; swards undersown with the oat crop were grazed once in the autumn. In the year of sowing tall fescue/clover mixtures produced yields similar to those of Italian ryegrass/clover swards. Tall fescue proved sensitive to competition, even from red clover, during establishment and sowing under a cover-crop is therefore not recommended. The yield of red- and white-clover was increased when sown under a cover-crop and when cut infrequently, compared with frequent grazing by sheep. The growth of white clover was greatly reduced by adding red clover to the mixture, particularly under a cover-crop and when defoliation was infrequent, but red clover increased total yield. The effect of N applied to the seed-bed varied with the method of establishment. Cereal yield was reduced under dry conditions by undersowing with a mixture containing red clover.     

The effect of cutting for conservation on a grazed perennial ryegrass-white clover pasture

Continuous stocking with sheep at high stocking rates may reduce the content of white clover (Trifolium repens) in mixed grass-clover swards. The present experiment was carried out to investigate the effects on sward production and composition of resting a perennial ryegrass (Lolium perenne)- white clover sward from grazing and taking a cut for conservation. Swards were set-stocked with 25 and 45 yearling wethers ha^?1 either throughout a grazing season, or on swards that were rested for a 6-week period and then cut in early, mid- or late season. In an additional treatment swards were cut only and not grazed. Net herbage accumulation was higher at the lower of the two stocking rates and was marginally increased by the inclusion of a rest period at the high but not the low stocking rate. Clover content was higher at the lower stocking rate and was increased by the inclusion of a rest period by 30% at 45 sheep ha^?1and by 11% at 25 sheep ha^?1 The effect was most marked at the end of the rest period before cutting. When rested from grazing the tiller density of ryegrass decreased although tiller length increased, and clover stolon length, petiole length and leaflet diameter increased though leaf and node number per unit length of stolon decreased; the reverse applied when the sward was returned to grazing after cutting. At the high stocking rate, rest periods in mid-season or later maintained the greatest clover content and marginally increased total net herbage accumulation. At the low stocking rate the timing of the rest period had no significant effect on total net herbage accumulation or on clover content. These results show that the combination of grazing and cutting is of benefit where the stocking rate is high enough to threaten clover survival and limit sheep performance. However, at such a stocking rate, feed reserves are at a minimum throughout the grazing season and so opportunities for resting the sward are probably low.     

Effects of cutting or grazing grass swards on herbage yield, nitrogen uptake and residual soil nitrate at different levels of N fertilization

On a Flemish sandy loam soil, cut and grazed swards were compared at different levels of mineral nitrogen (N) fertilization. Economically optimal N fertilization rates were 400 (or more) and 200 kg N ha^?1 yr^?1 on cut and grazed swards respectively. Considering the amounts of residual soil nitrate‐N in autumn, these N rates also met the current Flemish legal provisions, i.e. no more than 90 kg ha^?1 nitrate‐N present in the 0–90 cm soil layer, measured between 1 October and 15 November. The N use efficiency was considerably higher in cut grassland systems than in grazed systems, even when the animal component of a cut and conservation system was included. The results indicate that, for cut grasslands, two N application rates should be considered: intensively managed grasslands with high amounts of N (400 kg ha^?1 yr^?1 or more) or extensively managed grasslands with white clover and no more than 100 kg N ha^?1 yr^?1.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK. 5. The effect of stocking rate and reduced levels of nitrogen fertilizer

The implications for UK upland sheep systems of reducing nitrogen fertilizer application to perennial ryegrass/white clover swards were studied over 3 years. Sward height (3·5–5·5 cm) was controlled for ewes with lambs until weaning using surplus pasture areas for silage; thereafter, ewes and weaned lambs were grazed on separate areas, and sward height was controlled by adjusting the size of the areas grazed and using surplus pasture areas for silage if necessary. Combinations from three stocking rates [10, 6 and 4 ewes ha⁻¹ on the total area (grazed and ensiled)] and four nitrogen fertilizer levels (150, 100, 50 and 0 kg ha⁻¹) provided six treatments that were replicated three times. Average white clover content was negatively correlated with level of nitrogen fertilizer. The proportion of white clover in the swards increased over the duration of the experiment. Control of sward height and the contribution from white clover resulted in similar levels of lamb liveweight gain on all treatments. All treatments provided adequate winter fodder as silage. It is concluded that the application of nitrogen fertilizer can be reduced or removed from upland sheep pastures without compromising individual animal performance provided that white clover content and sward height are maintained. Resting pastures from grazing by changing ensiled and grazed areas from year to year sustained white clover content over a 3-year period.     

Out-of-season management of grass/clover swards to manipulate clover content

Two experiments are described in which the effect of grazing or defoliating mixed swards at different times over winter and spring on clover content and development was investigated. In the first experiment swards were grazed with sheep (to about 3 cm) for a short period in (a) November, (b) November, January and March, (c) March or (d) not at all, in three consecutive years. All swards were grazed intermittently during the grazing season with cattle and cut for silage once each year. Each plot received either 0 or 50 kg N ha^?1 in March. The effect of N fertilizer was to reduce clover content in each summer and clover growing point density in the third year. In two of the three years, treatments involving grazing in March had lower subsequent net annual herbage accumulation compared with the other two treatments and higher clover content in summer of the third year. Reduction in growing point density in all plots during the grazing season was associated with cattle grazing when conditions were wet, suggesting that stolon burial was implicated. Grazing with sheep in November, January and March resulted in significantly more visible (when counted in situ) clover growing points in April in year 2 and more total growing points (counted after dissection of turves) in the third year than the November grazed and ungrazed treatments which had, on occasions, higher grass tiller density. In a microplot experiment, high herbage mass standing over winter was associated with lower potential photosynthesis per unit clover lamina area and lower growing point density in March. Cutting herbage in March to 2-3 cm resulted in higher clover content and higher growing point number per unit stolon length. The latter was significantly correlated with total irradiance and red: far red at the canopy base. Potential photosynthesis of clover was not affected by cutting in March. It is concluded that growing point density can be increased by grazing or cutting during winter or spring. However, in order for these new stolons to contribute to clover yield during the summer, they have to be maintained until then by ensuring that competition from grass is minimized by keeping the sward short in winter and spring and avoiding the burial of stolons during grazing.     

How does the vertical and horizontal structure of a perennial ryegrass and white clover sward influence grazing?

Mixtures of perennial ryegrass ( Lolium perenne L.) and white clover ( Trifolium repens L.) sown in alternate rows or in a thoroughly mixed matrix were grazed by sheep, either continuously or during short grazing tests, and were used to investigate the influence of the vertical and horizontal components of the sward structure on defoliation by sheep.
In an experiment under continuous grazing, the defoliation intensity was greater for white clover compared with perennial ryegrass leaves (0·80 and 0·58 respectively). In spring, perennial ryegrass leaves were more defoliated than white clover leaves, whereas the reverse was observed in summer. The ratio of the proportion of white clover to perennial ryegrass leaves grazed was negatively correlated with the difference between the surface height of the perennial ryegrass and white clover rows in spring. In both spring and summer, white clover leaves of the same extended leaf length had a higher proportion of them grazed than perennial ryegrass leaves.
In another experiment, during short grazing tests with perennial ryegrass–white clover swards that were grazed at the same sward surface height and at the same white clover content as in the previous experiment, there were no significant differences in the proportion of white clover and perennial ryegrass leaves grazed between strips of the two species and thoroughly mixed structures. The proportion of white clover leaves grazed was higher than that of perennial ryegrass leaves.
These results show that the differential defoliation by sheep of perennial ryegrass and white clover leaves varies according to their vertical distribution in the mixed canopy, but is little affected by their horizontal distribution. Even small differences in sward surface height between mixed perennial ryegrass and white clover can affect diet selection by sheep to a rather large extent.     

The fitting of frequency distributions to height measurements on grazed swards

Herbage height was measured in ryegrass/white clover ( Lolium perenne/Trifolium repens ) swards grazed by cattle using the rising plate meter or the sward stick. Ordinary single normal and double normal distributions were fitted to the data obtained. After the first 6 weeks of continuous grazing the double normal distributions fitted data for the taller swards better than the single normal distribution did, but for a short sward there was no improvement. There was no improvement due to the use of the double normal distribution with data for swards grazed by sheep.     

The liveweight gain of Limousin × Friesian heifers grazing perennial ryegrass/white clover swards of different clover content and the effects of their grazing on sward botanical composition

An experiment was carried out in 1992 and 1993 to examine the effect of white clover content of perennial ryegrass/white clover swards on the performance of Limousin × Friesian heifers. Swards with low (L), medium (M) and high (H) white clover contents were established and managed by continuous variable stocking. A compressed sward height of 5·5 cm was maintained using a buffer fence to vary plot areas, with herbage surplus to grazing requirements cut, removed and yields measured. The mean white clover proportions for treatments L, M and H were 0·02, 0·19 and 0·18 in 1992 and 0·13, 0·16 and 0·31 in 1993 respectively. White clover contents of the swards reached a maximum in August and September, and differences between treatments diminished. There was no significant difference between treatments in the content of white clover in the swards in autumn 1993.
Liveweight gains of heifers increased asymptotically with increasing white clover content of the sward. Below a white clover herbage mass of 300 kg DM ha^–1, there was little effect on liveweight gain, which was 0·70 kg day^–1 over the grazing season. Between 400 and 450 kg DM ha^–1 white clover, liveweight gains were 0·85–0·90 kg day^–1. While clover content of the sward did not significantly affect utilized metabolizable energy output; the mean output over the grazing season in the two years from liveweight gain and herbage yield was 78 GJ ha^–1. It is suggested that, using this grazing system, white clover reached an equilibrium with a mean herbage mass of about 400 kg DM ha^–1 over the grazing season.     

Sward composition, animal performance and the potential production of grass/white clover swards continuously stocked with sheep

The potential productivity of perennial ryegrass/ white clover swards (GC) under continuous stocking management was assessed by comparing their performance, when grazed by sheep at sward surface heights of 3, 6 and 9 cm, with that of an all–grass sward (G) maintained at 6 cm and fertilized with 420 kg N ha^–1 The grass/clover swards received no nitrogen fertilizer. The different grazing treatments had a marked effect on animal performance. In the first year for example, for treatments GC3, GC6, GC9 and G6–420 respectively, mean stocking rates to weaning were 19–7, 14–3, 8–9 and 18–4 ewes ha^–1 (plus twin lambs); lamb growth rates were 223, 268, 295 and 260 g d^–1and so total lamb live weight gain was 1054, 920, 630 and 1148 kg h a^–1. The relative performance of the treatments was similar in all three years. All three grazing treatments had a similar effect on the composition of the grass/clover swards. Clover content increased in 1985, and was sustained in 1986 and 1987 during the main grazing season, although a marked decline in clover content during the winter led to a progressive long–term decline in both the proportion and the amount of clover.
It is suggested that a management based on maintaining a sward surface height close to 6 cm (as in all–grass swards) leads to optimum performance in grass/white clover swards grazed using continuous stocking with sheep. Despite the presence of a small and declining clover content, the output of the mixed grass/clover sward managed in this way was 80%, 80% and 82% of that of a grass sward supplied with 420 kg N ha^–1 in 1985, 1986, and 1987 respectively and, similarly, 83% of the output in 1987 of a grass sward receiving 210 kg N ha^–1.     

The diet ingested by sheep grazing swards differing in white clover and perennial ryegrass content

A series of twenty-four swards containing different proportions of white clover (0·20-0·25) and perennial ryegrass were created by using different seed mixtures, herbicide applications and previous cutting Frequencies. These swards were used to study the diet of oesophageally-fistulated wether sheep which grazed the various swards for a 30-min period after 1, 2 and 3 weeks of regrowth.
The proportion of white clover in the diet was generally greater than that in the sward. Fifty-seven percent of the variation in the proportion of white clover in the diet could be attributed to the proportion of white clover in the sward. White clover and perennial ryegrass leaf and stem were grazed to the same height and the proportion of white clover in the grazed horizon of the sward explained 83% of the variation in the proportion of white clover in the diet. The proportion of white clover in the diet was greater than the proportion in the grazed horizon of the sward in week 3 of regrowth, but not in weeks 1 and 2, and greater when the proportion of white clover in the grazed horizon was lower than 0·20. Both these observations were interpreted as indicating selection for white clover by the sheep within the grazed horizon.
There was a positive and linear relationship between the depth of the grazed horizon and sward height which, together with the relationship between the proportion of white clover in the grazed horizon and in the diet, would allow the prediction of the proportion of white clover of the diet from the height and the white clover content of the grazed horizon of the sward.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK: 6. The effect of reducing stocking rate and application of fertilizer nitrogen in Wales

Abstract The implications for the agricultural productivity of the UK upland sheep systems of reducing nitrogen fertilizer application and lowering stocking rates on perennial ryegrass/white clover swards were studied over 4 years at a site in Wales. The system involved grazing ewes and lambs from birth to weaning on swards maintained at a constant height with surplus herbage made into silage, thereafter ewes and weaned lambs grazed on separate areas until the onset of winter with adjustments to the size of the areas grazed and utilizing surplus pasture areas for silage. Four stocking rates [SR 18, 15, 12 and 9 ewes ha^?1 on the total area (grazed and ensiled)] and two levels of annual nitrogen fertilizer application (N 200 and 50 kg ha^?1) were studied in five treatments (N200/SR18, N200/SR15, N50/SR15, N50/SR12 and N50/SR9). Average white clover content was negatively correlated with the level of annual nitrogen fertilizer application. White clover content of the swards was maintained over the duration of the experiment with an increasing proportion of clover in the swards receiving 50 kg N ha^?1. Control of sward height and the contribution from white clover resulted in similar levels of lamb liveweight gain from birth to weaning in all treatments but fewer lambs reached the slaughter live weight by September at the higher stocking rates and with the lower level of fertilizer application. Three of the five treatments provided adequate winter fodder as silage (N200/SR15, N50/SR12 and N50/SR9). Because of the failure to make adequate winter fodder and the failure of white clover to fully compensate for reduction in nitrogen fertilizer application, it is concluded that nitrogen fertilizer can only be reduced on upland sheep pastures if accompanied by reduced stocking rates.     

Factors affecting the stolon growth of white clover in ryegrass/clover patches

The effects of spatial location of white clover ( Trifolium repens L.) within a perennial ryegrass ( Lolium perenne L.)/white clover pasture on stolon and petiole extension were investigated in two experiments, where patch size containing white clover (0·5 m, 1·5 m and 4 m diameter), location within the patch (inside and edge) and cutting height (4 cm and 8 cm) were varied. Stolon extension rate was greater on the edge of a patch (12·1 mm week⁻¹) than inside the patch (7·2 mm week⁻¹). Patch size affected both stolon and petiole extension rate, which were both greater in small and medium-sized than in large patches. It is suggested that the fastest spread of white clover in patchy sward environments should occur from small patches, which could double in diameter during a growing season. Manipulating the heights of vegetation within and outside large patches affected light quality (red-far red; R/FR) at ground level, which was greater under shorter than taller swards and greater under the canopy of the grass matrix than the grass/white clover patch. However, the height differences between adjacent vegetation had little effect on stolon or petiole growth. In May only, stolon extension at the patch boundary was greatest when both patches and the grass matrix had a height of 8 cm.     

The influence of sward canopy structure on foraging decisions by grazing cattle. I. Patch selection

Patch selection by grazing dairy cows in response to simultaneous variation in combinations of sward structural characteristics was examined in three experiments in which four mature dairy cows were offered a choice of patches (typically 0.9 m × 0.9 m) of perennial ryegrass (Lolium perenne) presented in a linear arrangement. Treatments involved combinations of variations in sward height, stubble height and/or depth of regrowth, prepared by preliminary cutting treatments. They were arranged in balanced sets of four to nine treatments, which were arranged in linear sequences of eighteen to twenty‐seven patches. Within experiments, sequences were balanced across replicate sets of patches, which were grazed separately by individual cows. The number of bites removed and the residence time for each patch were highly correlated in all three experiments, and the results are reported using number of bites per patch as an estimator of foraging behaviour. In the first experiment, with vegetative swards, cows preferentially selected the tallest swards. When swards comprising reproductive stem were offered in Experiment 2, cows selectively grazed short‐stubble swards rather than tall‐stubble swards, although both offered a similar depth of regrowth. Cows did not exhibit preference for swards comprising the greatest quantity of leaf mass, indicating that the spatial distribution of plant components assumed greater importance. In the third experiment, the number of bites removed increased with increasing depth of regrowth, and was negatively correlated with sward height. The three patch‐appraisal cues investigated were broadly ranked in order of importance as (i) depth of regrowth, (ii) sward maturity and (iii) sward height. There was no evidence, at least at a short temporal scale, that patch behaviour was influenced by conditions in adjacent patches, suggesting that the cows assessed grazing opportunities on a patch‐by‐patch basis.     

RESPONSE OF GRASSLAND TO NITROGENOUS FERTILIZER IN THE WEST OF ENGLAND

Extensive replicated plot experiments were carried out at Henley Manor Farm from 1956–1961 to obtain accurate measurements of herbage production and response to nitrogenous fertilizers under West of England conditions. 356 plots were used; use of the herbage by both cutting and grazing was studied. The principal results are: Grass/ clover swards (containing approx. 35% clover) grown without nitrogenous fertilizer averaged 75 cwt dry matter per acre per annum. Application of nitro-chalk to initially grass/clover swards reduced the clover content of the herbage. About 133 lb N/annum was required by an all-grass sward to equal the production of a grass/clover sward without N. Attempts to increase production of grass/clover swards by using N for early growth and relying on clover for mid-season growth were unsuccessful in 2 out of 3 years. Swards (initially grass/clover) gave significant increases in total dry-matter production from regular use of the lowest level of N (26.0 or 34.7 lb N/acre/cut or graze) in 44 out of 49 cases. Extremely high yields were obtained from the heaviest use of N (104.2 lb N/acre/silage cut). 52.1 lb N/acre/cut or graze was the optimum rate of application on ail-grass and grass/clover swards. Up to about 350 lb N/acre/annum the dosage-response curve was very nearly straight for all-grass and grass/clover swards. On the grazed plots herbage left ungrazed amounted on average to only 6% of the total. The health of all stock on the high nitrogen plots (as well as all others) was excellent.