Carbon isotope discrimination and water use efficiency in Iranian diploid,tetraploid and hexaploid wheats grown under well-watered conditions

Carbon isotope discrimination (Δ) has been proposed as physiological criterion to select C₃ crops for yield and water use efficiency. The relationships between carbon isotope discrimination (Δ), water use efficiency for grain and biomass production (WUE_G and WUE_B, respectively) and plant and leaf traits were examined in 20 Iranian wheat genotypes including einkorn wheat (Triticum monococcum L. subsp. monococcum) accessions, durum wheat (T. turgidum L. subsp. durum (Desf.) Husn.) landraces and bread wheat (T. aestivum L. subsp. aestivum) landraces and improved cultivars, grown in pots under well-watered conditions. Carbon isotope discrimination was higher in diploid than in hexaploid and tetraploid wheats and was negatively associated with grain yield across species as well as within bread wheat. It was also positively correlated to stomatal frequency. The highest WUE_G and grain yield were noted in bread wheat and the lowest in einkorn wheat. Einkorn and bread wheat had higher WUE_B and biomass than durum wheat. WUE_G and WUE_B were significantly negatively associated to Δ across species as well as within bread and durum wheat. The variation for WUE_G was mainly driven by the variation for harvest index across species and by the variation for Δ within species. The quantity of water extracted by the crop, that was closely correlated to root mass, poorly influenced WUE_G. Environmental conditions and genetic variation for water use efficiency related traits appear to highly determine the relationships between WUE_G and its different components (water consumed, transpiration efficiency and carbon partitioning).     

Ontogenetic-based sequential path analysis of grain yield and its related traits in several winter wheat cultivars

Studies that provide representative insights for determining yield through its related traits during the ontogeny of modern cultivars subjected to sources of environmental variation are limited for different crops, including wheat. Most of the empirical evidence on the relationships between the yield of small grain cereal crops and its contributing traits has been obtained under dry or semidry conditions. The aims of this paper were to (1) illustrate how an path analysis can be used to clarify and interpret the relationships between grain yield (GY), yield components, and other yield-related traits of 25 winter wheat cultivars subjected to sources of environmental variation and (2) determine how the yield-related traits contribute to the yield variation. The data used in this analysis were generated from multi-environment trials across wheat-growing areas in Poland. Using Ward’s clustering procedure was capable of identifying the most critical predictor traits of the yield components and their contributions to cultivar-focused GY variation. Our findings document, confirm, and improve the basic biological understanding of how to grow modern wheat cultivars for high GY through effectively stimulating the improvement of yield-related traits through the optimization of developmental stage-based agronomic strategies. Our results confirmed empirically that modern European wheat cultivars grown in a temperate climate require favorable conditions, the use of appropriate N fertilizer and growth regulators, and the application of fungicide to protect against leaf diseases and to provide conditions that effectively increase the time to anthesis, the Leaf Area Index per spike at anthesis, and the grain filling duration, and reduce plant height and flag leaf disease severity, thus leading to a high GY. A high yield level is obtained by the performance of preferred yield-related traits that can maintain the three yield components at relatively high levels.     

Relationships among important traits in the nitrogen economy of winter wheat

Identification and utilization of important attributes in the nitrogen economy of wheat (Triticum aestivum L.) should provide a basis for increasing grain protein percentage (GPP). The objectives of this study were to determine the magnitude of genetic variability of some factors important in the nitrogen economy of wheat, their relationship to one another, and their influence on grain yield (GY) and GPP. Twenty‐five hard red winter wheat genotypes representing a wide range of GY, GPP, and plant stature were grown in field trials over a 2‐year period. Significant differences were found for harvest index (HI) and nitrogen harvest index (NHI). Total nitrogen at anthesis (TNA), total nitrogen at maturity (TNM), and biological yield (BY) did not differ significantly. GPP was positively correlated with TNM and BY and negatively correlated with TNA, HI, and GY. GPP was not correlated with NHI, however, stepwise regression of GPP and grain protein yield revealed NHI as a common component, with a positive coefficient. In addition, NHI was positively correlated with GY. Selection of parents with complementary traits for nitrogen‐use efficiency may allow for simultaneous increases in GY and GPP.     

Predicting grain yield and protein content in winter wheat at different N supply levels using canopy reflectance spectra

A field experiment using a split-plot randomized complete block design with three replications was carried out to determine relationships between spectral indices and wheat grain yield （GY）, to compare the performance of four vegetation indices （VIs） for GY prediction, and to study the feasibility of VI to estimate grain protein content （GPC） in winter wheat. Two typical winter wheat （Triticum aestivum L.） cultivars ＇Xuzhou 26＇ （high protein content） and ＇Huaimai 18＇ （low protein content） were used as the main plot treatments and four N rates, i.e., 0, 120, 210, and 300 kg N ha^-1, as the sub-plot treatments. Increasing soil N supply significantly increased GY and GPC （P ≤ 0.05）. For the two cultivars combined, significant and positive correlations were found between four VIs and GY, with the strongest relationship observed when using the green ratio vegetation index （GRVI） at mid-filling. Cumulative VI estimates improved yield predictions substantially, with the best interval being heading to maturity stage. Similar results were found between VI and grain protein yield. However, when using cumulative VI, GPC showed no significant improvement. The strong relationship between leaf N status and GPC （R2 =0.9144 for ＇Xuzhou 26＇ and R2 = 0.8285 for ＇Huaimai 18＇） indicated that canopy spectra could be used to predict GPC. The strong fit between estimated and observed GPC （R2 = 0.7939） indicated that remote sensing techniques were potentially useful predictors of grain protein content and quality in wheat.     

Manganese relationships in spring wheat and spring barley production in Northern Idaho

Abstract

Manganese is often limiting in cereal crop production in the Kootenai River Valley of Northern Idaho; however, attempts to relate DTPA‐extractable Mn in soils to crop yield response have not been successful. Consequently, Mn plant tissue diagnosis may be an alternate diagnostic tool. The objectives of this study were to: (1) determine the critical nutrient concentration (CNC) of Mn in spring wheat and spring barley tissue in the Kootenai River Valley of northern Idaho, (2) examine yield response of spring wheat and spring barley to Mn fertilization, and (3) assess relative efficiencies of foliar and surface broadcast Mn applications to these crops. Paired plots with four replications consisting of a foliar Mn application rate of 1.5 kg ha^‐1 and a control were located at four sites in 1988 and at ten sites in 1989. Soil, plant tissue samples and grain yield data were evaluated by linear plateau regression to determine CNCs of Mn for spring wheat and spring barley tissue. In addition, five randomized complete block experiments were conducted in 1989 and 1990 to evaluate Mn fertilizer rates and sources (foliar vs. surface applied) on spring wheat and spring barley production. Tissue Mn was highly correlated by linear plateau regression to both spring wheat (r² = 0.74**) and spring barley (r² = 0.70**) grain yield. The Mn CNC was established at 11.0 mg Mn kg‐¹ plant tissue for spring wheat and 10.1 mg Mn kg^‐1 plant tissue for spring barley. The Mn CNCs were established at 92.4 and 93.0% of maximum yield for spring wheat and spring barley, respectively. DTPA‐extractable Mn was not significantly correlated to grain yield for either crop (r² = 0.02, NS). Based on study results, Mn analysis of spring wheat and spring barley plant tissue was diagnostic of eventual grain yield. When tissue diagnosis showed plants to be deficient in Mn, the deficiency was corrected by applying Mn fertilizer as a surface broadcast or a foliar spray. However, foliar application of Mn was more efficient than broadcast application.     

Effect of Phosphorus Additions and Arbuscular Mycorrhizal Fungal Inoculation on the Growth,Physiology, and Phosphorus Uptake of Wheat Under Two Water Regimes

Drought stress greatly affected the growth and development of wheat in the world, while wheat growth could benefit through improvement of water status and nutrient uptake by mycorrhizal symbiosis or addition of phosphorus (P). Experimental treatments were (a) phosphorus addition (0 and 90 kg/ha), (b) soil water condition (40% field capacity and 95% field capacity), and (c) arbuscular mycorrhizal fungi (AMF, Glomus intraradices) (noninoculation and inoculation) which were conducted in a growth chamber. The results showed that addition of phosphorus and AMF inoculation significantly increased the relative water content and specific leaf area of flag leaves especially under 40% field capacity (water deficit (WD)). The leaf gas exchange parameters were all decreased under WD. The water use efficiency (WUE) and instantaneous WUE (WUEi) was enhanced by WD, AMF inoculation, and phosphorus addition. AMF inoculation and WD significantly decreased the carbon isotope discrimination (CID) of leaf. The P concentrations in stem, grain, and leaf were significantly increased by phosphorus addition, WD, and AMF inoculation. Significant correlations were found between WUE and grain P, stem P, and leaf P concentrations. Leaf CID was significantly negatively correlated with WUE and stem P concentrations. Inoculation of AMF or phosphorus addition could improve the growth, physiology, and phosphorus uptake in spring wheat under drought conditions.     

不同水、氮处理对济麦20产量和蛋白质品质的影响

以强筋小麦济麦20为材料,在防雨棚肥水控制池条件下,研究了不同灌水和施氮处理对小麦产量和蛋白质品质的影响。结果表明,开花期灌水有利于抑制小麦生育后期旗叶叶绿素的降解,提高叶片含氮量,延长旗叶功能期;开花期灌水和灌浆期灌水有利于小麦产量和千粒重的提高,以春季灌3水(春5叶露尖+开花+灌浆)最高;在180kg.N/hm2条件下,籽粒产量及千粒重、容重和穗粒数均高于270kg.N/hm2的处理,对穗粒数的影响达显著水平。籽粒蛋白质及各组分的含量,除球蛋白外,均以B3处理(灌春2叶露尖水和春5叶露尖水)最高;增施氮肥有利于谷蛋白含量的提高。籽粒蛋白质产量以B4处理(春5叶露尖水+开花水+灌浆水)最高,与B2处理(春5叶露尖水+开花水)差异不显著;180kg.N/hm2处理的蛋白质产量高于270kg.N/hm2处理。在本试验条件下,180kg/hm2的施氮量处理结合春5叶露尖灌水和开花期灌水有利于强筋小麦产量和品质的共同提高。     

Estimation of yield and dry matter of winter wheat using logistic model under different irrigation water regimes and nitrogen application rates

Complex simulation models need considerable input data that may be unavailable. In this study, based on growing degree days, a simple and empirical logistic model was used to predict top dry matter (DM) and grain yield (GY) of winter wheat in a semiarid area. For calibration and validation of the model, winter wheat cv. Shiraz was cultivated for two growing seasons (2009–2010 and 2010–2011) under different levels of irrigation water and nitrogen application rates. The coefficients of the logistic model and the harvest index were related to the seasonal amount of applied water (I) plus rainfall (R) and applied nitrogen plus residual soil mineral nitrogen (N + N_r) with good accuracy in 2009–2010. The values of DM were estimated using the developed empirical logistic model during the growing season in 2010–2011. Results also indicated that there was a good agreement between the measured and predicted GY in 2010–2011 with fair accuracy. Therefore, it is concluded that the presented model is appropriate for prediction of DM and GY of winter wheat at the study region.     

Agronomic Assessment of Nitrogen Use Efficiency in Spring Wheat and Interrelations with Leaf Greenness Under Field Conditions

Four spring wheat genotypes (Triticum aestivum L.) were grown without (N0 = 0 kg N ha^?1) and under ample (N1 = 250 kg ha^?1) nitrogen (N) fertilizer in field experiments in two seasons. The aim was to assess genotypic variation in N use efficiency (NUE) components and N-related indices during grain filling thus to identify superior wheat genotypes. Leaf chlorophyll (SPAD) readings at crucial growth stages were employed to help differentiate genotypes. Interrelations between yield and N-related indices with SPAD, where also assessed to explain possible pathways of improving NUE early in the growing season. Results showed that genotypic effects on NUE were mostly evident in 2000, a year with drier preanthesis and wetter postanthesis than the normal periods. ‘Toronit’ almost always had the highest biomass yield (BY) and grain yield (GY). Except in 1999 under N0, ‘L94491? showed the highest % grain N concentration (GNC). Genotypes affected SPAD at almost all stages and N fertilization delayed leaf senescence for all genotypes and growth seasons. Correlations between SPAD at different growth stages and GY, N biomass yield at maturity (NBYM) and GNC were significant (P≤ 0.001), positive and strong/very strong (>r = 0.7). N translocation efficiency (NTE) was inversely related to PANU (~r = ? 0.77, P≤ 0.001), suggesting that N after anthesis is being preferentially transported to the ears to meet the N demand of the growing grains. It is concluded that there is still a large potential for increased NUE by improved N recirculation, use of fast and inexpensive crop N monitoring tools and high yielding, N uptake efficient genotypes.

Abbreviations: NUE, Nitrogen use efficiency; SPAD, Minolta SPAD-502 chlorophyll meter, NHI, nitrogen harvest index; HI, Harvest index; NTE, N translocation efficiency from vegetative plant parts to grain; DMTE, dry matter translocation efficiency; CPAY, contribution of pre-anthesis assimilates to yield; PANU, Post-anthesis N uptake, d.a.s., days after sowing, N0, zero (0) kg ha^?1 applied N fertilizer, N1, 250 kg ha^?1 applied N fertilizer.     

Use of Plant-Growth-Promoting Rhizobacteria (PGPR) Seed Inoculation as Alternative Fertilizer Inputs in Wheat and Barley Production

In this study we aimed to investigate the effects of plant-growth-promoting rhizobacteria (PGPR) on seed incubation of spring wheat and barley. Three bacterial strains were applied singly and in combinations. Seed inoculation with strains significantly affected grain yield (GY), straw (SWY), total yield (TY), and plant nutrient element (PNE) content. In field trials, compared to the control, single inoculations gave GY, SWY, and TY increases by 27.5–31.9%, 1.1–5.3%, and 1.3–11.3% in wheat and 15.1–27.8%, 10.8–15.5%, and 14.5–18.5% in barley, respectively, but mixtures of strains gave increases in GY, SWY, and TY by 54.7%, 2.1%, and 6.7% in wheat and 57.8%, 14.6%, and 17.5% in barley, respectively. According to the results, it was concluded that seed inoculations with PGPR and mixture inoculation might satisfy nitrogen requirements, but Bacillus megaterium M3 and MIX (Bacillus subtilis OSU142, B. megaterium M3, Azospirillum brasilense Sp245) inoculation provided greater PNE concentrations than mineral fertilizer application for wheat and barley under field conditions.     

Furrow diking and N management for dryland wheat production in permanent raised beds

The permanent bed planting system for wheat (Triticum aestivum L.) production has recently received additional attention. Studies using hard red spring wheat (cultivar Nahuatl F2000) were conducted at two locations in central Mexico. The studies included the installation of three furrow diking treatments, two granular N timing treatments and three foliar N rates applied at the end of anthesis. The objective was to evaluate the effect of these factors on wheat grain yield, yield components and grain N in a wheat–maize (Zea maize L.) rotation with residues of both crops left as stubble. Results indicated that diking in alternate furrows increased both grain yield and the final number of spikes per m². The split application of N fertilizer enhanced the number of spikes per m² and grain N uptake, but the effect on grain yield was inconsistent. Similarly, grain protein increased with the foliar application of 6 kg N ha^?1, depending upon the maximum temperature within the 10 days following anthesis. The normalized difference vegetative index (NDVI) readings collected at four growth stages were generally higher for the split N application than for the basal N application at planting. Grain N uptake was associated to NDVI readings collected after anthesis.     

Nitrogen Leaching Of Spring Wheat Genotypes (Triticum Aestivum L.) Varying In Nitrogen-Related Traits

Efficient use of nitrogen (N) by wheat crop and hence prevention of possible contamination of ground and surface waters by nitrates has aroused environmental concerns. The present study was conducted in drainage lysimeters for three years (1998–2000) to identify whether spring wheat genotypes (Triticum aestivum L.) that differ in N-related traits differ in N leaching and to relate parameters of N use efficiency (NUE) with parameters of N leaching. For this reason two spring wheat cultivars (‘Albis’ and ‘Toronit’) and an experimental line (‘L94491’) were grown under low (20 kg N ha^?1) and ample N supply (270 kg N ha^?1). The genotypes varied in parameters of NUE but not in N leaching. Grain yield of the high-protein line (‘L94491’) was, on average, 11% lower than that of ‘Toronit’ but among genotypes had significantly higher N in the grain (%), grain N yield, and N harvest index. Nitrogen lost through leaching was considerably low (0.42–0.52 g m^?2) mainly due to low volume of percolating water or the ability of the genotypes to efficiently exploit soil mineral N. There were no clear relationships between N-related genotype traits and N leaching, but across all treatments significantly negative correlations between volume of leachate and the amount of N in the total biomass and grain N yield existed.     

Utilizing Existing Sensor Technology to Predict Spring Wheat Grain Nitrogen Concentration

Optimum grain nitrogen (N) concentration and yield in spring wheat (Triticum aestivum L.) can be problematic without proper N fertilizer management. Sensor-based technologies have been used for application of fertilizers and also to predict yield in wheat, although little has been done in the prediction of grain N. Field studies were conducted in South Dakota in 2006 (Gettysburg, Bath, and Cresbard) and 2007 (Gettysburg, Aurora, Leola, and Artas). There were five N treatments (0, 56, 112, 168, and 224 kg N ha^?1) applied pre-plant with a second N application applied foliar at anthesis. Sensor readings were taken at growth stages Feekes 10, anthesis, and postfoliar application using the GreenSeeker Hand Held optical sensor. Grain samples were taken at maturity and analyzed for total N. Using similar information collected in 2003 and 2005, a critical normalized difference vegetation index (NDVI) value was determined using the Cate–Nelson procedure. The critical NDVI value needed to ensure optimum grain N was 0.70. In 2006 and 2007, the plots that received an application of N at anthesis had higher grain N than the plots not receiving N. There was also a significant response between applied N and grain yield. The results show that with further studies, the Greenseeker could be used to apply N to maximize yield and grain N in a precise and accurate manner.     

The variation of phosphorous content,grain yield,and rhizosphere microbial biomass among durum wheat cultivars under salinity stress

ABSTRACT

The effectiveness of plant–soil synergies is largely modulated by interaction between cultivar and rhizosphere microbiome. We evaluated the agronomic performance of six durum wheat cultivars, in two semi-arid locations in Tunisia that differed in their irrigation water salinity: S1 (6 dS m^?1) and S2 (12 dS m^?1). The two-consecutive-year field experiments assessed the effects of the microbial biomass carbon (MBC), leaf phosphorus (LP) and rhizosphere phosphorus (P) on the grain yield (GY) and yield components at tillering and flowering stages. Overall, in saline conditions, cultivars differed in above- and below-ground traits, particularly, with tolerant cultivars presenting relatively greater MBC, P and LP. Furthermore, in S2, GY positively correlated with MBC (r = 0.69), LP (r = 0.80) and P (r = 0.79). Additionally, in S2, MBC positively correlated with P (r = 0.87) and LP (r = 0.85) at flowering. This result was further confirmed by multiple linear regression (step-wise) analysis, which revealed that MBC and LP were the determinant components of GY variability under S2. The present study demonstrates that LP and soil P are mandatory for improving the management of durum wheat. Salinity tolerance was largely affected by the cultivars’ rhizosphere MBC.     

MAIZE RESPONSE TO YIELD AND YIELD TRAITS WITH DIFFERENT NITROGEN AND DENSITY UNDER CLIMATE VARIABILITY

To study yield and yield traits of maize, two experiments were conducted in 2006 as spring and summer crops and repeated in 2007. Three plant populations (43, 53, and 67 thousands ha^-1) and three nitrogen (N) rates (90, 120 and 150 kg N ha^-1) were compared in a completely randomized block design with split plot arrangement. The treatments plant population was assigned to the main and N to sub plots in three replications. Sowing of spring crop was done in March and harvested in July and likewise the summer crop in July and harvested in November. Each experimental unit comprised of 5 × 6 m area having eight rows spaced 0.75 m. Experimental results revealed that grain yield (GY) of summer was higher than spring season. Higher GY was associated with increases in the plant population and nitrogen rate. The treatment plant population of 53,000 and 67,000 ha^-1 did not differ. However, each increase in the N rate significantly increased GY. This increase in the GY by increasing N was due to increases in the ear length (EL), ear diameter (ED), grain number (GN) and thousand grain weight (TGW). Increasing plant population of maize did not show any remarkable changes in the yield traits. The study revealed that differences in GY due to seasonal climate cannot be rewarded with increase in either plant population or nitrogen rates. Moreover, spring and summer season maize crops can economically be planted with 150 and 120 kg N ha^?1 at 53,000 and 67,000 ha^?1 populations, respectively, to save environment and production cost.     

Boron and Amino Acid Foliar Application on Wheat-Soybean Intercropping in a Non-Tillage System

Micronutrient and amino acid (AA) foliar fertilization has generally been sprayed onto plants to increase the crop yield. The experiment had the aim of evaluating the foliar boron (B) and AA application on grain yield (GY), physiological characteristics, nutritional status, and yield components in wheat (Triticum aestivum L.) and soybean [Glycine max (L.) Merrill] intercropping in a non-tillage system (NTS). The experiment was set up as a randomized block design with eight treatments and four replicates. The treatments had the following boron (B) rates: [0, 1, 2, 4, and 8 kg ha^?1, source: boric acid (H₃BO₃)] + AAs (2 L ha^?1) applied by foliar spraying and the additional treatments [(Control - without B and AAs), 2 kg ha^?1 B, 2 L ha^?1 AAs and 2 kg ha^?1 B + 1 L ha^?1 AAs] applied at the end of the elongation and spike beginning of wheat plants and development growth stage (V5) of soybean for two growing seasons. Boron and AAs had no influence on the physiological and yield components and had no increases in the foliar and grain B content in wheat and soybean. No matter the dose, the foliar B + AAs (2 L ha^?1) application did not increase the GY in wheat-soybean under a rotational NTS in loamy soil with suitable available B.     

Within-Field Variability of Winter Wheat Yield and Grain Quality versus Soil Properties

Within-field variability in wheat grain yield and its quality always exists in production fields and depends, among other factors, mainly on various soil properties related to nutrients and water availability. The aim of the research was to examine the relationships between selected soil properties such as texture; pH; content of the available phosphorus (P), potassium (K), magnesium (Mg), and organic carbon; and winter wheat grain yield and quality under rainfed conditions. Six crop fields with winter wheat (Triticum aestivum, L.) in three sites located in different regions of Poland were examined during two seasons. The grain yield was mainly determined by the soil texture, and the majority of the chemical soil properties did not have a significant effect on grain yield. The grain quality traits were determined by the examined soil properties to a smaller degree than grain yield. The relationships were not consistent across sites and years.     

Alleviation of drought stress in winter wheat by late foliar application of zinc,boron, and manganese

In many regions, drought during flowering and grain‐filling inhibits micronutrient acquisition by roots resulting in yield losses and low micronutrient concentrations in cereal grains. A field and a greenhouse experiment were conducted to study the effect of foliar applications of zinc (Zn), boron (B), and manganese (Mn) at late growth stages of winter wheat (Triticum aestivum L.) grown with or without drought stress from booting to maturity. Foliar applications of Zn, B, and Mn did not affect grain yield in the absence of drought. However, under drought, foliar application of Zn and B in the field increased grain yield (15% and 19%, respectively) as well as raising grain Zn and B concentration, while Zn and Mn sprays in the greenhouse increased grain yield (13% and 10%, respectively), and also increased grain Zn and Mn concentrations. Furthermore, under drought stress both in the field and greenhouse experiment the rate of photosynthesis, pollen viability, number of fertile spikes, number of grains per spike, and particularly water‐use efficiency (WUE) were increased by late foliar application of micronutrients. These results indicate that by increasing WUE foliar application of Zn, B, and Mn at booting to anthesis can reduce the harmful effects of drought stress that often occur during the late stages of winter wheat production. These findings therefore are of high relevance for farmers' practice, the extension service, and fertilizer industry.     

Foliar application of silicon at different growth stages alters growth and yield of selected wheat cultivars

To evaluate the response of some selected wheat cultivars to silicon application at different growth stages under drought stress, an experiment was carried out in the greenhouse of College of Agriculture, Shiraz University, Iran, during 2012 using a completely randomized factorial design with four replications. Experimental treatments included drought stress (100% F.C. as control and 40% F.C. as drought) and foliar application of 6 mM sodium silicate (control, application at mid tillering stage, at anthesis stage, and application at tillering + anthesis stages) and wheat cultivars (Sirvan and Chamran, relatively drought-tolerant, and Shiraz and Marvdasht, drought-sensitive cultivars). Drought stress significantly reduced chlorophyll content, leaf area, relative water content, grains per spike, 1000-grain weight, grain yield and biomass of all wheat cultivars. Furthermore, drought stress increased electrolyte leakage of the flag leaves of all cultivars. In contrast, foliar-applied silicon significantly increased these parameters and reduced electrolyte leakage. Furthermore, highest positive influence of silicon application was observed at combined use of silicon both at the tillering + anthesis stages in wheat plants under both stress and non-stress conditions. Significant differences were found in physiological responses among wheat cultivars. The drought tolerant cultivars (Sirvan and Chamran) had significantly higher growth and yield than those of drought sensitive cvs. Shiraz and Marvdasht under drought stress. In conclusion, foliar application of silicon especially at the tillering + anthesis stages was very effective in promoting resistance in wheat plants to drought conditions by maintaining cellular membrane integrity and relative water content, and increasing chlorophyll content.     

EUF-extrahierbarer Stickstoff und dessen Beziehung zu Stickstoffaufnahme und Ertrag von Weizen

EUF extractable nitrogen and its relation to nitrogen uptake and yield of wheat The investigations were focussed on the question whether nitrogen extracted from soils by EUF may reflect nitrogen availability for cereals. Soil samples were taken from the upper soil layer of farmers' fields differing in soil type and texture. Samples still taken in the autumn after the harvest of sugar beets as well as samples taken under cereals in January had relatively high contents of EUF extractable organic N and low contents of NO₃. Decrease in EUF extractable N during spring (March until end of May) under winter wheat was significantly correlated with the N uptake of the wheat and with grain yield. In the plots not treated with fertilizer N the organic N of the EUF-extracts was as a mineralizable N source more important for crop nutrition than the EUF extractable NO₃. The correlation between the EUF extractable N in spring and the grain yield of wheat was highly significant (r = 0.79***). In this correlation 20 different locations were implicated. Similar good correlations were found between the EUF extractable N and the N uptake of wheat.